Abstract
The oldest representative of the ceraphronoid family Megaspilidae is described and illustrated based on a well-preserved female from Early Cretaceous Lebanese amber. Alphaspilus heliades gen. et sp. nov. shows the characteristic anatomical features of the family and, despite superficial similarity, clearly differs from the genera Megaspilus Westwood and Conostigmus Dahlbom. This discovery proves that both stem and crown ceraphronoid groups coexisted in Early Cretaceous forest ecosystems. An updated checklist of all known fossil Megaspilidae species is provided.
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