Abstract
The Early Cretaceous genus Mesolygaeus Ping, 1928 is revised based on numerous specimens from the Lower Cretaceous Laiyang Formation in the Laiyang Basin, eastern Shandong Province, China. The genus Enicocoris Popov, 1980 is recognized as a junior synonym of Mesolygaeus syn. nov. and the family Mesolygaeidae, 1990 is therefore recognized as a junior synonym of Enicocoridae Popov, 1980 syn. nov. Also, the genus Longyanteclypea Zhang et al., 2013 is considered as a junior synonym of Mongolocoris Ryzhkova, 2012 syn. nov. Additionally, a new species, Mongolocoris polhemi Ryzhkova sp. nov., is described from the Lower Cretaceous Tsagaantsab Formation of Tsagaan-Suburga, Dornogovi Province, southeastern Mongolia.
References
- Grabau, A.W. (1923) Cretaceous fossils from Shantung. Bulletin of the Geological Survey of China, 5, 164–181.
- Hong, Y.C. (1976) Insecta. In: Tianjin Institute of Geology and Mineral Resourses (Ed.), Paleontological atlas of North China Region: Inner Mongolia. Vol. 2. Geological Publishing House, Beijing, 81–87. [In Chinese]
- Hong, Y.C. (1981) Discovery of new Early Cretaceous insects from Xishan, Beijing. Bulletin of the Tianjin Institute of Geology and Mineral Resourses, 4, 87–94. [In Chinese]
- Hong, Y.C. (1982) Mesozoic fossil insects of Jiuquan basin in Gansu province. Geological Publishing House, Beijing, 187 pp. [In Chinese]
- Hong, Y.C. (1984) New fossil insects of Laiyang Group from Laiyang basin, Shandong Province. Professional Papers on Stratigraphy and Palaeontology, 11, 31–41. [In Chinese]
- Hong, Y.C. (1988) New fossil insects of Lenshuiwu Formation, northeastern Jiangxi. Professional Papers of Stratigraphy and Palaeontology, 21, 172–178. [In Chinese]
- Hong, Y.C. (1995a) Fossil insects of the Southern Ordos Basin. Acta Geologica Gansu, 4 (1), 1–13. [In Chinese]
- Hong, Y.C. (1995b) Fossil insects of the Northern Ordos Basin. Acta Geologica Gansu, 4 (2), 1–9. [In Chinese]
- Hong, Y.C. & Wang, W. (1990) Insects of Laiyang Formation. In: The stratigraphy and paleontology of Laiyang Basin, Shandong. Geology Publishing House, Beijing, 89–105. [In Chinese]
- Hong, Y.C. & Ren, D. (1992) Supplement characteristics of the family Mesolygaeidae Hong et Ren, 1990. Memoirs of Beijing Natural History Museum, 51 (3), 1–53.
- Huang, D.Y., Cai, C.Y. & Chen, A.L. (2012) The homonymy of Xishania with reference to Xishania fusiformis Hong, 1981 and X. longiusula Hu, 2002. Acta Palaeontologica Sinica, 51 (1), 102–106. [In Chinese with English abstract]
- International Code of Zoological Nomenclature (1999) Available from: https://www.iczn.org/the-code/the-code-online/ (Accessed 12 January, 2025).
- Lin, Q.B. (1976) The Jurassic fossil insects from western Liaoning. Acta Palaeontologica Sinica, 15 (1), 97–116. [In Chinese]
- Lin, Q.B. (1982a) Insecta. In: Nanjing Institute of Geology and Mineral Resourses (Ed.), Paleontological atlas of East China. part III, vol. 1 (Volume of Mesozoic and Cenozoic). Geological Publishing House, Beijing, pp. 148–155. [In Chinese]
- Lin, Q.B. (1982b) Class Insecta. In: Xi’an Institute of Geology and Mineral Resourses (Ed.), Palaeontological atlas of Northwest China, Shaanxi-Gansu-Ningxia volume. part II, Mesozoic and Cenozoic. Geological Publishing House, Beijing, pp. 70–83. [In Chinese]
- Lin, Q.B. (1983) Cretaceous succession of insect assemblages in China. Zitteliana, 10, 393–349.
- Ping, C. (1928) Cretaceous fossils from Shantung. Palaeontologia Sinica Series B, 13 (1), 1–56.
- Ponomarenko, A.G. (2007) Evolution of continental aquatic ecosystems. In: Problems of aquatic entomology of Russia and adjacent countries. Voronezh State University, Voronezh, pp. 228–259. [In Russian]
- Popov, Yu.A. (1980) Heteroptera from the Lower Cretaceous deposits of the Manlay locality. In: Early Cretaceous Lake Manlay. Proceedings of joint Soviet-Mongolian paleontological expedition. Vol. 13. Nauka, Moscow, 48–51. [In Russian]
- Popov, Yu.A. (1986) Peloridiidae and Bugs: Suborders Peloridiina (= Coleorrhyncha) et Cimicina (= Heteroptera). In: Insects in the Early Cretaceous ecosystems of Western Mongolia. Proceedings of Joint Soviet-Mongolian Paleontological Expedition. Vol. 28. Nauka, Moscow, 50–84. [In Russian]
- Popov, Yu.A. (1988) New Mesozoic Coleorrhyncha and Heteroptera (Hemiptera) of Eastern Transbaikalia. Paleontological Journal, 4, 67–77. [In Russian]
- Popov, Yu.A., Dolling, W.R. & Whalley, P.E.S. (1994) British Upper Triassic and Lower Jurassic Heteroptera and Coleorrhyncha (Insecta: Hemiptera). Genus, 5 (4), 307–347.
- Ryzhkova, O.V. (2012) New saldoid bugs of the family Enicocoridae (Hemiptera: Heteroptera: Leptopodomorpha) from the Lower Cretaceous of Mongolia. Paleontological Journal, 46, 485–494. https://doi.org/10.1134/S0031030112050085
- Ryzhkova, O.V. (2013) Mesozoic stage of the historical development of bugs of the infraorder Leptopodomorpha (Insecta, Hemiptera, Heteroptera) of Siberia and Mongolia. PhD in Biology thesis. Moscow, 168 pp. [In Russian]
- Shcherbakov, D.E. & Popov, Y.A. (2002) Superorder Cimicida Laicharting, 1781, order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. In: Rasnitsyn, A.P. & Quicke, D.L. (Eds), History of insects. Kluwer Academic Publishers, Dordrecht, pp. 143–157.
- Schuh, R.T. & Weirauch, C. (2020) True bugs of the world (Hemiptera: Heteroptera): Classification and natural history. Second ed. Siri Scientific Press, Rochdell, 800 pp.
- Wang, W.L. (1980) Insecta. In: Shenyang Institute of Geology and Minerals (Ed.), Paleontological atlas of Northeast China. II. Mesozoic and Cenozoic. Geological Publishing House, Beijing, pp. 130–153. [In Chinese]
- Wootton, R.J. & Betts, C.R. (1986) Homology and function in the wings of Heteroptera. Systematic Entomology, 11, 389–400. https://doi.org/10.1111/j.1365-3113.1986.tb00191.x
- Zhang, J.F. (1986) Some fossil insects from the Jurassic of northern Hebei, China. In: The Paleontology and Stratigraphy of Shandong. Ocean Press, Beijing, pp. 74–84. [In Chinese]
- Zhang, J.F. (1989) Discussion of the genus Mesolygaeus Ping. Abstracts of 15th Annual Symposium of the Palaeontological Society of China. pp. 2–3.
- Zhang, J.F. (1991) Going further into Late Mesozoic mesolygaeids (Heteroptera, Insecta). Acta Palaeontologica Sinica, 30 (6), 679–704.
- Zhang, J.F. (1992) Late Mesozoic entomofauna from Laiyang, Shandong Province, China, with discussion of its palaeoecological and stratigraphical significance. Cretaceous Research, 13 (2), 133–145. https://doi.org/10.1016/0195-6671(92)90031-K
- Zhang, J.F. (1993) A contribution to the knowledge of insects from the Late Mesozoic in Southern Shaanxi and Henan provinces, China. Palaeoworld, 2, 49–56.
- Zhang, J.F., Golub, V.B., Popov, Y.A. & Shcherbakov, D.E. (2005) Ignotingidae fam. nov. (Insecta: Heteroptera: Tingoidea), the earliest lace bugs from the upper Mesozoic of eastern China. Cretaceous Research, 26 (5), 783–792. https://doi.org/10.1016/j.cretres.2005.05.004
- Zhang, W.T., Engel, M.S., Yao, Y.Z., Ren, D. & Shih, C.K. (2013) The Mesozoic family Archegocimicidae and phylogeny of the infraorder Leptopodomorpha (Hemiptera). Journal of Systematic Palaeontology, 12 (1), 93–111. https://doi.org/10.1080/14772019.2012.744109
