Ptychopteridae is an ancient and rather diverse nematoceran family with three extant and 12 fossil genera subdivided into 80 extinct species belonging to five subfamilies (Eskov & Lukashevich, 2015; Lukashevich, 2019, 2020; Liu & Huang, 2020). Several undescribed ptychopterid larvae from the Upper Triassic of Germany represent the oldest record of the family (Barth et al., 2011). Crenoptychoptera Kalugina, 1985, an extinct genus belonging to the subfamily Eoptychopterinae Handlirsch, 1906, includes seven described species. All these species were recorded from the Lower Jurassic to the Lower Cretaceous of Eurasia: C. dobbertinensis Ansorge, 1998 from the Lower Jurassic of the Former Clay in Germany (Lukashevich et al., 1998), C. conspecta Lukashevich, 1995 from the Lower Jurassic of Ust-Baley in Russia (Lukashevich, 1995), the type species C. antica Kalugina, 1985 and C. defossa Kalugina, 1985 from the Middle Jurassic of the Kubekovo in Russia (Kalugina & Kovalev, 1985), C. bavarica Krzemiński & Ansorge, 1995 from the Upper Jurassic of Wegscheid in Germany (Krzemiński & Ansorge, 1995), C. liturata Lukashevich, 2011 from Upper Jurassic of Shar Teg in Mongolia, and C. gronskayae Kalugina, 1989 from the Upper Jurassic to Lower Cretaceous of Kempendyay in Russia (Kalugina, 1989; Lukashevich, 2011). Three species discovered from the Daohugou beds of China have been assigned to Crenoptychoptera (Hao et al., 2009), but a following study suggested that they should be placed in Axymyiidae (Zhang, 2010).
Barth, G., Ansorge, J. & Brauckmann, C. (2011) First record of the genus Ipsvicia (Hemiptera: Ipsviciidae) outside Gondwana—an Australian genus from the Upper Triassic of Germany. Polish Journal of Entomology, 80, 645–657.
Eskov, K.Y. & Lukashevich, E.D. (2015) On the history of ranges of two relict nematoceran families, Ptychopteridae and Tanyderidae (Insecta: Diptera): a biogeographical puzzle. Russian Entomological Journal, 24, 257–270.
Hao, J.Y., Dong, K.Q. & Ren. D. (2009) Middle Jurassic Eoptychopteridae from Daohugou, Inner Mongolia, China (Insecta, Diptera, Eoptychopteridae). Acta Zootaxonomica Sinica, 34, 106–110 [In Chinese].
Huang, D.Y. (2019) Jurassic integrative stratigraphy and timescale of China. Science in China: Earth Science, 662, 223–255.
Huang, D.Y., Fu, Y.Z., Gao, J. & Nel, A. (2018) A new damsel-dragonfly of the small family Selenothemistidae from the earliest Late Jurassic of China (Odonata: Isophlebioptera). Palaeoentomology, 1 (1), 37–41.
Kalugina, N.S. & Kovalev, V.G. (1985) [Dipterous insects of Jurassic Siberia.] Paleontological Institute, Moscow, 1–198. [In Russian]
Krzemiński, W. & Ansorge, J. (1995) New Upper Jurassic Diptera (Limoniidae, Eoptychopteridae) from the Solnhofen Lithographic Limestone (Bavaria, Germany). Stuttgarter Beiträge zur Naturkunde (B), 221, 1–7.
Liu, Y.M. & Huang, D.Y. (2020) New materials of trichocerid and ptychopterid dipterans from the earliest Late Jurassic of Jiyuan Basin, China. Palaeoentomology, 3 (5), 461–465.
Lukashevich, E.D. (1995) First pupae of the Eoptychopteridae and Ptychopteridae from the Mesozoic of Siberia (Insecta: Diptera). Paleontological Journal, 29, 164–171.
Lukashevich, E.D. (2019) Review of the fossil record of Bittacomorphinae (Diptera: Ptychopteridae). Zootaxa, 4661 (3): 566–578.
Lukashevich, E.D. (2020) Rare nematoceran dipterans (Insecta: Diptera) from the Khasurty Locality, Transbaikalia. Paleontological Journal, 54 (6): 600–612.
Lukashevich, E.D., Ansorge, J., Krzemiński, W. & Krzemińska, E. (1998) Revision of Eoptychopterinae (Diptera: Eoptychopteridae). Polskie Pismo Entomologiczne, 67, 311–343.
Lukashevich, E.D. (2011) New nematocerans (Insecta: Diptera) from the Late Jurassic of Mongolia. Paleontological Journal, 45, 620–628.
Lukashevich, E.D. (2012) Phylogeny of Ptychopteroidea (Insecta: Diptera). Paleontological Journal, 46, 476–484.
Zhang, J.F. (2010) Two new genera and one new species of Jurassic Axymyiidae (Diptera: Nematocera), with revision and redescription of the extinct taxa. Annals of the Entomological Society of America, 103, 455–464.