Abstract
The rich late Oligocene fossil deposit of Enspel (Westerwald, Germany) provided two new species of soldier beetles (Cantharidae), namely Cantharis doernerorum sp. nov. and Cantharis zabolica sp. nov. Furthermore, C. doernerorum sp. nov. appears to have preserved traces of the original coloration, a character not found until now in soldier beetles preserved as compression fossils in rocks, and C. zabolica sp. nov. shows a metathoracic wing with veins. The new species are compared to those of the adjacent and almost contemporaneous Rott Lagerstätte (late Oligocene) and other fossiliferous deposits.
References
Barden, H.E., Bergmann, U., Edwards, N.P., Egerton, V.M., Manning, P.L., Perry, S., van Veelen, A., Wogelius, R.A. & van Dongen, B.E. (2015) Bacteria or melanosomes? A geochemical analysis of micro-bodies on a tadpole from the Oligocene Enspel Formation of Germany. Palaeobiodiversity and Palaeoenvironments, 95 (1), 33–45.
https://doi.org/10.1007/s12549-014-0177-5
Brancucci, M. (1980) Morphologie comparée, évolution et systématique des Cantharidae (Insecta: Coleoptera). Entomologica Basiliensia, 5, 215–388.
Comte, B. & Vianey-Liaud, M. (1987) Nouveaux Eomyidae du Paléogène d’Europe occidentale. Comptes Rendus de l’Académie des Sciences, Paris, Série II , 304 (15), 951–955.
Delkeskamp, K. (1977) Coleopterorum Catalogus Supplementa. Pars 165 (Fasc. I), Cantharidae. W. Junk, The Hague, 485 pp.
Fabricius, J.C. (1798) Supplementum entomologiae systematicae. Proft et Storch, Hafniae, [2] + 572 pp.
Fanti, F. (2014) Catalogo critico delle Cantharidae d’Italia (Insecta, Coleoptera). Memorie della Società Entomologica Italiana, 91 (1–2), 61–132.
https://doi.org/10.4081/MemorieSEI.2014.61
Fanti, F. (2017) Catalogo Cantharidae fossili del mondo. Fossils & Minerals Review, 2, 1–18 (abbreviated Italian version. Published: 12 March 2017) / World catalog of fossil Cantharidae. Fossils & Minerals Review, 2 (Special Issue), 1–52 (extended English version. Published: 25 May 2017).
Fanti, F. & Damgaard, A.L. (2018) Fossil soldier beetles from Baltic amber of the Anders Damgaard amber collection (Coleoptera Cantharidae). Baltic Journal of Coleopterology, 18 (1), 1–32.
Fanti, F. & Vitali, F. (2017) Key to fossil Malthininae, with description of two new species in Baltic amber (Coleoptera Cantharidae). Baltic Journal of Coleopterology, 17 (1), 19–27.
Felder, M., Weidenfeller, M. & Wuttke, M. (1998) Lithologische Beschreibung einer Forschungsbohrung im Zentrum des oberoligozänen, vulkano-lakustrinen Beckens von Enspel/Westerwald (Rheinland-Pfalz; Bundesrepublik Deutschland). Mainzer geowissenschaftliche Mitteilungen, 27, 101–136.
Gunkel, S. & Wappler, T. (2015) Plant-insect interactions in the upper Oligocene of Enspel (Westerwald, Germany), including an extended mathematical framework for rarefaction. Palaeobiodiversity and Palaeoenvironments, 95, 55–75.
https://doi.org/10.1007/s12549-014-0176-6
Heyden, C.H.G. von & Heyden, L.F.J.D. von (1866) Käfer und Polypen aus der Braunkohle des Siebengebirges. Palaeontographica, 15, 131–156 (+ p. 157: Dipteren-Larve aus dem Tertiär-Thon von NiederFlörsheim in Rhein-Hessen) + plates 22–24. [separatum under the title: Käfer und Polypen aus der Braunkohle des Siebengebirges. -Dipteren-Larve aus dem Tertiär-Thon von Nieder-Flörsheim in Rhein-Hessen. Besonderer Abdruck aus den Palaeontographicis, XV. Verlag von Theodor Fischer, Cassel, 1866, pp. 1–29 + plates 1–3].
Imhoff, L. (1856) Versuch einer Einführung in das Studium der Koleoptern. In zwei Theilen und einem, 25 Tafeln lithographirter Abbildungen nebst Text enthaltenden, Anhange. Schweighauser, Basel, xxxi + [2] + 114 + [2] + 272 pp. + 25 plates.
Kazantsev, S.V. (2018) New Baltic amber soldier beetles (Coleoptera, Cantharidae, Cantharinae). Euroasian Entomological Journal, 17 (2), 146–152.
Kӧhler, F. & Klausnitzer, B. (1998) Verzeichnis der Käfer Deutschlands. Entomologische Nachrichten und Berichte (Dresden), 4, 1–185.
Köhler, J. & Uhl, D. (2014) Die Blatt-und Karpoflora der oberoligozänen Fossillagerstätte Enspel (Westerwald, Rheinland-Pfalz, W-Deutschland). Mainzer Naturwissenschaftliches Archiv, Beiheft 35, 1–87.
Kuśka, A. (1992) Three new species of beetles (Coleoptera: Cantharidae, Anobiidae, Curculionidae) from the Baltic amber. Annals of the Upper Silesian Museum-Entomology, 3, 107–113.
Kuśka, A. (1996) New beetle species (Coleoptera: Cantharidae, Curculionidae) from the Baltic amber. Prace Muzeum Ziemi, 44, 13–18.
Leach, W.E. (1815) Entomology. In: Brewster, D. (Ed.), The Edinburgh Encyclopaedia; or dictionary of arts, sciences, and miscellaneous literature. Volume IX, part I. William Blackwood, Edinburgh, pp. 57–172.
Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii, Holmiae, 823 pp.
Magis, N. & Wittmer, W. (1974) Nouvelle répartition des genres de la sous-famille des Chauliognathinae (Coleoptera, Cantharoidea: Cantharidae). Bulletin de la Société Royale des Sciences de Liège, 43, 78–95.
McKey Fender, D. (1950) Notes on Cantharis III (Coleoptera: Cantharidae). The Pan-Pacific Entomologist, 26, 25–33, 61–79.
McNamara, M.E., Briggs, D.E.G. & Orr, P.J. (2012) The controls on the preservation of structural color in fossil insects. Palaios, 27 (7), 443–454.
https://doi.org/10.2110/palo.2012.p12-027r
Mertz. D.F., Renne, P.R., Wuttke, M. & Mödden, C. (2007) A numerically calibratedt reference level (MP28) for the terrestrial mammal-based biozonation of the European Upper Oligocene. International Journal of Earth Sciences (Geologische Rundschau), 96 (2), 353–361.
https://doi.org/10.1007/s00531-006-0094-6
Meunier, F.A. (1915) Über einige fossile Insekten aus den Braunkohlenschichten (Aquitanien) von Rott im Siebengebirge. 3. Teil. Zeitschrift der Deutschen Geologischen Gesellschaft (A), 67, 219–230, plates XXVI–XXVII.
Pirrung, M., Büchel, G. & Jacoby, W. (2001) The Tertiary volcanic basins of Eckfeld, Enspel and Messel (Germany). Zeitschrift der deutschen geologischen Gesellschaft, 152 (1), 27–59.
Poschmann, M., Schindler, T. & Uhl, D. (2010) Fossil-Lagerstätte Enspel—a short review of current knowledge, the fossil association, and a bibliography. Palaeobiodiversity and Palaeoenvironments, 90 (1), 3–20.
https://doi.org/10.1007/s12549-009-0017-1
Procházka, R. (1936) Studie k. morphologickému a systematickému významu nervatury křidel u Malacodermat. Etude sur l’importance morphomatique et systematique de la nervation des ailes des Malacodermata. Sborník Entomologického oddelění Národního Musea v Praze, 14, 100–132 + 2 plates.
Ramsdale, A.S. (2002) Family 64. Cantharidae Imhoff 1856. In: Arnett, R.H., Jr., Thomas, M.C., Skelley, P.E. & Frank, J.H. (Eds.), American Beetles, 2. Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press, Boca Raton, pp. 202–218.
Roček, Z. & Wuttke, M. (2010) Amphibia of Enspel (Late Oligocene, Germany). Palaeobiodiversity and Palaeoenvironments, 90 (4), 321–340.
https://doi.org/10.1007/s12549-010-0042-0
Sagegami-Oba, R., Takahashi, N. & Oba, Y. (2007) The evolutionary process of bioluminescence and aposematism in cantharoid beetles (Coleoptera: Elateroidea) inferred by the analysis of 18S ribosomal DNA. Gene, 400, 104–113.
https://doi.org/10.1016/j.gene.2007.06.004
Schindler, T. & Wuttke, M. (2010) Geology and limnology of the Enspel Formation (Chattian, Oligocene; Westerwald, Germany). Palaeobiodiversity and Palaeoenvironments, 90 (1), 21–27.
https://doi.org/10.1007/s12549-009-0014-4
Schindler, T. & Wuttke, M. (2015) A revised sedimentological model for the late Oligocene crater lake Enspel (Enspel Formation, Westerwald Mountains, Germany). Palaeobiodiversity and Palaeoenvironments, 95, 5–16.
https://doi.org/10.1007/s12549-014-0178-4
Storch, G., Engesser, B. & Wuttke, M. (1996) Oldest fossil record of gliding in rodents. Nature, 379 (6564), 439–441.
https://doi.org/10.1038/379439a0
Uhl, D. (2015) Preliminary note on fossil flowers and inflorescences from the late Oligocene of Enspel (Westerwald, W-Germany). Palaeobiodiversity and Palaeoenvironments, 95, 47–53.
https://doi.org/10.1007/s12549-014-0172-x
Uhl, D. & Poschmann, M. (2018) Groenlandia pescheri sp. nov. (Potamogetonaceae) from the Late Oligocene Fossil-Lagerstätte Enspel (Westerwald, Germany). Acta Palaeobotanica, 58 (1): 61–72.
https://doi.org/10.2478/acpa-2018-0001
Wedmann, S. (1998) Taphonomie der Bibionidae (Insecta: Diptera) aus der oberoligozänen Fossillagerstätte Enspel (Deutschland). Neues Jahrbuch für Geologie und Paläontologie Monatshefte, 1998 (9), 513–528.
Wedmann, S. (2000) Die Insekten der oberoligozänen Fossillagerstätte Enspel (Westerwald, Deutschland). Systematik, Biostratinomie und Paläoökologie. Mainzer Naturwissenschaftliches Archiv, Beiheft 23, 154 + 24 pp.
Wedmann, S., Poschmann, M. & Hörnschemeyer, T. (2010) Fossil insects from the Late Oligocene Enspel Lagerstätte and their palaeobiogeographic and palaeoclimatic significance. Palaeobiodiversity and Palaeoenvironments, 90 (1), 49–58.