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Type: Article
Published: 2025-08-29
Page range: 428-438
Abstract views: 133
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Anaspis Geoffroy, 1762 subgenus Spanisa Emery, 1876 (Coleoptera: Scraptiidae) as Paleogene relicts

Department of Life Sciences, Natural History Museum, Cromwell Road, SW7 5BD, London, United Kingdom; Coleopterological Research Center, Institute of Life Sciences and Technology, Daugavpils University, LV-5401, Daugavpils, Latvia; Institute of Biology, University of Latvia, O. Vācieša iela 4, LV-1004, Rīga, Latvia
Natural History Museum of Denmark, Universitetsparken 15, DK-2100, Copenhagen, Denmark; I.I. Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kyiv 01030, Ukraine
Coleoptera Scraptiidae Anaspidinae fossils taxonomy morphology palaeodiversity key distribution

Abstract

Anaspis (Spanisa) solodovnikovi sp. nov., the second known representative of its genus and subgenus from the late Eocene Rovno amber of Ukraine, is described and illustrated. The taxonomic position is confirmed, and supplementary description and images are provided for the Eocene Baltic amber A. (Silaria) parva Abdullah, 1964. A key to extant and extinct species of the subgenus Spanisa Emery, 1876 is presented for the first time. Morphology and new taxonomic position of Anaspis (incertae sedis) subhumerosa Franciscolo, 1954 are briefly discussed; this species is excluded from the subgenus Spanisa. Extant Spanisa species are allopatric, but in Rovno amber they are sympatric likely because of their adaptation to the ‘extinct’ equable Eocene climate. New faunistic records are provided for the extant A. (Spanisa) labiata A. Costa, 1854 and A. (Spanisa) subtilis Hampe, 1871.

References

  1. Abdullah, M. (1964) New heteromerous beetles (Coleoptera) from the Baltic amber of eastern Prussia and gum copal of Zanzibar. Transactions of the Royal Entomological Society of London, 116, 329–346. https://doi.org/10.1111/j.1365-2311.1964.tb02300.x
  2. Alekseev, V., Kirejtshuk, A.G. & Bukejs, A. (2025) The second species of Lobanoviella Kirejtshuk & Reid, 2021 (Coleoptera: Megalopodidae) in Eocene amber. Palaeoentomology, 8 (2), 166–173. https://doi.org/10.11646/palaeoentomology.8.2.7
  3. Archibald, S.B. & Farrell, B.D. (2003) Wheeler’s dilemma. Acta Zoologica Cracoviensia, 46 (Supplement—Fossil Insects), 17–23.
  4. Belokobylskij, S.A., Simutnik, S.A., Vasilenko, D.V. & Perkovsky, E.E. (2023) First record of the parasitoid subfamily Doryctinae (Hymenoptera, Braconidae) in Rovno amber: description of a new genus and species with stigma-like enlargement on the hind wing of the male. Journal of Hymenoptera Research, 95, 59–72. https://doi.org/10.3897/jhr.95.96784
  5. Chemyreva, V.G., Legalov, A.A. & Perkovsky, E.E. (2024) A new genus of Ambositrinae (Hymenoptera, Diapriidae) from Rovno amber and remarks on the Eocene distribution of the subfamily. Ecologica Montenegrina, 79, 104–112. https://doi.org/10.37828/em.2024.79.9
  6. Emery, M.C. (1876) Essai monographique sur les mordellides de l´Europe et des contrées limitrophes. L’Abeille, Journal d’Entomologie, 14 [1874–1876], 1–128.
  7. Ermisch, K. (1950) Die Gattungen der Mordelliden der Welt. Entomologische Blätter, 45-46 (1), 34–92.
  8. Esser, J. (2001) Bemerkenswerte Käferfunde aus Sachsen-Anhalt. Entomologische Nachrichten und Berichte, 45 (3-4), 228–230.
  9. Fedotova, Z.A. & Perkovsky, E.E. (2008) New taxa of gall midges (Diptera, Cecidomyiidae) from Dubrovitsa (Rovno amber). Vestnik Zoologii, 42 (1), 27–40. [In Russian with English abstract] https://doi.org/10.1007/s11492-008-2007-6
  10. Franciscolo, M.E. (1954) On two species of Anaspidinae (Coleoptera: Scraptiidae) taken on flowers of Protea abyssinica in Natal. (xxviii. Contribution to the knowledge of Mordellidae and Scraptiidae. Proceedings of the Entomological Society of London B, 23 (3-4), 63–73. https://doi.org/10.1111/j.1365-3113.1954.tb00103.x
  11. Franciscolo, M.E. (1972) Su alcuni generi poco noti di Anaspidinae. Memorie della Società Entomologica Italiana, 51, 123–155.
  12. ICZN (International Commission on Zoological Nomenclature) (1984) Opinion 1273 Anaspis, Luperus, Lampyris and Clerus (Insecta, Coleoptera): Determination of authorship and fixation of type species. Bulletin of Zoological Nomenclature, 41 (1), 28–31.
  13. Ivanov, V.D., Melnitsky, S.I. & Perkovsky, E.E. (2016) Caddisflies from Cenozoic resins of Europe. Paleontological Journal, 50 (5), 485–493. https://doi.org/10.1134/S0031030116050063
  14. Iwan, D. & Kubisz, D. (2020) Family Scraptiidae Mulsant, 1856. In: Iwan, D. & Löbl, I. (Eds), Tenebrionoidea. Revised and updated second edition. Catalogue of Palaearctic Coleoptera. Volume 5. Brill, Leiden & Boston, pp. 631–641. https://doi.org/10.1163/9789004434998_004
  15. Janák, J. (2024) Four new species and new records of Paederinae from Baltic and Ukrainian (Rovno) amber (Coleoptera: Staphylinidae). Studies and Reports, Taxonomical Series, 20 (1), 97–113.
  16. Johnston, M.A., Naczi, R.F.C. & Gimmel, M.L. (2024) Review of North American Scraptiidae (Coleoptera: Tenebrionoidea), with a catalog of world genera. The Coleopterists Bulletin, 78 (2), 171–200. https://doi.org/10.1649/0010-065X-78.2.171
  17. Kazantsev, S.V. (2025) A review of the fossil genus Mimoplatycis Kazantsev, 2013 (Coleoptera Cantharidae: Malthininae). Zootaxa, 5618 (2), 267–274. https://doi.org/10.11646/zootaxa.5618.2.5
  18. Kazantsev, S.V., Legalov, A.A. & Perkovsky, E.E. (2025) A new species of the fossil genus Sucinorhagonycha Kuśka, 1996 (Coleoptera: Cantharidae: Cantharinae) from late Eocene Rovno amber, with a taxonomic note. Ecologica Montenegrina, 81, 54–62. https://doi.org/10.37828/em.2025.81.7
  19. Koch, K. (1989) Die Käfer Mitteleuropas. Ökologie. Band 2. Goecke & Evers, Krefeld, 382 pp.
  20. Köhler, F. & Klausnitzer, B. (1998) Verzeichnis der Käfer Deutschlands. Entomologische Nachrichten und Berichte, Beiheft 4, 185 pp.
  21. Legalov, A.A., Vasilenko, D.V. & Perkovsky, E.E. (2023a) From abundance to extinction: evolutionary history of European Aedemonini (Curculionidae) with a description of the first representative from Rovno amber. Diversity, 15, 376. https://doi.org/10.3390/d15030376
  22. Legalov, A.A., Vasilenko, D.V. & Perkovsky, E.E. (2023b) New proxy for Moraceae in Priabonian of Europe: first record of the genus Demimaea Pascoe, 1870 (Coleoptera: Curculionidae) from Eocene Rovno amber. Historical Biology, 35 (8), 1322–1328. https://doi.org/10.1080/08912963.2022.2089983
  23. Lyubarsky, G.Yu., Perkovsky, E.E. & Vasilenko, D.V. (2023) Unexpected diversity of Xenoscelinae in Priabonian European amber: the third xenosceline species from Rovno amber. Life, 13 (3), 636. https://doi.org/10.3390/life13030636
  24. Lyubarsky, G.Yu., Legalov, A.A., Vasilenko, D.V. & Perkovsky, E.E. (2024a) The first fossil of subgenus Neomicrambe (Coleoptera: Cryptophagidae: Micrambe) from Danish amber. Ecologica Montenegrina, 72, 227–234. https://doi.org/10.37828/em.2024.72.21
  25. Lyubarsky, G.Yu., Legalov, A.A., Vasilenko, D.V. & Perkovsky, E.E. (2024b) A second species of the subgenus Neomicrambe (Coleoptera: Cryptophagidae: Micrambe) from Danish amber. Ecologica Montenegrina, 79, 125–132. https://doi.org/10.37828/em.2024.79.12
  26. Mänd, K., Muehlenbachs, K., McKellar, R.C., Wolfe, A.P. & Konhauser, K. (2018) Distinct origins for Rovno and Baltic ambers: evidence from carbon and hydrogen stable isotopes. Palaeogeography, Palaeoclimatology, Palaeoecology, 505, 265–283. https://doi.org/10.1016/j.palaeo.2018.06.004
  27. Matalin, A.V., Perkovsky, E.E. & Vasilenko, D.V. (2021) First record of tiger beetles (Coleoptera, Cicindelidae) from Rovno amber with the description of a new genus and species. Zootaxa, 5016 (2), 243–256. https://doi.org/10.11646/zootaxa.5016.2.5
  28. Medvedev, L.N. (1969) New Mesozoic beetles (Cucujoidea) from Asia. Paleontological Journal, 3 (1), 19–25.
  29. Moragues y de Manzanos, F. (1889) Coleópteros de Mallorca. Anales de la Sociedad Espanola de Historia natural, 18, 11–34.
  30. Odnosum, V.K. (2013) About the study of fossil mordellid and scraptiid-beetle (Coleoptera: Mordellidae, Scraptiidae) of the Old Word. Ukrayins’kyy entomolohichnyy zhurnal, 1 (6), 43–49. [In Russian with English abstract]
  31. Olmi, M., Guglielmino, A., Vasilenko, D.V. & Perkovsky, E.E. (2022) Discovery of the first apterous pincer wasp from amber, with description of a new tribe, genus and species of Apodryininae (Hymenoptera, Dryinidae). Zootaxa, 5162 (1), 54–66. https://doi.org/10.11646/zootaxa.5162.1.3
  32. Perkovsky, E.E. & Odnosum, V.K. (2009) A new species of the scraptiid beetle genus Anaspis (Insecta: Coleoptera: Scraptiidae) from the Baltic and Rovno ambers (Upper Eocene of Eastern Europe). Paleontological Journal, 43 (9), 1092–1094. https://doi.org/10.1134/S0031030109090081
  33. Perkovsky, E.E., Legalov, A.A. & Háva, J. (2024) Trinodes puetzi Háva et Prokop (Coleoptera, Dermestidae, Trinodinae) from Klesov: first record for the Ukrainian Eocene. Ecologica Montenegrina, 73, 428–433. https://doi.org/10.37828/em.2024.73.28
  34. Polchaninova, N.Yu., Terekhova, V.V. & Drogvalenko, O.M. (2023) The first stage of the study of ground-dwelling arthropods of the main habitats of Rzhyshchiv CATC. In: Kuzemko, A., Kutsokon’, Yu. & Vasilyuk, O. (Eds), Biodiversity of Rzhyshchiv city amalgamated territorial community. Issue 2. Studies of “Hlyboki Balyky” Ecological Research Station, separate unit of Ukrainian ecostations network. Druk Art, Chernivtsi, pp. 225–247. [In Ukrainian with English abstract]
  35. Radchenko, A.G. & Perkovsky, E.E. (2021) Wheeler’s dilemma revisited: first Oecophylla-Lasius syninclusion and other ant syninclusions in the Bitterfeld amber (late Eocene). Invertebrate Zoology, 18 (1), 47–65. https://doi.org/10.15298/invertzool.18.1.05
  36. Shamshev, I.V., Perkovsky, E.E. & Legalov, A.A. (2025) A new species of hybotid dance fly of the genus Eodromyia (Diptera: Hybotidae, Tachydromiinae) from the late Eocene Rovno amber. Ecologica Montenegrina, 81, 14–21. https://doi.org/10.37828/em.2025.81.2
  37. Shavrin, A.V., Vasilenko, D.V. & Perkovsky, E.E. (2025) The first record of omaliine rove beetles (Coleoptera: Staphylinidae: Omaliinae) in Eocene Rovno amber. Palaeoentomology, 8 (2), 121–124. https://doi.org/10.11646/palaeoentomology.8.2.1
  38. Telnov, D, Perkovsky, E.E., Kundrata, R., Kairišs, K., Vasilenko, D.V. & Bukejs, A. (2023) Revealing Palaeogene distribution of the Ptilodactylidae (Insecta: Coleoptera): the first Ptilodactyla Illiger, 1807 records from Rovno amber of Ukraine. Historical Biology, 35 (11), 2171–2180. https://doi.org/10.1080/08912963.2022.2136034
  39. Tenenbaum, S. (1915) Fauna koleopterologiczna wysp Balearskich. (Faune coléoptérologique des ȋles Baléares). Skład główny w księgarni Gebethnera i Wolffa, Warszawa, 150 + iv pp.