Abstract
Chalcocopris hesperus (originally Scarabaeus hesperus) (Coleoptera, Scarabaeidae, Scarabaeinae, Dichotomiini) is a common and unmistakable dung beetle species from the Atlantic Forest of Brazil, Paraguay and Argentina. In 2015, a neotype was designated for its valid name based on the belief that the original type series had been lost. However, the designation was invalid for failing to comply with Article 75.3.1 of the Code. Moreover, even if it had been originally valid, the neotype would now lose its type status for we found a syntype precisely where it was originally said to be, in the material of the Joseph Banks collection, now preserved in the Natural History Museum, London. We present the evidence supporting the recognition of this specimen as a syntype and designate it as the lectotype. The specimen, from Rio de Janeiro, Brazil, may have been collected by Sir Joseph Banks himself during the first Cook voyage round the world (1768–1771), but we discuss aspects that might indicate that it may actually have been caught later and by someone else. Finally, we show that Scarabaeus hesperus was fixed as the type species of Chalcocopris not in 2016, as recently argued by one of us, but in 1998 by Olivier Montreuil. The data gathered and discussed here will hopefully prove to be valuable to anyone investigating the history and type status of specimens from the Banksian entomological collection.
References
- Anonymous (1811) Donations to the museum of the Linnean Society, exclusive of many presents of single specimens of animals, plants, and minerals. The Transactions of the Linnean Society of London, 10: 413‒414. <https://doi.org/10.1111/j.1096-3642.1810.tb00034.x>.
- Anonymous (1815) Donations to the museum of the Linnean Society, exclusive of many presents of single specimens of animals, plants, and minerals. Continued from page 414 of Vol. X. of the Society’s transactions. The Transactions of the Linnean Society of London, 11: 430.
- Anonymous [International Commission on Zoological Nomenclature] (1999) International code of zoological nomenclature. Fourth edition adopted by the International Union of Biological Sciences. London (International Trust for Zoological Nomenclature): i–xxix +1–306.
- Almeida, S. S. P. & Louzada, J. N. C. (2009) Estrutura da comunidade de Scarabaeinae (Scarabaeidae: Coleoptera) em fitofisionomias do Cerrado e a sua importância para a conservação. Neotropical Entomology, 38 (1): 32‒43. <https://doi.org/10.1590/S1519-566X2009000100003>.
- Alonso, C. B. G., Zurita, G. A. & Bellocq, M. I. (2022) Livestock grazing impact differently on the functional diversity of dung beetles depending on the regional context in subtropical forests. Scientific Reports, 12 (1636): 1‒11. <https://doi.org/10.1038/s41598-022-05616-x>.
- Anderson, K. (2023) The Mantodea of Johan Fabricius. Soothsayer, 4 (1): 27‒61.
- Andrews, J. R. H. & Gibbs, G. W. (1989) The first New Zealand insects collected on Cook’s Endeavour voyage. Pacific Science, 43 (1): 102‒114.
- Banks, J. (1896) Journal of the Right Hon. Sir Joseph Banks during Captain Cook’s first voyage in H.M.S. Endeavour in 1768‒71 to Terra del Fuego, Otahite, New Zealand, Australia, the Dutch East Indies, etc. Edited by Sir Joseph D. Hooker. With portraits and charts. London (MacMillan and Co.): [i]–[lii] + [1]–466. [Joseph Banks died in 1820. Edited by Joseph Dalton Hooker, this work, Banks’ Endeavour journal, was published posthumously 76 years later]. <https://doi.org/10.5962/bhl.title.29416>.
- Bousquet, Y. (2016) Litteratura Coleopterologica (1758–1900): a guide to selected books related to the taxonomy of Coleoptera with publication dates and notes. Zookeys, 583: 1–776. <https://doi.org/10.3897/zookeys.583.7084>.
- Braga, P. B., Carvalho, R. L., Louzada, J. N. C. & Audino, L. D. (2019) Dung type or intraspecific variation in biomass: What is more important in determining dung beetle ecological function? Oecologia australis, 23 (3): 536–547. <https://doi.org/10.4257/oeco.2019.2303.12>.
- Burmeister, H. (1846) [Heft 10]. • Burmeister, H. (1840–1846) Genera quaedam insectorum. Iconibus illustravit et descripsit. Volumen I. Continet tabulas XL. Berlin [Berolini] (A. Burmeister): [8 unnumbered pages], 4 pl. [Publication date following Bousquet 2016].
- Cambefort, Y. (2006) Des coléoptères, des collections & des hommes. Paris (Publications scientifiques du Muséum national d’Histoire naturelle): 1–375. <https://doi.org/10.4000/books.mnhn.2265>.
- Cassenote, S., da Silva, P. G., Di Mare, R. A. & Paladini, A. (2019) Seasonality of dung beetles (Coleoptera: Scarabaeinae) in Atlantic Forest sites with different levels of disturbance in southern Brazil. Iheringia, Série Zoologia, 109: 1–9. <https://doi.org/10.1590/1678-4766e2019035>.
- Castelnau, Comte de (1840) Histoire naturelle des animaux articulés, annelides, crustacés, arachnides, myriapodes et insectes. Histoire naturelle des insectes coléoptères. Tome deuxième. Paris (Duménil): 1–563 + [i], 38 pl. <https://doi.org/10.5962/bhl.title.47104>.
- Cook, J. (1893) Captain Cook’s journal during his first voyage round the world made in H.M. Bark “Endeavour” 1768‒71. A literal transcription of the original MSS. With notes and introduction edited by Captain W.J.L. Wharton, R.N., F.R.S., hydrographer of the Admiralty. Illustrated by maps and facsimiles. London (Elliot Stock): [i]–lvi + 1–400. [Captain Cook died in 1779 during his third voyage round the world. His first voyage’s diary, the work cited here, was published in full only 114 years later, edited by W. J. L. Wharton].
- Culot, L., Bovy, E., Vaz-de-Mello, F. Z., Guevara, R. & Galetti, M. (2013) Selective defaunation affects dung beetle communities in continuous Atlantic rainforest. Biological Conservation, 163: 79‒89. <https://doi.org/10.1016/j.biocon.2013.04.004>.
- Cupello, M., Ribeiro-Costa, C. S., Gill, B. D., Julio, C. A. & Costa-Silva, V. (2023) New country records for dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) in Bolivia and Paraguay. The Coleopterists Bulletin, 77 (1): 83‒92. <https://doi.org/10.1649/0010-065X-77.1.83>.
- Cupello, M., Rossini, M. & Vaz-de-Mello, F. Z. (2016) On the type species of the South American dung beetle genus Chalcocopris Burmeister, 1846 (Coleoptera: Scarabaeidae: Scarabaeinae), with some comments on the type locality of C. hesperus (Olivier, 1789). Zootaxa, 4061 (3): 274–276. <https://doi.org/10.11646/zootaxa.4061.3.5>.
- Erichson, W. F. (1847) Conspectus Insectorum Coleopterorum, quae in Republica Peruana observata sunt. Archiv für Naturgeschichte, 13 (1): 67–185.
- Evans, G. O. & Hyatt, K. H. (1963) Mites of the genus Macrocheles Latr. (Mesostigmata) associated with coprid beetles in the collections of the British Museum (Natural History). Bulletin of the British Museum (Natural History), Zoology, 9 (9): 325–401. <https://doi.org/10.5962/bhl.part.16344>.
- Fabricius, J. C. (1775) Systema entomologiae, sistens insectorvm classes, ordines, genera, species, adiectis synonymis, locis, descriptionibvs, observationibvs. Flensburg & Leipzig [“Flensbvrgi et Lipsiae”] (Kortius): [i–xxxii] + 1–832. <https://doi.org/10.5962/bhl.title.36510>.
- Fabricius, J. C. (1781) Species insectorvm exhibentes eorvm differentias specificas, synonyma avctorvm, loca natalia, metamorphosin adiectis observationibvs, descriptionibvs. Tomus 1. Hamburg & Kiel [“Hambvrgi et Kilonii”] (Carol. Ernest. Bohn): i–viii + 1–552. <https://doi.org/10.5962/bhl.title.36509>.
- Fabricius, J. C. (1787) Mantissa insectorvm sistens eorvm species nvper detectas adiectis characteribvs genericis, differentiis specificis, emendationibvs, observationibvs. Tomus 1. Copenhagen [“Hafniae”] (Christ. Gottl. Proft.): [i]–xx + [1]–348. <https://doi.org/10.5962/bhl.title.11657>.
- Fabricius, J. C. (1792a) Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tomus 1. Copenhagen [“Hafniae”] (Christ. Gottl. Proft.): [i]‒xx + [1]‒330. [Publication date following Bousquet 2016] <https://doi.org/10.5962/bhl.title.122153>.
- Fabricius, J. C. (1792b) Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tomus 1. Pars 2. Copenhagen [“Hafniae”] (Christ. Gottl. Proft.): [1]‒538. [Publication date following Bousquet 2016]
- Fabricius, J. C. (1801a) Systema eleutheratorum, secundum ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus. Tomus 1. Kiel [“Kiliae”] (Bibliopolii Academici Novi): [i]–xxiv + [1]–506. [Publication date following Bousquet 2016].
- Fabricius, J. C. (1801b) Systema eleutheratorum, secundum ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus. Tomus 2. Kiel [“Kiliae”] (Bibliopolii Academici Novi): [1]–687. [Publication date following Bousquet 2016].
- Fabricius, J. C. (1804) [Autobiography]. • Hope, F. W. (1845) The auto-biography of John Christian Fabricius, translated from the Danish with additional notes and observations. Transactions of the entomological Society of London, 4 (1): i‒xvi. • [Hope does not inform when Fabricius wrote this version of his autobiography—another was written earlier, in 1798 (Tuxen 1967). We assume it was in the spring or summer of 1804 because [1] Fabricius mentions events in his life up to that point in time (e.g., his 1798–1804 spring visits to Copenhagen to see the collections of his friends Sehested and Tønder Lund, as well as the publication of his Systema Piezatorum in 1804), and, at the same time, [2] he says “this summer I expect the excellent Humboldt to return from South America”, and that eventually happened in August 1804 (Papavero 1971). Therefore, he certainly wrote it after [1] the start of the spring of 1804 and [2] before August 1804].
- Falqueto, S. A., Vaz-de-Mello, F. Z. & Schoereder, J. H. (2005) Are fungivorous Scarabaeidae less specialist? Ecología austral, 15: 17‒22.
- Gage, A. T. & Stearn, W. T. (1988) A bicentenary history of the Linnean Society of London. London (Academic Press): i–ix + 1–242.
- Garner, B. H. & Hancock, E. G. (2018) The “lost” types of Carabus pallens Fabricius, 1775 (Coleoptera: Carabidae: Lebiinae) from the Banks and Hunter collections: lectotype designation, redescription, and distribution. The Coleopterists Bulletin, 72 (4): 845‒857. <https://doi.org/10.1649/0010-065X-72.4.845>.
- Genes, L., Fernandez, F. A. S., Vaz-de-Mello, F. Z., Rosa, P., Fernandez, E. & Pires, A. S. (2018) Effects of howler monkey reintroduction on ecological interactions and processes. Conservation Biology, 33 (1): 88‒98. <https://doi.org/10.1111/cobi.13188>.
- Geoffroy, É. L. (1762) Histoire abrégée des insectes qui se trouvent aux environs de Paris; dans laquelle ces animaux sont rangés suivant un ordre méthodique. Tome premier. Paris (Durand): i–xxviii + [1]–523, 10 pl. [Publication date following Bousquet 2016]. <https://doi.org/10.5962/bhl.title.154767>.
- Giménez Gómez, V. C., Verdú, J. R., Alonso, C. B. G. & Zurita, G. A. (2018) Relationship between land uses and diversity of dung beetles (Coleoptera: Scarabaeinae) in the southern Atlantic forest of Argentina: which are the key factors? Biodiversity and Conservation, 27: 3201–3213. <https://doi.org/10.1007/s10531-018-1597-8>.
- Giménez Gómez, V. C., Verdú, J. R., Casanoves, F. & Zurita, G. A. (2022) Functional responses to anthropogenic disturbance and the importance of selected traits: a study case using dung beetles. Ecological Entomology, 47 (4): 503–514. <https://doi.org/10.1111/een.13135>.
- Giménez Gómez, V. C., Verdú, J. R. & Zurita, G. A. (2019) Factores claves que afectan a la preferencia trófica y diversidad de coleópteros copro-necrófagos del bosque Atlántico de Argentina: una propuesta de conservación. Cuadernos de Biodiversidad, 56: 26–47. <https://doi.org/10.14198/cdbio.2019.56.03>.
- Günther, A. (1912) The history of the collections contained in the natural history departments of the British Museum. Vol. II. Appendix. General history of the Department of Zoology from 1856 to 1895. London (The Trustees of the British Museum): i–ix + 1–109.
- Halffter, G. & Matthews, E. G. (1966) The natural history of dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Folia entomológica mexicana, 12–14: 1–312.
- Hancock, E. G., Pearce, N. & Campbell, M. (2015) William Hunter’s world: the art and science of eighteenth-century collecting. Farnham (Ashgate Publishing): i‒xxvii + 1‒392.
- Harold, E. (1859) Beitrage zur Kenntnis einiger coprophagen Lamellicornien. Berliner entomologische Zeitschrift, 3: 193–224. <https://doi.org/10.1002/mmnd.18590030208>.
- Harold, E., Le Baron de (1869) Révision des espèces qui rentrent dans le genre Pinotus Erichs. L’Abeille, 6: 123–144.
- Hernández, M. I. M., Silva, P. G., Niero, M. M., Alves, V. M., Bogoni, J. A., Brandl, A. L., Bugoni, A., Campos, R. C., Condé, P. A., Marcon, C. B., Simões, T., Terhorst, L. H. & Vaz-de-Mello, F. Z. (2019) Ecological characteristics of Atlantic Forest dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) in the state of Santa Catarina, southern Brazil. The Coleopterists Bulletin, 73 (3): 693–709. <https://doi.org/10.1649/0010-065X-73.3.693>.
- Heyne, A. & Taschenberg, O. (1908) Die exotischen Käfer in Wort und Bild. Esslingen & Munich (J. F. Schrieber): i–vii + 1–262 + [i], 39 pl.
- Hope, F. W. (1838) Observations on the Lamellicorns of Olivier. Entomological Magazine, 5: 312–326.
- Kerr, J. G. (1910) Remarks upon the Zoological Collection of the University of Glasgow, made on the occasion of the visit by the Natural History Society, on March 12, 1910. The Glasgow Naturalist, 2 (4): [97]–111.
- Lacordaire, J. T. (1855) Histoire naturelle des insectes. Genera des coléoptères, ou exposé méthodique et critique de tous les genres proposés jusqu’ici dans cet ordre d’insectes. Tome troisième. Contenant les familles des Pectinicornes et Lamellicornes. Paris (Librairie Encyclopédique de Roret): 1–594. [Publication date following Bousquet 2016]. <https://doi.org/10.5962/bhl.title.8864>.
- Lange, R. B. (1947) Ensaio da zoogeografia dos Scarabaeidae do Paraná com algumas notas eto-ecológicas. Arquivos do Museu paranaense, 6: 305‒315.
- Lima, J. D. N., Costa Silva, V., Bianchi, V., Silva, P. G. & Di Mare, R. A. (2015) Estrutura e organização de assembleias de Scarabaeinae (Coleoptera, Scarabaeidae) em diferentes fitofisionomias no sul do Brasil. Iheringia, Série Zoologia, 105 (4): 393‒402. <https://doi.org/10.1590/1678-476620151054393402>.
- Lima, R. C., Hoffmann, M. & Vicente, R. E. (2022) Community structure of dung beetles (Scarabaeidae: Scarabaeinae) in the Atlantic Forest of the Reserva Natural Vale (Linhares, Espírito Santo, Brazil). Biodiversidade brasileira, 12 (4): 1–15. <https://doi.org/10.37002/biobrasil.v12i4.2186>.
- Louzada, J. N. C. (1998) Considerations on the perching behavior of tropical dung beetles (Coleoptera, Scarabaeidae). Revista brasileira de Entomologia, 41 (2‒4): 125‒128.
- Louzada, J. N. C. & Carvalho e Silva, P. R. (2009) Utilisation of introduced Brazilian pastures ecosystems by native dung beetles: diversity patterns and resource use. Insect Conservation and Diversity, 2: 45–52. <https://doi.org/10.1111/j.1752-4598.2008.00038.x>.
- Louzada, J. N. C. & Lopes, F. S. (1997) A comunidade de Scarabaeidae copro-necrófagos (Coleoptera) de um fragmento de Mata Atlântica. Revista brasileira de Entomologia, 41 (1): 117‒121.
- Luederwaldt, H. (1929) As espécies brasileiras do gênero Pinotus (Coleoptera - Lamellicornidae - Coprini), com algumas considerações também sobre outras espécies. Revista do Museu paulista, 16: 603–776.
- Luederwaldt, H. (1931) O gênero Ontherus (Coleop.) (Lamellic. - Coprid. - Pinot.), com uma chave, para a determinação dos Pinotides americanos. Revista do Museu paulista, 17: [363]–422.
- Maldaner, M. E., Cupello, M., Ferreira, D. C. & Vaz-de-Mello, F. Z. (2017) Type specimens and names assigned to Coprophanaeus (Megaphanaeus) d’Olsoufieff, 1924, the largest New World dung beetles (Coleoptera: Scarabaeidae: Phanaeini). Zootaxa, 4272 (1): 83–102. <https://doi.org/10.11646/zootaxa.4272.1.4>.
- Maldaner, M. E., Valois, M. and Vaz-de-Mello, F. Z. (2018) A revision of Dichotomius (Homocanthonides) Luederwaldt, 1929 (Coleoptera: Scarabaeidae: Scarabaeinae). Revista brasileira de Entomologia, 62: 237‒242. <https://doi.org/10.1016/j.rbe.2018.05.001>.
- Martínez, A. (1951) Notas coleopterologicas. V. La invalidez del nombre generico Pinotus Erichson y dos nuevas sinonimias (Col. Scarab.). Anales de la Sociedad científica argentina, 92: 138–142.
- Martínez, A. (1959) Catálogo de los Scarabaeidae argentinos. Revista del Museo argentino de Ciencias naturales “Bernardino Rivadavia”, Ciencias Zoológicas, 5: 1–126.
- Martins, D. S., Ferreira, P. S. F., Fornazier, M. J. & Santos, J. S. (2014) Insetos da Reserva Natural Vale, Linhares, Espírito Santo, Brasil. Ciência & Ambiente, 49: [219]–235.
- Montreuil, O. (1998) Analyse phylogénétique et paraphylie des Coprini et Dichotomiini (Coleoptera: Scarabaeidae). Scénario biogéographique. Annales de la Société entomologique de France, 34 (2): 135‒148.
- O’Brian, P. (1988) Joseph Banks, a life. London (Collins Harvill): 1–328.
- Olivier, G. A. (1789a) Nº 3. Scarabé. Scarabaeus. • Olivier, G. A. (1789) Entomologie, ou histoire naturelle des insectes, avec leurs caractères génériques et spécifiques, leur description, leur synonymie, et leur figure enluminée. Coléoptères. Tome premier. Paris (Baudouin): 1–190. [Publication date following Bousquet 2016]. <https://doi.org/10.5962/bhl.title.61905>.
- Olivier, G. A. (1789b) Nº 3. Scarabé, Scarabaeus. Pl. XIV. • Olivier, G. A. (1789‒1795) Entomologie, ou histoire naturelle des insectes, avec leurs caractères génériques et spécifiques, leur description, leur synonymie, et leur figure enluminée. Coléoptères. Tome septième. Planches. Genres 1 à 65. Paris (Desray): pl. 14. [Publication date following Bousquet 2016. Note that Olivier (1789c) mentioned having delivered to “eager” subscribers “several illuminated plates” in August or September of presumably the previous year, 1788, before going to London to study its entomological collections. Even if the identities of these plates are eventually retrieved and new names are discovered to have been made available on them—thereby, changing their publication year from 1789 to 1788— this will certainly not be the case of plate XIV and Scarabaeus hesperus, for Olivier only discovered the species in London, thus, after delivering the first batch of plates he was referring to]. <https://doi.org/10.5962/bhl.title.49479>.
- Olivier, G. A. (1789c) Avertissement. • Olivier, G. A. (1789) Entomologie, ou histoire naturelle des insectes, avec leurs caractères génériques et spécifiques, leur description, leur synonymie, et leur figure enluminée. Coléoptères. Tome premier. Paris (Baudouin): [i]‒iv. [Publication date following Bousquet 2016]. <https://doi.org/10.5962/bhl.title.61905>.
- Olivier, G. A. (1790) [Livraison 41]: 1‒368. • Olivier, G. A. (1790‒1791) Encyclopédie méthodique, ou par ordre de matières. Histoire naturelle. Insectes. Tome cinquième. Paris (Panckoucke): 1–793. [Publication date following Bousquet 2016].
- Orchiston, W. (2016) Exploring the history of New Zealand astronomy. Trials, tribulations, telescopes and transits. Cham, Heidelberg, New York, Dordrecht & London (Springer): i–xlv + 1–688. <https://doi.org/10.1007/978-3-319-22566-1>.
- O’Shea, R. (1973) Type designations of some Fabrician Coreidae in the Banks collection at the British Museum (Natural History). Annals of the entomological Society of America, 66 (5): 1177‒1178. <https://doi.org/10.1093/aesa/66.5.1177>.
- Papavero, N. (1971) Essays on the history of Neotropical dipterology, with special reference to collectors (1750–1905). Vol. 1. São Paulo (Museu de Zoologia, Universidade de São Paulo): i‒vii + [1]‒216. <https://doi.org/10.5962/bhl.title.101715>.
- Papavero, N. (1994) Levantamento de localidades. • Papavero, N. (ed.) (1994) Fundamentos práticos de taxonomia zoológica (coleções, bibliografia, nomenclatura). São Paulo (Editora Unesp): 107‒119.
- Parkinson, S. (1773) A journal of a voyage to the south seas, in his majesty ship, the Endeavour. London (Stanfield Parkinson): 1–212 + [i‒ii]. [Sidney Parkinson died in 1771 during the Endeavour voyage. The notes he prepared during the voyage were later edited and published under the sponsorship of his brother Stanfield Parkinson as the book cited here].
- Pereira, F. S. & Halffter, G. (1961) Nuevos datos sobre Lamellicornia mexicanos con algunas observaciones sobre saprofagia. Revista brasileira de Entomologia, 10: 53–66.
- Pessôa, S. B. & Lane, F. (1941) Coleópteros necrófagos de interêsse médico-legal. Ensáio monográfico sôbre a família Scarabaeidae de S. Paulo e regiões vizinhas. Arquivos de Zoologia do Estado de São Paulo, 2: 389–504.
- Perty, J. A. M. (1830) Insecta Brasiliensia. • Perty, J. A. M. (1830–1833) Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCXVII-MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae regis augustissimi peracto collegerunt Dr. J. B. de Spix et Dr. C. F. Ph. de Martius. Munich [“Monachii”] (C. F. P. von Martius): 1–60, pl. 1–12. <https://doi.org/10.5962/bhl.title.102991>.
- Radford, W. P. K. (1981) The Fabrician types of the Australian and New Zealand Coleoptera in the Banks collection at the British Museum (Natural History). Records of the South Australian Museum, 18 (8): [155]–197.
- Rossini, M. and Vaz-de-Mello, F. Z. (2015) A review of the genus Chalcocopris Burmeister, 1846 (Coleoptera: Scarabaeidae: Scarabaeinae), with description of a new species. Zootaxa, 3920: 291–300. <https://doi.org/10.11646/zootaxa.3920.2.5>.
- Schiffler, G., Vaz-de-Mello, F. Z. & Azevedo, C. O. (2003) Scarabaeidae s. str. (Coleoptera) do delta do rio Doce e Vale do Suruaca no município de Linhares, estado do Espírito Santo, Brasil. Revista brasileira de Zoociências, 5 (2): 205‒211.
- Schubert, E. (2014) Olivier, Antoine Guillaume. • Anonymous (2014) Image collection. <https://sdei.senckenberg.de/photothek/>. [Accessed on 19th January 2013].
- Sekerka, L. & Barclay, M. V. L. (2014) Fabrician types of Cassidinae (Coleoptera: Chrysomelidae) deposited in the Natural History Museum, London. Acta entomologica Musei nationalis Pragae, 54 (2): 657‒684.
- Staig, R. A. (1931) The Fabrician types of insects in the Hunterian collection at Glasgow University. Coleoptera. Part I. Cambridge (Cambridge University Press): [i]‒xv + [1]–110, pl. 1‒28.
- Tarasov, S. & Dimitrov, D. (2016) Multigene phylogenetic analysis redefines dung beetles relationships and classification (Coleoptera: Scarabaeidae: Scarabaeinae). BMC evolutionary Biology, 16 (257): 1–19. <https://doi.org/10.1186/s12862-016-0822-x>.
- Tarasov, S. & Génier, F. (2015) Innovative Bayesian and parsimony phylogeny of dung beetles (Coleoptera, Scarabaeidae, Scarabaeinae) enhanced by ontology-based partitioning of morphological characters. PLoS One, 10 (3): 1–86. <https://doi.org/10.1371/journal.pone.0116671>.
- Tuxen, S. L. (1967) The entomologist, J. C. Fabricius. Annual Review of Entomology, 12: 1–14. <https://doi.org/10.1146/annurev.en.12.010167.000245>.
- Vaz-de-Mello, F. Z. & Cupello, M. (2018) The type specimens of South American dung beetles, Part I: On the species described in the genus Canthon Hoffmannsegg, 1817 (Coleoptera: Scarabaeidae: Scarabaeinae) by the German entomologist Adolf Schmidt (1856–1923). Spixiana, 41 (1): 33‒76.
- Vaz-de-Mello, F. Z., Edmonds, W. D., Ocampo, F. C. & Schoolmeesters, P. (2011) A multilingual key to the genera and subgenera of the subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae). Zootaxa, 2854: 1–73. <https://doi.org/10.11646/zootaxa.2854.1.1>.
- Waterhouse, C. O. (1906) Insecta. • Anonymous (ed.) (1906) The history of the collections contained in the natural history department of the British Museum. Vol. II. Separate historical accounts of the several collections included in the department of zoology. London (The Trustees of the British Museum): 551–653. [Note that the authorship of the Insecta part of the work is not given on its first page as done for the other groups, but was misplaced on page 579 following the heading of the subsection “Alphabetical list of the previous owners of collections [...]”].
- Wilkins, G. L. (1955) A catalogue and historical account of the Banks shell collection. Bulletin of the British Museum (Natural History), Historical Series, 1 (3): [69]–119, pl. 14–19.
- Whitehead, P. J. P. (1969) Zoological specimens from Captain Cook’s voyages. Journal of the Society for the Bibliography of natural History, 5 (3): 161–201. <https://doi.org/10.3366/jsbnh.1969.5.3.161>.
- Zimsen, E. (1964) The type material of I. C. Fabricius. Copenhagen (Munksgaard): [1]‒656.