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Type: Correspondence
Published: 2026-04-29
Page range: 121-124
Abstract views: 89
PDF downloaded: 2

First record of the genus Minyohelea (Diptera: Ceratopogonidae) from the Lower Cretaceous amber of Spain with description of a new species

Department of Invertebrate Zoology and Parasitology, University of Gdańsk, ul. Wita Stwosza 59, 80-308, Gdańsk, Poland
Departamento de Biodiversidad, Ecología y Evolución, Facultad de Biología, Universidad Complutense, 28040 Madrid, Spain
Nematocera Cretaceous amber inclusion fossil synonymy

Abstract

Fossil Ceratopogonidae are common in Spanish amber. To date eight species have been described (Peñalver et al., 2025); two of them have been described from San Just amber: Atriculicoides hispanicus (Szadziewski & Arillo, 2016) and Atriculicoides sanjusti (Szadziewski & Arillo, 2016), both in the same piece as the new species. Gerontodacus skalskii (Szadziewski & Arillo, 1998), originally described from Álava amber is also recorded in San Just amber (Arillo et al., 2008).

References

  1. Arillo, A., Peñalver, E. & Delclòs, X. (2008) Microphorites (Diptera: Dolichopodidae) from the Lower Cretaceous amber of San Just (Spain), and the co-occurrence of two ceratopogonid species in Spanish amber deposits. Zootaxa, 1920 (1), 29–40. https://doi.org//10.11646/zootaxa.1920.1.2
  2. Barrón, E., Peyrot, D., Bueno-Cebollada, C.A., Kvaček, J., Álvarez-Parra, S., Altolaguirre, Y. & Meléndez, N. (2023) Biodiversity of ecosystems in an arid setting: The late Albian plant communities and associated biota from eastern Iberia. PloS ONE, 18 (3), e0282178. https://doi.org//10.1371/journal.pone.0282178
  3. Borkent, A. (1995) Biting midges in the Cretaceous amber of North America (Diptera: Ceratopogonidae). Backhuys Publishers, Leiden, 237 pp.
  4. Borkent, A. (1997) Upper and Lower Cretaceous biting midges (Ceratopogonidae: Diptera) from Hungarian and Austrian amber and the Koonwarra Fossil Bed of Australia. Stuttgarter Beiträge zur Naturkunde, Serie B, 249, 1–10.
  5. Borkent, A. (2000) Biting midges (Ceratopogonidae: Diptera) from Lower Cretaceous Lebanese amber with a discussion of the diversity and patterns found in other ambers. In: Grimaldi, D. (Ed.), Studies on fossils in amber, with particular reference to the Cretaceous of New Jersey. Backhuys Publishers, Leiden, pp. 355–451.
  6. Corral, J.C., López del Valle, R. & Alonso, J. (1999) El ámbar cretácico de Álava (Cuenca Vasco-Cantábrica, norte de España). Su colecta y preparación. Estudios del Museo de Ciencias Naturales de Álava, 14 (Special Publication, number 2), 7–21. https://doi.org//10.33349/1999.29.914
  7. Hennig, W. (1948) Die Larvenformen der Dipteren: eine Übersicht über die bisher bekannten Jungendstadien der zweiflügeligen Insekten, 1. Akademie-Verlag, Berlin, 184 pp. https://doi.org//10.1515/9783112766613
  8. Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum caracteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. L. Salvii, Holmiae [= Stockholm], 824 pp. https://doi.org//10.5962/bhl.title.542
  9. Nascimbene, P. & Silverstein, H. (2000) The preparation of fragile Cretaceous ambers for conservation and study of organismal inclusions. In: Grimaldi, D. (Ed.), Studies on fossils in amber, with particular reference to the Cretaceous of New Jersey. Backhuys Publishers, Leiden, pp. 93–102.
  10. Newman, E. (1834) Attempted division of British insects into natural orders. Entomological Magazine, 2, 379–431.
  11. Noè, G. (1907) Due nuove specie di ditteri appartenenti ad un genere nuovo. Archivio Zoologico Italiano: Pubblicato Sotto gli Auspicii Della Unione Zoologica, 3, 101–163, 2 pls.
  12. Peñalver, E., Arillo, A., López Del Valle, R. & Delclòs, X. (2025) Cretaceous Spanish amber: History, research and checklist of taxa. Palaeoentomology, 8 (2), 195–234. https://doi.org//10.11646/palaeoentomology.8.2.10
  13. Peñalver, E., Delclòs, X. & Soriano, C. (2007) A new rich amber outcrop with palaeobiological inclusions in the Lower Cretaceous of Spain. Cretaceous Research, 28 (5), 791–802. https://doi.org//10.1016/j.cretres.2006.12.004
  14. Pielowska-Ceranowska, A., Azar, D. & Szwedo, J. (2021) New fossil biting midge from the Lower Cretaceous amber of Lebanon (Diptera: Ceratopogonidae). Cretaceous Research, 129, 105004, 1–9. [2022] https://doi.org//10.1016/j.cretres.2021.105004
  15. Pielowska-Ceranowska, A., Azar, D. & Szwedo, J. (2022) Redescription of Minyohelea nexuosa (Diptera: Ceratopogonidae) based on a new specimen from Lebanese amber. Palaeoentomology, 5 (2), 137–141. https://doi.org//10.11646/palaeoentomology.5.2.5
  16. Szadziewski, R. (1996) Biting midges from Lower Cretaceous amber of Lebanon and Upper Cretaceous Siberian amber of Taimyr (Diptera, Ceratopogonidae). Studia Dipterologica, 3, 23–86.
  17. Szadziewski, R. & Arillo, A. (1998) Biting midges (Diptera: Ceratopogonidae) from the Lower Cretaceous amber from Álava, Spain. Polskie Pismo Entomologiczne, 67, 291–298.
  18. Szadziewski, R., Arillo, A., Urbanek, A. & Sontag, E. (2016) Biting midges of the extinct genus Protoculicoides Boesel from Lower Cretaceous amber of San Just, Spain and new synonymy in recently described fossil genera (Diptera: Ceratopogonidae). Cretaceous Research, 58, 1–9. https://doi.org//10.1016/j.cretres.2015.09.016
  19. Villanueva-Amadoz, U., Pons, D., Diez, J.B., Ferrer, J. & Sender, L.M. (2010) Angiosperm pollen grains of San Just site (Escucha Formation) from the Albian of the Iberian Range (north-eastern Spain). Review of Palaeobotany and Palynology, 162, 362–381. https://doi.org//10.1016/j.revpalbo.2010.02.014