Skip to main content Skip to main navigation menu Skip to site footer
Short Communication
Published: 2021-09-24

Re-description of the Cretaceous wasp Humiryssus leucus Lin, 1980 (Hymenoptera: Evanioidea: Baissidae)

State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China
Hymenoptera Evanioidea Baissidae


Humiryssus Lin, 1980 is a Cretaceous genus with Humiryssus leucus Lin, 1980 as its type species established based on a tiny wasp from the Lower Cretaceous (Hauterivian-Barremian) Laocun Formation at Laocun, Jiande, Zhejiang Province, eastern China and originally placed in the extinct family Paroryssidae (Hymenoptera: Tenthredinoidea) (Lin, 1980). It was later treated jointly with the genus Manlaya Rasnitsyn, 1980 (Evanioidea: Gasteruptiidae: Baissinae) because the characters shown in the line drawing by Lin (1980) indicate a close relationship of the genus to Manlaya (Rasnitsyn et al., 1998). However, Zhang & Rasnitsyn (2004) considered it to be a separate genus within the Baissinae, which is now considered as an independent family under the Evanioidea (e.g., Li et al., 2018; Jouault et al., 2020, 2021).


  1. Engel, M.S. (2013) A new genus and species of Baissidae in Late Cretaceous amber from New Jersey (Hymenoptera: Evanioidea). Novitates Paleoentomologicae, 3, 1–8.

  2. Hong, Y.C. & Wang, W.L. (1990) Fossil insects from the Laiyang Basin, Shandong Province. In: The Regional Geological Surveying Team, Shandong Bureau of Geology and Mineral Resources (Eds). The stratigraphy and paleontology of Laiyang Basin, Shandong Province. Beijing: Geological Publishing House, pp. 157–158 [In Chinese with English Summary].

  3. Jouault, C., Nel, A. & Perrichot, V. (2020) New evanioid wasps (Hymenoptera: Praeaulacidae, Aulacidae) from Cenomanian Burmese amber. Cretaceous Research, 110, 104407.

  4. Jouault, C., Maréchal, A., Condamine, F.L., Wang, B., Legendre, F. & Perrichot, V. (2021) Including fossils in phylogeny: a glimpse into the evolution of the superfamily Evanioidea (Hymenoptera: Apocrita) under tip-dating and the fossilized birth–death process. Zoological Journal of the Linnean Society (in press).

  5. Li, L.F., Rasnitsyn, A.P., Shih, C.K., Labandeira, C.C., Buffington, M., Li, D.Q. & Ren, D. (2018) Phylogeny of Evanioidea (Hymenoptera, Apocrita), with descriptions of new Mesozoic species from China and Myanmar. Systematic Entomology, 43, 810e842.

  6. Lin, Q.B. (1980) Mesozoic insects from Zhejiang and Anhui provinces. In: Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences (Ed.). Division and correlation of the Mesozoic volcano-sedimentary strata in Zhejiang and Anhui Provinces. Beijing: Science Press, pp. 211–234 [In Chinese].

  7. Rasnitsyn, A.P. (1975) Hymenoptera Apocrita of the Mesozoic. Transactions of the Paleontological Institute, Academy of Sciences of the USSR, 147, 1–134 [In Russian].

  8. Rasnitsyn, A.P. (1980a) Origin and evolution of the Hymenoptera. Transactions of the Paleontological Institute, Academy of Sciences of the USSR, 174, 1–192 [In Russian].

  9. Rasnitsyn, A.P. (1980b) On system of the family Aulacidae (Hymenoptera) in connection with a new finding in the Cretaceous of Manlay. In: Kalugina, N.S. (Ed.), The Early Cretaceous Lake Manlay. Transactions of the Joint Soviet-Mongolian Paleontological Expedition, 13, 65–67 [In Russian].

  10. Rasnitsyn, A.P. (1986) Vespida (= Hymenoptera). In: Rasnitsyn, A.P. (Ed.), Insects in the Early Cretaceous Ecosystems of West Mongolia. Transactions of the Joint Soviet-Mongolian Paleontological Expedition, 28, 154–164 [In Russian].

  11. Rasnitsyn, A.P., Jarzembowski, E.A. & Ross, A.J. (1998) Wasps (Insecta: Vespida = Hymenoptera) from the Purbeck and Wealden (Lower Cretaceous) of southern England and their biostratigraphical and palaeoenvironmental significance. Cretaceous Research, 19, 329–391.

  12. Zhang, J.F. & Rasnitsyn, A.P. (2004) Minute members of Baissinae (Insecta: Hymenoptera: Gasteruptiidae) from the Upper Mesozoic of China and limits of the genus Manlaya Rasnitsyn, 1980. Cretaceous Research, 25, 797–805.