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Type: Monograph
Published: 2017-07-14
Page range: 1–65
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Benthic hydroids (Cnidaria, Hydrozoa) from the Ross Sea (Antarctica) collected by the New Zealand Antarctic expedition BioRoss 2004 with RV Tangaroa

Instituto Cavanilles de Biodiversidad y Biología Evolutiva / Departamento de Zoología, Universidad de Valencia, Valencia, Spain, Apdo. Correos 22085, 46071 Valencia, Spain.
Coelenterata Southern Ocean Biodiversity Continental shelf New species New records Checklist

Abstract

During the New Zealand BioRoss 2004 survey, with RV Tangaroa, sampling of marine communities on the Ross Sea shelf was undertaken. Samples were obtained employing several sampling gears (Van Veen grab, epibenthic sled, rough bottom trawl, and beam trawl). Among the numerous benthic samples obtained, a large and important collection of hydroids was present. Sixty-one species, four of them new to science (Monocoryne antarctica sp. nov., Halecium tangaroa sp. nov., Staurotheca gracilis sp. nov. and Symplectoscyphus densus sp. nov.) have been recorded. Oswaldella blanconae sp. nov. is also described. “Anthoathecata” are represented by ten species belonging to the families Bougainvilliidae, Candelabridae, Clathrozoellidae, Hydractiniidae, Eudendriidae and Tubulariidae. Leptothecata are dominant, with 51 species belonging to the families Campanulariidae, Campanulinidae, Lafoeidae, Haleciidae, Hebellidae, Kirchenpaueriidae, Schizotrichidae, Staurothecidae, Symplectoscyphidae and Zygophylacidae. Symplectoscyphidae is the richest family with 19 species (31%), followed by Staurothecidae with eight (13%) and Haleciidae with seven species (11%). At the generic level, Symplectoscyphus with 14 species (two of them identified only at the genus level), Staurotheca with eight species (one identified to genus level) and Halecium with seven species are the most speciose genera. Twenty-two species (including the four new species) constitute new records for the Ross Sea, thus raising the number of valid, known species of the area to 77.

 

References

  1. Allman, G.J. (1888) Report on the Hydroida dredged by H.M.S. Challenger during the years 1873–76. II. Tubularinae, Corymorphynae, Campanularinae, Sertularinae and Thalamophora. Report of the Scientific Results of the Voyage of H.M.S. Challenger 1873–1876, 23 (70), i–xix + 1–90, 40 pls.

    Bale, W.M. (1884) Catalogue of the Australian Hydroid Zoophytes. Australian Museum, Sydney, 198 pp., 19 pls.

    Billard, A. (1906) Hydroïdes. In: Joubin, L. (Ed.), Expédition Antarctique Française (1903–1905) commandée par le Dr Jean Charcot. Masson et Cie, Paris, pp. 1–20.

    Billard, A. (1914) Hydroïdes. In: Deuxième Expédition Antarctique Française (1908–1910), commandée par le Dr Jean Charcot, Paris, 34 pp.

    Blanco, O.M. (1969) Un nuevo Sertulárido antártico, Symplectoscyphus naumovi nov. sp. Neotrópica, 15 (46), 14–16.

    Blanco, O.M. (1976) Hidrozoos de la expedición Walther Herwig. Revista del Museo de La Plata, Zoología, Nueva Serie, 12 (113), 27–74, pls 1–8.

    Blanco, O.M. (1977a) Nuevos hidrozoos antárticos. Contribuciones del Instituto Antártico Argentino, 205, 3–15.

    Blanco, O.M. (1977b) Una nueva especie de Symplectoscyphus argentina. Contribuciones del Instituto Antártico Argentino, 206, 1–12.

    Blanco, O.M. (1982) Un nuevo sertulárido de Isla Baja. Revista del Museo de La Plata, Zoología, Nueva Serie, 13 (133), 39–43, figs. 1–7 on 2 pls.

    Blanco, O.M. (1984) Contribución al conocimiento de hidrozoos antárticos y subantárticos. Contribuciones del Instituto Antártico Argentino, 294, 1–53, pls. 1–47.

    Bonnevie, K. (1899) Hydroida. Norske Nordhavs-Expedition 1876–1878, Zoologi, 26, 1–104, pls. 1–8, map.

    Briggs, E.A. (1938) Hydroida. Scientific Reports Australasian Antarctic Expedition 1911–1914, Series C, 9 (4), 1–46, pls. 15–16.

    Brinckmann Voss, A. & Lindner, A. (2008) Monocoryne colonialis sp. nov., a colonial candelabrid hydroid (Cnidaria: Hydrozoa: Candelabridae) from the north Pacific. Journal of the Marine Biological Association of the United Kingdom, 88 (8), 1631–1635.
    https://doi.org/10.1017/S002531540800180X

    Broch, H. (1948) Antarctic Hydroids. Scientific Results of the Norwegian Antarctic Expedition 1927–1928, et seq., 28, 1–23.

    Cerrano, C., Puce, S., Chiantore, M. & Bavestrello, G. (2000) Unusual trophic strategies of Hydractinia angusta (Cnidaria, Hydrozoa) from Terra Nova Bay, Antarctica. Polar Biology, 23, 488–494.
    https://doi.org/10.1007/s003000000110

    Cerrano, C., Puce, S., Chiantore, M., Bavestrello, G. & Cattaneo-Vietti, R. (2001) The influence of the epizoic hydroid Hydractinia angusta on the recruitment of the Antarctic scallop Adamussium colbecki. Polar Biology, 24, 577–581.

    https://doi.org/10.1007/s003000100254

    Clark, S.F. (1875) Descriptions of new and rare species of hydroids from the New England coasts. Transactions of the Connecticut Academy of Arts and Sciences, 3, 58–66, pls. 9–10.

    Clark, S.F. (1877) Report on the hydroids collected on the coast of Alaska and the Aleutian Islands, by W.H. Dall, U.S. Coast Survey, and Party, from 1871 to 1874 inclusive. Proceedings of the Academy of Natural Sciences of Philadelphia, 1876, 209–238, pls 7–16.

    Clark, M. & Rowden, A. (2004) BioRoss 2004: expanding our knowledge of marine life in the Ross Sea. Water & Atmosphere, 12 (3), 24–25.

    Dayton, P.K., Robilliard, G.A. & Paine, R.T. (1970) Benthic faunal zonation as a result of anchor ice at McMurdo Sound, Antarctica. In: Holdgate, W. (Ed.), Antarctic Ecology. Academic Press, London, pp. 244–257.

    El Beshbeeshy, M. (1991) Systematische, Morphologische und Zoogeographische Untersuchungen an den Thekaten Hydroiden des Patagonischen Schelfs. University of Hamburg, Hamburg, 390 pp.

    Fleming, J. (1820) Observations on the natural history of the Sertularia gelatinosa of Pallas. Edinburgh Philosophical Journal, 2, 82–89.

    Fraser, C.M. (1943) New species of hydroids, mostly from the Atlantic Ocean, in the United States National Museum. Proceedings of the United States National Museum, 91, 77–89, pls. 13–17.

    Galea, H.R. & Schories, D. (2012) Some hydrozoans (Cnidaria) from King George Island, Antarctica. Zootaxa, 3321, 1–21.

    González Molinero, A. & Peña Cantero, A.L. (2015) SEM study of species of Oswaldella Stechow, 1919 (Cnidaria, Hydrozoa, Kirchenpaueriidae), with an annotated checklist of the species of the genus. Zootaxa, 4052 (4), 401–441.
    https://doi.org/10.11646/zootaxa.4052.4.1

    Hartlaub, C. (1904) Hydroiden. In: Expédition antarctique Belge. Résultats du voyage du S.Y. Belgica en 1897–1899. Rapports scientifiques, Zoologie, 1–19.

    Hickson, S.J. & Gravely, F.H. (1907) Coelenterata. II. Hydroid zoophytes. National Antarctic Expedition (S.S. Discovery) 1901–1904, Natural History, 3, 1–34, pls. 1–4.

    Hincks, Th. (1861) A catalogue of the zoophytes of South Devon and South Cornwall. Annals and Magazine of Natural History, Series 3, 8, 152–161, 251–262, 290–297, pls. 6–8.

    Jäderholm, E. (1903) Aussereuropäischen Hydroiden im schwedischen Reichsmuseum. Arkiv för Zoologi, 1, 259–312, pls. 12–15.

    Jäderholm, E. (1904) Mitteilungen ueber einige von der Schwedischen Antarctic‑Expedition 1901–1903 eingesammelte Hydroiden. Archives de Zoologie Expérimentale et Générale, Series 4, 3 (Notes et revue 1), 1–14.

    Jäderholm, E. (1905) Hydroiden aus antarktischen und subantarktischen Meeren, gesammelt von der schwedischen Sudpolarexpedition. Wissenschaftliche Ergebnisse der Schwedischen Südpolar-Expedition 1901–1903, 5 (8), 1–41, pls 1–14.

    Jäderholm, E. (1917) Hydroids from the South Seas. Redogörelse för Norrköpings Högre Allmänna Läroverk Läsaret, 1916–1917, 1–25, pls. 1–2.

    Jäderholm, E. (1926) Ueber einige antarktische und subantarktische Hydroiden. Arkiv för Zoologi, Series A, 18 (14), 1–7.

    Kayal, E., Bentlage, B., Cartwright, P., Yanagihara, A.A., Lindsay, D.J., Hopcroft, R.R. & Collins, A.G. (2015) Phylogenetic analysis of higher-level relationships within Hydroidolina (Cnidaria: Hydrozoa) using mitochondrial genome data and insight into their mitochondrial transcription. PeerJ, 3, e1403.
    https://doi.org/10.7717/peerj.1403

    Maronna, M.M., Miranda, T.P., Peña Cantero, A.L., Barbeitos, M.S. & Marques, A.C. (2016) Towards a phylogenetic classification of Leptothecata (Cnidaria, Hydrozoa). Scientific Reports, 6, 18075.
    https://doi.org/10.1038/srep18075

    Marques, A.C., Mergner, H., Höinghaus, R., Santos C.M.D. & Vervoort, W. (2000) Morphological study and taxonomical notes on Eudendriidae (Cnidaria: Hydrozoa: Athecatae/Anthomedusae). Zoologische Mededelingen, Leiden, 74 (5), 75–118.

    Marques, A.C., Peña Cantero, A.L. & Migotto, A.E. (2005) Redescription and systematic status of the Antarctic genus Abietinella Levinsen, 1913 (Lafoeidae, Hydrozoa, Cnidaria). Journal of Natural History, 39 (18), 1443–1455.
    https://doi.org/10.1080/00222930400005765

    Millard, N.A.H. (1964) The Hydrozoa of the south and west coasts of South Africa. Part II. The Lafoeidae, Syntheciidae and Sertulariidae. Annals of the South African Museum, 48, 1–56.

    Millard, N.A.H. (1966) The Hydrozoa of the south and west coasts of South Africa. Part III. The Gymnoblastea and small families of the Calyptoblastea. Annals of the South African Museum, 48 (18), 427–487, pl. 1.

    Millard, N.A.H. (1975) Monograph on the Hydroida of southern Africa. Annals of the South African Museum, 68, 1–513.

    Millard, N.A.H. (1977) Hydroids from the Kerguelen and Crozet shelves, collected by the cruise MD.03 of the Marion‑Dufresne. Annals of the South African Museum, 73 (1), 1–47.

    Miranda, T.P., Cunha, A.F. & Marques, A.C. (2016) Status of the names of some hydroid species (Cnidaria, Hydrozoa) described from the Atlantic coast of Patagonia. Zootaxa, 4171 (3), 595–600.
    https://doi.org/10.11646/zootaxa.4171.3.13

    Naumov, D.V. & Stepanjants, S.D. (1962) Hydroida (Thecaphora) collected by the Soviet Antarctic Expedition on the M/V Ob in antarctic and subantarctic waters. In: Biological results of the Soviet Antarctic Expedition, 1955–1958, 1. Issledovaniya Fauny Morei, 1 (9), 68–106.

    Naumov, D.V. & Stepanjants, S.D. (1972) Marine invertebrates from Adélie Land collected by the XIIth and XVth French Antarctic Expeditions. 3. Hydroida. Téthys Supplément, 4, 25–60.

    Peña Cantero, A.L. (1998) Two new antarctic species of the genus Schizotricha Allman, 1883 (Cnidaria, Hydrozoa). Polar Biology, 19 (2), 77–84.
    https://doi.org/10.1007/s003000050217

    Peña Cantero, A.L. (2006) Benthic hydroids from the south of Livingston Island (South Shetland Islands, Antarctica) collected by the Spanish Antarctic expedition Bentart 94. Deep-Sea Research II, 53, 932–948.
    https://doi.org/10.1016/j.dsr2.2006.02.005

    Peña Cantero, A.L. (2007) Breaking molds: Oswaldella laertesi, sp. nov., a unique Antarctic species of Oswaldella Stechow, 1919 (Cnidaria: Hydrozoa: Kirchenpaueriidae). Zootaxa, 1612, 63–68.

    Peña Cantero, A.L. (2008) Benthic hydroids (Cnidaria: Hydrozoa) from the Spanish Antarctic expedition Bentart 95. Polar Biology, 31, 451–464.
    https://doi.org/10.1007/s00300-007-0371-2

    Peña Cantero, A.L. (2009) Benthic hydroids (Cnidaria, Hydrozoa) from the Balleny Islands (Antarctica). Polar Biology, 32, 1743–1751.
    https://doi.org/10.1007/s00300-009-0673-7

    Peña Cantero, A.L. (2010a) On a new Antarctic species of Symplectoscyphus Marktanner-Turneretscher, 1890 (Cnidaria, Hydrozoa, Sertulariidae), with an annotated checklist of the Antarctic species of the genus. Zootaxa, 2494, 29–44.

    Peña Cantero, A.L. (2010b) Benthic hydroids (Cnidaria: Hydrozoa) from Peter I Island (Southern Ocean, Antarctica). Polar Biology, 33, 761–773.
    https://doi.org/10.1007/s00300-009-0754-7

    Peña Cantero, A.L. (2012) Filling biodiversity gaps: benthic hydroids from the Bellingshausen Sea (Antarctica). Polar Biology, 35, 851–865.
    https://doi.org/10.1007/s00300-011-1130-y

    Peña Cantero, A.L. (2013) Benthic hydroids from off Low Island (Southern Ocean, Antarctica). Marine Ecology, 34 (Supplement 1), 123–142.
    https://doi.org/10.1111/maec.12031

    Peña Cantero, A.L. (2014a) Benthic hydroids (Cnidaria, Hydrozoa) from the continental shelf and slope off Queen Mary Coast (East Antarctica). Polar Biology, 37, 1711–1731.
    https://doi.org/10.1007/s00300-014-1556-0

    Peña Cantero, A.L. (2014b) Revision of the Antarctic species of Halecium Oken, 1815 (Cnidaria, Hydrozoa, Haleciidae). Zootaxa, 3790 (2), 243–280.
    https://doi.org/10.11646/zootaxa.3790.2.2

    Peña Cantero, A.L. (2015) Review of some little-known benthic hydroids (Cnidaria, Hydrozoa) from the Southern Ocean. Zootaxa, 3972 (3), 369–392.
    https://doi.org/10.11646/zootaxa.3972.3.4

    Peña Cantero, A.L., Boero, F. & Piraino, S. (2013) Shallow-water benthic hydroids from Tethys Bay (Terra Nova Bay, Ross Sea, Antarctica). Polar Biology, 36, 731–753.
    https://doi.org/10.1007/s00300-013-1299-3

    Peña Cantero, A.L. & García Carrascosa, A.M. (1993) The coppinia of Abietinella operculata (Lafoeidae: Hydrozoa Leptomedusae) and its systematic position. Journal of Natural History, 27, 1003–1011.
    https://doi.org/10.1080/00222939300770631

    Peña Cantero, A.L. & García Carrascosa, A.M. (1994) Hidrozoos de la campaña ‘Antártida 8611’. In: Castellví, J. (Ed.), Actas del cuarto simposio español de estudios antárticos. CICYT, Madrid, pp. 117–140.

    Peña Cantero, A.L. & García Carrascosa, A.M. (1995) Hidrozoos bentónicos de la campaña Antártida 8611. Publicaciones Especiales del Instituto Español de Oceanografía, 19, 1–148.

    Peña Cantero, A.L. & García Carrascosa, A.M. (1998) Oswaldella vervoorti spec. nov. (Cnidaria: Hydrozoa), a new benthic hydroid from the South Shetland Islands, Antarctica. Zoologische Verhandelingen, 232, 175–180.

    Peña Cantero, A.L. & García Carrascosa, A.M. (1999) Biogeographical distribution of the benthic thecate hydroids collected during the Spanish ‘Antártida 8611’ expedition and comparison between Antarctic and Magellan benthic hydroid faunas. Scientia Marina, 63 (Supplement 1), 209–218.
    https://doi.org/10.3989/scimar.1999.63s1209

    Peña Cantero, A.L., García Carrascosa, A.M. & Vervoort, W. (1997) On Antarctoscyphus (Cnidaria, Hydrozoa), a new genus of antarctic hydroids and the description of two new species. Polar Biology, 18, 23–32.
    https://doi.org/10.1007/s003000050155

    Peña Cantero, A.L., García Carrascosa, A.M. & Vervoort, W. (1998) On the species of Filellum Hincks, 1868 (Cnidaria: Hydrozoa) with the description of a new species. Journal of Natural History, 32, 297–315.
    https://doi.org/10.1080/00222939800770151

    Peña Cantero, A.L. & Gili, J.M. (2006) Benthic hydroids (Cnidaria, Hydrozoa) from off Bouvet Island (Antarctic Ocean). Polar Biology, 29, 764–771.
    https://doi.org/10.1007/s00300-006-0113-x

    Peña Cantero, A.L. & Marques, A.C. (1999) Phylogenetic analysis of the Antarctic genus Oswaldella Stechow, 1919 (Hydrozoa, Leptomedusae, Kirchenpaueriidae). Contributions to Zoology, 68, 83–93.

    Peña Cantero, A.L. & Ramil, F. (2006) Benthic hydroids associated with volcanic structures from Bransfield Strait (Antarctica) collected by the Spanish Antarctic expedition GEBRAP96. Deep-Sea Research II, 53, 949–958.
    https://doi.org/10.1016/j.dsr2.2006.02.007

    Peña Cantero, A.L., Svoboda, A. & Vervoort, W. (1996) Species of Schizotricha Allman, 1883 (Cnidaria, Hydrozoa) from recent antarctic expeditions with R.V. ‘‘Polarstern’’, with the description of a new species. Zoologische Mededelingen, Leiden, 70, 411–435.

    Peña Cantero, A.L., Svoboda, A. & Vervoort, W. (1997a) Species of Staurotheca Allman, 1888 (Cnidaria, Hydrozoa) from recent Antarctic expeditions with R.V. ‘Polarstern’, with the description of six new species. Journal of Natural History, 31, 329–381.
    https://doi.org/10.1080/00222939700770171

    Peña Cantero, A.L., Svoboda, A. & Vervoort, W. (1997b) Species of Oswaldella Stechow, 1919 (Cnidaria, Hydrozoa) from recent Antarctic expeditions with R.V. “Polarstern”, with the description of eight new species. Zoological Journal of the Linnean Society, 119, 339–388.

    Peña Cantero, A.L., Svoboda, A. & Vervoort, W. (1999) Species of Antarctoscyphus Peña Cantero, García Carrascosa and Vervoort, 1997 (Cnidaria, Hydrozoa, Sertulariidae) from recent Antarctic expeditions with R.V. Polarstern, with the description of two new species. Journal of Natural History, 33, 1739–1765.
    https://doi.org/10.1080/002229399299707

    Peña Cantero, A.L., Svoboda, A. & Vervoort, W. (2002) Species of Symplectoscyphus Marktanner-Turneretscher, 1890 (Cnidaria: Hydrozoa, Sertulariidae) from recent Antarctic expeditions with R.V. Polarstern, with the description of four new species. Journal of Natural History, 36, 1509–1568.
    https://doi.org/10.1080/00222930110051716

    Peña Cantero, A.L., Svoboda, A. & Vervoort, W. (2004) Antarctic hydroids (Cnidaria: Hydrozoa) of the families Campanulinidae, Lafoeidae and Campanulariidae from recent Antarctic expeditions with R.V. Polarstern, with the description of a new species. Journal of Natural History, 38, 2269–2303.
    https://doi.org/10.1080/00222930310001647361

    Peña Cantero, A.L. & Vervoort. W. (1996) On two new species of Oswaldella Stechow, 1919: O. terranovae spec. nov. and O. tottoni spec. nov. (Cnidaria Hydrozoa) (Notes on Antarctic hydroids, III). Zoologische Mededelingen, 70, 135–143.

    Peña Cantero, A.L. & Vervoort. W. (1998) On two new species of Oswaldella Stechow, 1919 (Cnidaria, Hydrozoa) from Bransfield Strait (Antarctica). Polar Biology, 20, 33–40.
    https://doi.org/10.1007/s003000050273

    Peña Cantero, A.L. & Vervoort. W. (1999) Review of the genus Schizotricha Allman, 1883 (Cnidaria, Hydrozoa, Halopterididae). Journal of Natural History, 33, 351–386.
    https://doi.org/10.1080/002229399300290

    Peña Cantero, A.L. & Vervoort, W. (2003) Species of Staurotheca Allman, 1888 (Cnidaria: Hydrozoa: Sertulariidae) from US Antarctic expeditions, with the description of three new species. Journal of Natural History, 37, 2653–2722.
    https://doi.org/10.1080/00222930210155701

    Peña Cantero, A.L. & Vervoort, W. (2004) Species of Oswaldella Stechow, 1919 (Cnidaria: Hydrozoa: Kirchenpaueriidae) from US Antarctic expeditions, with the description of three new species. Journal of Natural History, 38, 805–861.
    https://doi.org/10.1080/0022293021000045154

    Peña Cantero, A.L. & Vervoort, W. (2005) Species of Schizotricha Allman, 1883 (Cnidaria: Hydrozoa: Halopterididae) from US Antarctic expeditions with the description of two new species. Journal of Natural History, 39, 795–818.
    https://doi.org/10.1080/00222930400001541

    Peña Cantero, A.L. & Vervoort, W. (2009) Benthic hydroids (Cnidaria: Hydrozoa) from the Bransfield Strait area (Antarctica) collected by Brazilian expeditions, with the description of a new species. Polar Biology, 32, 83–92.
    https://doi.org/10.1007/s00300-008-0506-0

    Peña Cantero, A.L., Vervoort. W. & Watson, J.E. (2003) On Clathrozoellidae (Cnidaria, Hydrozoa, Anthoathecatae), a new family of rare deep-water leptolids, with the description of three new species. Zoologische Verhandelingen, 345, 281–296.

    Puce, S., Cerrano, C. & Bavestrello, G. (2002) Eudendrium (Cnidaria, Anthomedusae) from the Antarctic Ocean with description of two new species. Polar Biology, 25, 366–373.

    Ritchie, J. (1907) The hydroids of the Scottish National Antarctic Expedition. Transactions of the Royal Society of Edinburgh, 45 (2), 519–545, pls. 1–3.
    https://doi.org/10.1017/s0080456800022821

    Ritchie, J. (1913) The hydroid zoophytes collected by the British Antarctic Expedition of Sir Ernest Shackleton, 1908. Proceedings of the Royal Society of Edinburgh, 33 (1), 9–34.

    Schuchert, P., Sanamyan, N. & Sanamyan, K. (2016) Observations on two large athecate hydroids (Cnidaria: Hydrozoa) from the Kamchatka Peninsula (NW Pacific). Revue suisse de Zoologie, 123 (1), 165–178.

    Soto Àngel, J.J. & Peña Cantero, A.L. (2015) On the benthic hydroids from the Scotia Arc (Southern Ocean): new insights into their biodiversity, ecology and biogeography. Polar Biology, 38, 983–1007.
    https://doi.org/10.1007/s00300-015-1660-9

    Stechow, E. (1921) Über Hydroiden der Deutschen Tiefsee-Expedition, nebst Bemerkungen über einige andre Formen. Zoologischer Anzeiger, 53 (9–10), 223–236.

    Stechow, E. (1922) Zur Frage des bipolaren Vorkommen mariner Tiere. Verhandlungen der Deutschen Zoologischen Gesellschaft, 27, 95–96.

    Stechow, E. (1923a) Über Hydroiden der Deutschen Tiefsee-Expedition, nebst Bemerkungen über einige andere Formen. Zoologischer Anzeiger, 56, 97–119.

    Stechow, E. (1923b) Zur Kenntnis der Hydroidenfauna des Mittelmeeres, Amerikas und anderer Gebiete. II. Teil. Zoologische Jahrbücher, Abteilung für Systematik, 47 (1), 29–270.

    Stechow, E. (1932) Neue Hydroiden aus dem Mittelmeer und dem Pazifischen Ozean, nebst Bemerkungen über einige wenig bekannte Formen. Zoologischer Anzeiger, 100 (3–4), 81–92.

    Stepanjants, S.D. (1972) Hydroidea of the coastal waters of the Davis Sea (collected by the XIth Soviet Antarctic Expedition of 1965–1966). In: Biological results of the Soviet Antarctic Expedition, 5. Issledovaniya Fauny Morei, 11 (19), 56–79. [in Russian]

    Stepanjants, S.D. (1979) Hydroids of the antarctic and subantarctic waters. In: Biological results of the Soviet Antarctic Expedition, 6. Issledovaniya Fauny Morei, 20 (30), 1–200, pls. 1–25. [in Russian]

    Svoboda, A. & Stepanjants, S.D. (2001) Redescription of two Antarctic Corymorphidae species and the reestablishment of the genus Monocaulus (Cnidaria: Hydrozoa). Pubblicazione Stazione Zoologica Napoli, Marine Ecology, 22 (1–2), 53–70.
    https://doi.org/10.1046/j.1439-0485.2001.00742.x

    Svoboda, A., Stepanjants, S. & Smirnov, I. (1997) Two polar Hydractinia species (Cnidaria), epibiotic on two closely related brittle stars (Echinodermata): an example for taxonomic and ecological bipolarity. In: Battaglia, B., Valencia, J. & Walton, D.W.H. (Eds.), Antarctic Communities: Species, Structure and Survival. Cambridge University Press, Cambridge, pp. 22–25.

    Totton, A.K. (1930) Coelenterata. Part V. Hydroida. Natural History Report, British Antarctic 'Terra Nova' Expedition, 1910, Zoology, 5 (5), 131–252, pls 1–3.

    Vanhöffen, E. (1910) Die Hydroiden der Deutschen Südpolar‑Expedition 1901–1903. Deutsche Südpolar-Expedition 1901–1903, 11 (Zoology 3), 269–340.

    Vervoort, W. (1972a) Hydroids from the Theta, Vema and Yelcho cruises of the Lamont‑Doherty geological observatory. Zoologische Verhandelingen, 120, 1–247.

    Vervoort, W. (1972b) Hydroids from submarine cliffs near Arthur Harbour, Palmer Archipelago, Antarctica. Zoologische Mededelingen, 47 (25), 337–357.

    Vervoort, W. & Watson, J.E. (1996) A rare hydroid, Clathrozoella drygalskii (Vanhöffen, 1910) in the waters of Australia and New Zealand. Scientia Marina, 60 (1), 117–120.

    Von Lendenfeld, R. (1885) The Australian Hydromedusae. II. Proceedings of the Linnean Society of New South Wales, 9, 345–353, pl. 6.

    Watson, J.E. (2008) Hydroids of the BANZARE expeditions, 1929–1931: the family Haleciidae (Hydrozoa, Leptothecata) from the Australian Antarctic Territory. Memoirs of the Museum of Victoria, 65, 165–178.
    https://doi.org/10.24199/j.mmv.2008.65.9