Abstract
The Mesozoic chironomid genus Jurochlus Kalugina, 1985, known only as pupae, is reviewed. Four new species ofJurochlus are described from the Upper Jurassic and Lower Cretaceous deposits of Mongolia, viz. J. trivittatus sp. nov.,J. limbatus sp. nov., J. lineatus sp. nov. (Shar Teg, J 3 ) and J. adustus sp. nov. (Khutel Khara, J 3 /K 1 ). Both previouslydescribed species, J. sibiricus Kalugina, 1985 and J. rigor Kalugina, 1985, are re-described in detail based on re-examination of the holotypes. The diagnosis of the genus Jurochlus is emended and its systematic position (probably Podonominae or Tanypodinae) is discussed.References
Ansorge, J. (1996) Insekten aus dem oberen Lias von Grimmen (Vorpommern, Norddeutschland). Neue Paläontologische Abhandlungen, 2, 1–132.
Ansorge, J. (1999) Aenne liasina gen. et sp. n.—the most primitive non biting midge (Diptera: Chironomidae: Aenneinae subfam. n.)—from the Lower Jurassic of Germany. Polskie Pismo Entomologiczne, 68, 431–443.
Azar, D., Veltz, I. & Nel, A. (2008) Mandibulate chironomids: primitive or derived? (Diptera: Chironomidae). Systematic Entomology, 33, 688–699.
Azar, D. & Nel, A. (2010) Two new non-biting midges from the Early Cretaceous Lebanese amber (Diptera: Chironomidae). Annales de la Société entomologique de France (n.s.), 46, 198–203.
Brundin, L. (1962) Insects of Macquarie island. Diptera: Chironomidae. Pacific Insects, 4, 945–954.
Brundin, L. (1966) Transantarctic relationships and their significance, as evidenced by chironomid midges with a monograph of the subfamilies Podonominae and Aphroteniinae and the Austral Hertagyiae. Kunglica Svenska Vetenskapsakademiens Handlingar, Fjärde Serien, 11(1), 1– 472.
Brundin, L. (1986) The pupae of Podonominae (Diptera: Chironomidae) of Holarctic region - Keys and diagnoses. In: Wiederholm, T. (Ed.), Chironomidae of the Holarctic region. Keys and diagnoses. Part 2. Pupae. Entomologica scandinavica, supplement 28, 19–30.
Cranston, P.S., Edward, D.H.D. & Colless, D.H. (1987) Archaeochlus Brundin: a midge out of time (Diptera: Chironomidae). Systematic Entomology, 12, 313–334.
Cranston, P.S. & Edward, D.H.D. (1992) A systematic reappraisal of the Australian Aphroteniinae (Chironomidae) with dating from vicariance biogeography. Systematic Entomology, 17, 41–54.
Cranston, P.S., Hardy, N.B., Morse, G.E., Puslednik, L. & McCluen, S.R. (2010) When molecules and morphology concur: the “Gondwanan” midges (Diptera: Chironomidae). Systematic Entomology, 35, 636–648.
Cranston, P.S., Hardy, N.B. & Morse G.E. (2012) A dated molecular phylogeny for the Chironomidae (Diptera). Systematic Entomology, 37, 172–188.
Fittkau, E.J. (1962) Die Tanypodinae (Diptera: Chironomidae) (Die Tribus Anatopyniini, Macropelopiini und Pentaneurini). Abhandlungen zur Larvalsystematik der Insekten, 6, 1–453.
Fittkau, E.J. & Murray, D.A. (1986) The pupae of Tanypodinae (Diptera: Chironomidae) of the Holarctic region - Keys and diagnoses. In: Wiederholm, T. (Ed.). Chironomidae of the Holarctic region. Keys and diagnoses. Part 2. Pupae. Entomologica scandinavica, supplement 28, 31–113.
Gubin, Y.M. & Sinitza, S.M. (1996) Shar Teg: a unique Mesozoic locality of Asia. Museum of Northern Arizona Bulletin, 60, 311–318.
Jell, P.A. & Duncan, P.M. (1986) Invertebrates, mainly insects, from the freshwater, Lower Cretaceous, Koonwarra Fossil Bed (Korumburra Group), South Gippsland, Victoria. Memoirs of the Association of Australasian Palaeontologists, 3, 111–205.
Kalugina, N.S. (1993) Chaoborids and non-biting midges from the Mesozoic of Eastern Transbaikalia (Diptera: Chaoboridae and Chironomidae). Trudy Paleontologicheskogo Instituta, 252, 117–139. Nauka, Moscow. [In Russian].
Kalugina, N.S., Kovalev, V.G. (1985) Dvukrylye nasekomye Yury Sibiri [Jurassic Diptera of Siberia]. Nauka, Moscow, 198 pp. [In Russian].
Langton, P.H. (1991) A key to pupal exuviae of West Palaearctic Chironomidae. 386 pp.
Lukashevich, E.D. (2009) Limoniidae (Diptera) in the Upper Jurassic of Shar Teg, Mongolia. Zoosymposia, 3, 131–154.
Lukashevich, E.D. (2011) New nematocerans (Insecta: Diptera) from the Late Jurassic of Mongolia. Paleontologicheskii Zhurnal, 6, 26–33.
Lukashevich, E.D. & Przhiboro, A.A. (2011) New Chironomidae (Diptera) with elongate proboscises from the Late Jurassic of Mongolia. In: Shcherbakov, D., Engel, M., Sharkey, M. (Eds.) Advances in the systematics of fossil and modern insects: honouring Alexandr Rasnitsyn. Zookeys, 130, 307–322.
Lukashevich, E.D. (in press). Pupae of Mesozoic Oryctochlus Kalugina, 1985 (Chironomidae: Podonominae), with description of two new species. Fauna norvegica.
Rasnitsyn, A.P. (1996) Conceptual issues in phylogeny, taxonomy and nomenclature. Contributions to Zoology, 66, 3–41.
Rasnitsyn, A.P. & Quicke, L.J. (2002) History of insects. Kluwer Academic Publishers, Dordrecht, Boston, London, 517 pp.
Roback, S.S. (1978). The immature chironomids of the Eastern United States. III. Tanypodinae-Anatopyniini, Macropelopiini and Natarsiini. Proceedings of the Academy of Natural Sciences of Philadelphia, 129, 151–202.
Sæther, O.A. (1980) Glossary of chironomid morphology terminology (Diptera: Chironomidae). Entomologica scandinavica, supplement 14, 1–51.
Veltz, I., Azar, D. & Nel, A. (2007) New chironomid flies in Early Cretaceous Lebanese amber (Diptera: Chironomidae). African Invertebrates, 48, 169–191.
Zhang, J. (1991) New genera and new species of Chironomidae (Diptera, Insecta) from Late Jurassic of China. Acta Paleontologica Sinica, 30, 556–569.