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Type: Correspondence
Published: 2026-02-25
Page range: 5-8
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New Sycoracinae (Diptera: Psychodidae) from the Early Cretaceous Lebanese amber

Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China; University of Chinese Academy of Sciences, Beijing 100049, China
Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China; University of Chinese Academy of Sciences, Beijing 100049, China
Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China; Lebanese University, Faculty of Sciences II, Natural Sciences Department, Fanar - El-Matn, PO box 26110217, Lebanon
Diptera Psychodidae

Abstract

Lebanese amber provides one of the most significant windows to terrestrial arthropod diversity of the Early Cretaceous (Azar, 1997; Maksoud et al., 2022). Dated to approximately ~125 Ma (lower Barremian), these amber deposits have yielded an exceptionally rich and often very well preserved palaeofauna (Ross, 2010; Granier et al., 2016; Maksoud et al., 2017). To date, about 272 insect species in 108 families have been formally described from Lebanese amber, representing a broad spectrum of Lower Cretaceous entomofaunal diversity (Maksoud & Tabakian, 2025).

References

  1. Azar, D. (1997) Lebanese amber. Meganeura, 1, 26–27.
  2. Azar, D., Azar, S. & Maksoud, S. (2018) Libanosycorax dimyi gen. et sp. nov., the earliest sycoracine fly from the Lower Cretaceous amber of Lebanon (Diptera: Psychodidae: Sycoracinae). Palaeoentomology, 1 (1), 42–46. https://doi.org/10.11646/palaeoentomology.1.1.5
  3. Azar, D., Nel, A. & Perkovsky, E. (2013) A new Sycorax species from Eocene Ukrainian Rovno amber (Diptera: Psychodida: Sycoracinae). In: Azar, D., Engel, M.S., Jarzembowski, E., Krogmann, L. & Santiago-Blay J. (Eds), Insect evolution in an amberiferous and stone alphabet. Brill, Leiden, pp. 27–46. https://doi.org/10.1163/9789004210714
  4. Azar, D., Perrichot, V., Néraudeau, D. & Nel, A. (2003) New Psychodids from the Cretaceous ambers of Lebanon and France, with a discussion of Eophlebotomus connectens Cockerell, 1920 (Diptera, Psychodidae). Annals of the Entomological Society of America, 96 (2), 117–126. https://doi.org/10.1603/0013-8746(2003)096[0117:NPFTCA]2.0.CO;2
  5. Azar, D. & Salamé, Y. (2015) A new genus of Sycoracinae (Diptera: Psychodidae) from upper Cretaceous amber of New Jersey. Cretaceous Research, 52, 539–547.
  6. http://doi.org/10.1016/j.cretres.2014.05.002
  7. Azar, D., Tahchy, A. & Perrichot, V. (2007) The oldest Sycoracinae (Diptera: Psychodidae) from the French Cretaceous amber. Alavesia, 1, 5–10.
  8. Bejarano, E.E., Duque, P. & Vélez, I.D. (2008) A new species of Sycorax (Diptera, Psychodidae, Sycoracinae) from the Pacific coast of Colombia. Revista Brasileira de Entomologia, 52 (1), 28–31. https://doi.org/10.1590/S0085-56262008000100005
  9. Bravo, F., Araújo, M.X. & Vilarinho, N. (2023) A new genus and new species of Sycoracinae (Diptera: Psychodidae) from the Neotropical region with keys to the extant genera of the subfamily and males of Neotropical species of Sycorax. Neotropical Entomology, 52, 697–708. https://doi.org/10.1007/s13744-023-01040-6
  10. Cordeiro, D.P. & Wagner, R. (2018) Family Psychodidae. In: Hamada, N., Thorp, J.H. & Rogers, D.C. (Eds), Thorp and Covich’s freshwater invertebrates, Fourth Edition. Volume 3: Keys to Neotropical Hexapoda. Academic Press, New York, 765–770. https://doi.org/10.1016/B978-0-12-804223-6.00039-1
  11. Granier, B.R.C., Toland, C., Gèze, R., Azar, D. & Maksoud, S. (2016) Some steps toward a new story for the Jurassic—Cretaceous transition in Mount Lebanon. Carnets de Géologie, 16 (8), 247–269. https://doi.org/10.4267/2042/59924
  12. Kvifte, G.M. & Wagner, R. (2017) Psychodidae (sand flies, moth flies or owl flies). In: Kirk-Spriggs, A.H. & Sinclair, B.J. (Eds), Manual of Afrotropical Diptera. Nematocerous Diptera and lower Brachycera. Suricata, 5. South African National Biodiversity Institute, Pretoria, pp. 607–632.
  13. Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio Decima, Reformata. Laurentii Salvii, Holmiae, [4] + 823 + [1] pp. https://doi.org/10.5962/bhl.title.542
  14. Maksoud, S., Azar, D., Granier, B.R.C. & Gèze, R. (2017) New data on the age of the Lower Cretaceous amber outcrops of Lebanon. Palaeoworld, 26 (2), 331–338. https://doi.org/10.1016/j.palwor.2016.03.003
  15. Maksoud, S., Granier, B.R.C. & Azar, D. (2022) Palaeoentomological (fossil insects) outcrops in Lebanon. Carnets de Géologie, 22 (16), 699–743. https://doi.org/10.2110/carnets.2022.2216
  16. Maksoud, S. & Tabakian, H. (2025) Lebanese amber: A time capsule from the dawn of modern ecosystems. Palaeoworld, 34 (6), 201015. https://doi.org/10.1016/j.palwor.2025.201015
  17. Meunier, F. (1905) Monographie des Psychodidae de l’ambre de la Baltique. Annales Historico-Naturales Musei Nationalis Hungarici, 3, 235–255.
  18. Newman, E. (1834) Attempted division of British insects into natural orders. Entomological Magazine, 2, 379–431.
  19. Petrulevičius, J., Salas-Gismondi, R., Flynn, J., Nel, A., Antoine, P.O. & De Franceschi, D. (2011) First fossil blood sucking Psychodidae in South America: a sycoracine moth fly (Insecta: Diptera) in the middle Miocene Amazonian amber. Insect Systematics & Evolution, 42 (1), 87–96. https://doi.org/10.1163/187631211X560919
  20. Rondani, C. (1856) Dipterologiae Italicae. Vol. 1. Genera italica ordinis dipterorum ordinatim disposita et distincta et in familias et stirpes aggregata. Tipographia Alexandri Stoschi, Parma, 228 pp. https://doi.org/10.5962/bhl.title.8160
  21. Ross, A.J. (2010) Amber: the natural time capsule. Natural History Museum, London, 112 pp.
  22. Stebner, F., Solórzano-Kraemer, M.M., Ibáñez-Bernal, S. & Wagner, R. (2015) Moth flies and sand flies (Diptera: Psychodidae) in Cretaceous Burmese amber. PeerJ, 3 (1254), 34. https://doi.org/10.7717/peerj.1254
  23. Tabakian, H. (2025) Psychodid flies (Diptera: Psychodidae) from the Lower Cretaceous Lebanese amber. Palaeoworld, 34 (6), 201014. https://doi.org/10.1016/j.palwor.2025.201014
  24. Wagner, R., Agnihotri, P. & Singh, H. (2022) A new species of Sycorax (Sycoracinae: Psychodidae) from the Lower Eocene amber of Tadkeshwar, Gujarat, India. Palaeoentomology, 5 (4), 319–326. https://doi.org/10.11646/palaeoentomology.5.4.4