Abstract
Anatopynia vittigera Edwards is transfered to Alotanypus. The male and female of A. vittigera comb. nov. are redescribed and immatures are described and illustrated. A cladistic analysis including one species of each Macropelopiini genus was conducted in order to assess the phylogenetic position of Alotanypus and to provide the first phylogenetic hypothesis for the genus. Adults and immatures were included in the analysis where discrete and continuos characters were considered. The cladistic analysis demonstrated that Alotanypus is a monophyletic genus, with Guassutanypus oliveirai as the sister group.References
Almeida, M.T. & Bisby, F.A. (1984) A simple method for establishing taxonomic characters from measurements data. Taxon, 33, 405–409.
Archie, J.W. (1985) Methods for coding variable morphological features for numerical taxonomic analysis. Systematic Zoology, 34, 326–345.
Colless, D.H. (1980) Congruence between morphometric and allozyme data for Menidia species: a reappraisal. Systematic Zoology, 29, 288–299.
Cranston, P.S. & Humphries, C.J. (1988) Cladistics and computers: A chironomid conundrum? Cladistics, 4, 72–92.
Donato, M., Massaferro, J. & Brooks S.J. (2008) Chironomid (Chironomidae: Diptera) checklist from Nahuel Huapi National Park, Patagonia, Argentina. Revista de la Sociedad Entomológica Argentina, 67, 163 –170.
Edwards, F.W. (1931) Chironomidae. In: Diptera of patagonia and South Chile. Part II, Nematocera. Trustees of the British Museum (Natural History), London, pp. 233–331.
Epler, J.H. (2001) Identification Manual for the larval Chironomidae (Diptera) of North and South Carolina. A guide to the taxonomy of the midges of the southeastern United States, including Florida. Special Publication SJ2001-SP13. North Carolina Department of Environment and Natural Resources, Raleigh, NC, and St. Johns River Water Management District, Palatka, FL. 526 pp. http://home.earthlink.net/~johnepler/index.html (accessed: 20 November 2010)
Farris, J. (1970) Methods for computing Wagner trees. Systematic Zoology, 19, 83–92.
Farris, J. (1990) Phenetics in camouflage. Cladistics, 6, 91–100.
Fittkau, E.J. (1962) Die Tanypodinae (Diptera: Chironomidae) (Die Tribus Anatopyniini, Macropelopiini und Pentaneurini). Abhandlungen zur Larvalsystematik der Insekten, 6, 1–453.
Fittkau, E.J. & Roback, S.S. (1983) The larvae of Tanypodinae (Diptera: Chironomidae) of the Holarctic region - Keys and diagnoses In: Wiederholm, T. (ed.), Chironomidae of the Holarctic region - Keys and diagnoses. Part 1. Larvae. Entomologica scandinavica, Supplement, 19, 33–110.
Fittkau, E.J. & Murray, D.A. (1986) The pupae of Tanypodinae (Diptera: Chironomidae) of the Holarctic region - Keys and Diagnoses In: Wiederholm, T. (ed.), Chironomidae of the Holarctic region - Keys and diagnoses. Part 2. Pupae. Entomologica scandinavica, Supplement, 28, 31–117.
Goloboff, P.A. (1993) Estimating character weights during tree search. Cladistics, 9, 83–91.
Goloboff, P.A., Carpenter, J.M., Arias, J.S. & Miranda Esquivel, D.R. (2008b) Weighting against homoplasy improves phylogenetic analysis of morphological data sets. Cladistics, 24, 758–773.
Goloboff, P.A., Farris, J., Källersjö, M., Oxelmann, B, Ramirez, M. & Szumik, C. (2003) Improvements to resampling measures of group support. Cladistics, 19, 324–332.
Goloboff, P.A., Farris, J.S. & Nixon, K.C. (2008a) TNT, a free program for phylogenetic analysis. Cladistics, 24, 1–13.
Goloboff, P.A., Mattoni, C.I. & Quinteros, A.S. (2006) Continuous characters analyzed as such. Cladistics, 22, 589–601.
Niitsuma, H. (2005) Taxonomic and distribution of Alotanypus kuroberobustus comb. nov. (Insecta: Diptera: Chironomidae) from the Paleartic region. Species Diversity, 10, 135–144.
Mickevich, M.F. & Johnson, M.F. (1976) Congruence between morphological and allozyme data in evolutionary inference and character evolution. Systematic Zoology, 25, 260–270.
Murray, D.A. & Fittkau, E.J. (1989) The adult males of Tanypodinae (Diptera: Chironomidae) of the Holarctic region - keys and diagnoses. In: Wiederholm, T. (ed.), Chironomidae of the Holarctic region - Keys and diagnoses. Part 3. Adult males. Entomologica scandinavica, Supplement, 19, 37–123.
Pimentel, R.A. & Riggins, R. (1987) The nature of cladistic data. Cladistics, 3, 201–209.
Rae, T.C. (1998) The logical basis for the use of continuous characters in phylogenetic systematics. Cladistics, 14, 221–228.
Raffaele, E. (1996) Relationships between seed and spore banks and vegetation of a mountain flood meadow (mallín) in Patagonia, Argentina. Wetlands, 16, 1–9.
Rieradevall, M. & Brooks, S.J. (2001) An identification guide to subfossil Tanypodinae larvae (Insecta: Diptera: Chironomidae) based on cephalic setation. Journal of Paleolimnology, 25, 81–99.
Roback, S.S. (1971) The adults of the subfamily Tanypodinae (= Pelopiinae) in North America (Diptera: Chironomidae). Monograph of the Academy of Natural Sciences of Philadelphia, 17, 1–419.
Roback, S.S. (1976) The Immature Chironomids of the Eastern United States I. Introduction and Tanypodinae-Coelotanypodini. Proceedings of The Academy of Natural Sciences, 127, 147–201.
Roback, S.S. (1977) The immature chironomids of the eastern United States. II. Tanypodinae-Tanypodini. Proceedings of The Academy of Natural Sciences, 128, 55–87.
Roback, S.S. (1978) The immature chironomids of the eastern United States III. Tanypodinae-Anatopyniini, Macropelopiini and Natarsiini. Proceedings of The Academy of Natural Sciences, 129, 151–202.
Roback, S.S. & Moss, W.W. (1978) Numerical taxonomic studies on the congruence of classifications for the genera and subgenera of Macropelopiini and Anatopyniini (Diptera: Chironomidae: Tanypodinae). Proceedings of The Academy of Natural Sciences, 129, 125–150.
Sæther, O.A. (1977) Female genitalia in Chironomidae and other Nematocera: morphology, phylogenies, keys. Bulletin of the Fisheries Research Board of Canada, 197, 1–211.
Sæther, O.A. (1980) Glossary of chironomid morphology terminology (Diptera: Chironomidae). Entomologica Scandinavica, Supplement, 14, 1–51.
Spies, M. & Reiss, F. (1996) Catalog and bibliography of Neotropical and Mexican Chironomidae (Insecta, Diptera). Spixiana, Supplement, 22, 61–119.
Strait, D.S., Moniz, M.A. & Strait, P.T. (1996) Finite mixture coding: a new approach to coding continuous characters. Systematic Biology, 45, 67–78.
Thiele, K. (1993) The holy grail of the perfect character: the cladistic treatment of morphometric data. Cladistics, 9, 275–304.
Thorpe, R.S. (1984) Coding morphometric characters for constructing distance Wagner networks. Evolution, 38, 244–355.
Watson, C.N. & Heyn, M.W. (1992) A preliminary survey of the Chironomidae (Diptera) of Costa Rica, with emphasis on the lotic fauna. Netherlands journal of Aquatic Ecology, 26, 257–262.
Wiens, J.J. (2001) Character analysis in morphological phylogenetics: problems and solutions. Systematic Biology, 50, 689–699.