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Published: 2021-06-03

Herpetological higher taxa nomina. 5. Anura, Urodela, Batrachia: addenda and corrigenda

Institut de Systématique, Évolution, Biodiversité, ISYEB – UMR 7205 – CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 30, 75005, Paris, France
Association RACINE, 5 allée des Cygnes, 35750 Saint Maugan, France
Availability authorship allocation priority validity hoplonym anoplonym haplorhizonym diplorhizonym Anuromorpha Anuromorphes Batrachi Batrachiacea Batrachiformes Batrachiformia Batrachoidacea Batrachoidaria Batrachoidiformes Batrachoidif


The nomenclatural status of 17 class-series nomina (13 diplorhizonyms and 4 haplorhizonyms), based on the single or primary stems Anura, Urodela and Batrachia that had been set aside in our previous papers of this series is examined. None of them is currently valid under the Duplostensional Nomenclatural System, four of them are unavailable (including "Batrachoidaria" Betancur-R. et al., 2017), and the status of four of them, based on fossil nucleogenera, is still unsettled because of missing information concerning the phylogenetic relationships of some of these genera.


  1. Abel, O. (1919) Die Stämme der Wirbelthiere. Berlin & Leipzig (Walter de Gruyter & Co.): i–xviii + 1–914.

    Amalitzky, V. P. (1921) Seymouridae. Petrograd (Akademii Nauk): 1–14. [In Russian; see Amalitzky (1924) for translation].

    Amalitzky, V. P. (1924) On a new Cotylosauria of the family Seymouridae from the Permian of North Russia. Annals and Magazine of natural History, (9), 13: 64–77. <>.

    Berg, L. S. (1940) Sistema ryboobraznykh i ryb, nyne zhivushchikh i iskopaemykh. [Classification of fishes, both recent and fossil]. Trudy zoologicheskogo Instituta, Akademiia Nauk SSSR, 5 (2): 87–345. [In Russian].

    Bertin, L. & Arambourg, C. (1958) Super-ordre des Téléostéens (Teleostei). In: P.-P. Grassé (ed.), Traité de zoologie, Agnathes et Poissons, Anatomie, systématique, biologie, Paris (Masson), 13 (3): 2204–2500.

    Betancur-R., R., Wiley, E. O., Arratia, G., Acero, A., Bailly, N., Miya, M., Lecointre, G. & Ortí, G. (2017) Phylogenetic classification of bony fishes. BMC evolutionary Biology, 17 (162): 1‒40 + Additional files 1‒6. <>.

    Blainville, H. de (1816) Prodrome d’une nouvelle distribution systématique du règne animal. Bulletin des Sciences de la Société philomatique de Paris, juillet 1816: ‘105–112’ [113–120] + 121–124.

    Bloch, M. E. & Schneider, J. G. (1801) Systema Ichthyologiae iconibus ex illustratum. Berolini (Sanderiano): i–lx + 1–584.

    Broili, F. (1904) Permische Stegocephalen und Reptilien aus Texas. Palaeontographica, 51: 1–48 + 49–121.

    Brongniart, A. (1800) Essai d’une classification naturelle des Reptiles. Iere partie. Établissement des ordres. Bulletin des Sciences, par la Société philomatique, 2 (35): 81–82.

    Cantino, P. D. & Queiroz, K. de (2020) International code of phylogenetic nomenclature (PhyloCode). Boca Raton, London & New York (CRC Press): i‒xl + 1‒149. <>.

    Cope, E. D. (1871) Contribution to the ichthyology of the Lesser Antilles. Transactions of the American philosophical Society, 14: 445‒483. <>.

    Cope, E. D. (1878) Descriptions of extinct Vertebrata from the Permian and Triassic formations of the United States. Proceedings of the American philosophical Society, 17: 182–193. <>.

    Dubois, A. (2005) Proposed Rules for the incorporation of nomina of higher-ranked zoological taxa in the International Code of Zoological Nomenclature. 1. Some general questions, concepts and terms of biological nomenclature. Zoosystema, 27 (2): 365–426. <>.

    Dubois, A. (2007) Naming taxa from cladograms: some confusions, misleading statements, and necessary clarifications. Cladistics, 23: 390–402. <>.

    Dubois, A. (2011) The International Code of Zoological Nomenclature must be drastically improved before it is too late. Bionomina, 2: 1–104. <>.

    Dubois, A. (2013) Zygoidy, a new nomenclatural concept. Bionomina, 6: 1–25. <>.

    Dubois, A. (2015) The Duplostensional Nomenclatural System for higher zoological nomenclature. Dumerilia, 5: 1–108.

    Dubois, A. (2020) Allocation of nomina to taxa in zoological nomenclature. Bionomina, 18: 1–43. <>.

    Dubois, A., Bauer, A. M., Ceríaco, L. M. P., Dusoulier, F., Frétey, T., Löbl, I., Lorvelec, O., Ohler, A., Stopiglia, R. & Aescht, E. (2019) The Linz Zoocode project: a set of new proposals regarding the terminology, the Principles and Rules of zoological nomenclature. First report of activities (2014‒2019). Bionomina, 17: 1–111. <>

    Dubois, A. & Frétey, T. (2020a) Herpetological higher taxa nomina. 1. Introduction and methodology. Bionomina, 19: 1–56. <>.

    Dubois, A. & Frétey, T. (2020b) Herpetological higher taxa nomina. 2. Anura Duméril, 1805. Bionomina, 20: 1–16. <>.

    Dubois, A. & Frétey, T. (2020c) Herpetological higher taxa nomina. 3. Urodela Duméril, 1805. Bionomina, 20: 17–35. <>.

    Dubois, A. & Frétey, T. (2020d) Herpetological higher taxa nomina. 4. Batrachia Brongniart, 1800. Bionomina, 20: 36–46. <>.

    Dubois, A. & Frétey, T. (2021) Herpetological higher taxa nomina. 6. Gymnophiona Rafinesque, 1814. Bionomina, 21: 39‒72. <>.

    Dubois, A. & Ohler, A. (2019) The nomina Anura, Urodela, Ecaudata and Caudata, credited to ‘Fischer von Waldheim, 1813’, do not exist, with comments on the nomenclature of higher zoological taxa and on the authorships and dates of other amphibian nomina. Bionomina, 14: 1‒68. <>.

    Dubois, A., Ohler, A. & Pyron, R. A. (2021) New concepts and methods for phylogenetic taxonomy and nomenclature in zoology, exemplified by a new ranked cladonomy of recent amphibians (Lissamphibia). Megataxa, 5: 1‒738.

    Duméril, A. M. C. (1805) Zoologie analytique, ou méthode naturelle de classification des animaux, rendue plus facile à l’aide de tableaux synoptiques. Paris (Allais), ‘1806’: i–xxxiii + 1–544. <>.

    Efremov, I. A. (1946) On the subclass Batrachosauria—an intermediary group between Amphibians and Reptiles. Doklady Akademii Nauk USSR, 1946 (6): 615‒638. [In Russian].

    Fitzinger, L. I. (1843) Systema Reptilium. Fasc. 1. Amblyglossae. Vindobonae (Braumüller & Seidel): 1–106 + i–ix.

    Gadow, H. (1898) A classification of Vertebrata, recent and extinct. London (Adam & Charles Black): i‒xvii + 1‒82. <>.

    Garsault, F. A. P., de (1764) Les Figures des plantes et animaux d’usage en médecine, décrits dans la Matière Médicale de Mr. Geoffroy Médecin. Paris (Mrs. Defehrt, Prevost, Duflos, Martinet &c.): [i–v] + 1–20, pl. 644–729. <>.

    Girard, C. F. (1854) Enumeration of the species of marine Fishes, collected at San Francisco, California, by Dr. C. B. R. Kennerly, naturalist attached to the survey of the Pacific R. R. route, under Lieut. A. W. Whipple. Proceedings of the Academy of natural Sciences of Philadelphia, 7: 141–142.

    Goodrich, E. S. (1909) Vertebrata Craniata (First fascicle: Cyclostomes and Fishes). In: R. Lankaster, A treatise on zoology, London (Adam and Charles Black), 9: i–xvi + 1–518. <>.

    Goodrich, E. S. (1930) Studies on the structure & development of Vertebrates. London (Macmillan and co.): i–xxx + 1–837. <>.

    Gray, J. E. (1842) The northern zoological gallery. In: Synopsis of the contents of the British Museum, 44th edition, London (British Museum): 97–157.

    Huene, F. F. von (1946) Die großen Stämme der Tetrapoden in den geologischen Zeiten. Biologisches Zentralblatt, 65 (7): 266–275.

    Huene, F. R. von (1948a) Ein Versuch, die paläozoischen Urodeloidea zu entwirren. Neues Jahrbuch für Mineralogie, Geologie und Paläontologie, (B), 1945–1948: 111–126.

    Huene, F. R. von (1948b) Short review of the lower tetrapods. In: A. L. Du Toit, Robert Broom commemorative volume, Special publication of the royal Society of South Africa, Cape Town (Royal Society of South Africa): 65–106.

    Huene, F. R. von (1952) Die Saurierwelt und ihre geschichtlichen Zusammenhänge. Jena (Fischer): 1–64.

    Huene, F. R. von (1956) Paläontologie und Phylogenie der niederen Tetrapoden. Jena (Gustav Fischer Verlag): i–xii + 1–716 + [i–ii], 1 tab.

    Huxley, T. H. (1863) Description of Anthracosaurus russelli, a new labyrinthodont from the Lanarkshire coal-field. The quarterly Journal of the geological Society of London, 19 (1): 56–68. <>.

    Kuhn, O. (1959) Die Ordnungen der fossilen ‘Amphibien’ und ‘Reptilien’. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte, 1959: 337–347.

    Kuhn, O. (1961) Die Familien der rezenten und fossilen Amphibien. Bamberg (Verlagshaus Meisenbach KG): 1–79.

    Kuhn, O. (1967) Namensänderungen bei einigen höheren Taxa der Amphibia und Reptilia. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte, 1967: 185–187.

    La Cepède, [B. G. E.] (1800) Histoire naturelle des poissons. Volume 2. Paris (Plassan): i–lxiv + 1–632, pl. 1–20.

    Latreille, [P. A.] (1825) Familles naturelles du règne animal. Paris (Baillière): [i–v] + 1–570.

    Laurenti, J. N. (1768) Specimen medicum, exhibens synopsin Reptilium emendatam cum experimentis circa venena et antidota Reptilium austriacorum. Viennae (Joan. Thom. Nob. de Trattnern): i–ii + 1–215, pl. 1–5. <>.

    Leuckart, F. S. (1840) Einige zoologische Bemerkungen. 2. Ueber das Gen. Cryptobranchus. Neue Notizen aus dem Gebiete der Natur- und Heilkunde, 13 (2): 19–20.

    Lydekker, R. (1889) Part III. Palaeozoology. Vertebrata. In: H.A. Nicholson & R. Lydekker (ed.), A manual of palaeontology for the use of students, with a general introduction on the principles of palaeontology, Volume 2, Edinburgh & London (William Blackwood & Sons): i–xi + 887–1624.

    Marjanović, D. & Laurin, M. (2019) Phylogeny of Paleozoic limbed vertebrates reassessed through revision and expansion of the largest published relevant data matrix. PeerJ, 6 [e5565]: 1–191. <>.

    Merrem, B. (1820) Versuch eines Systems der Amphibien. Tentamen systematis Amphibiorum. Marburg (Iohann Christian Krieger): [i–xv + 1–191] × 2, 1 pl. <>.

    Miranda-Ribeiro, A. de (1924) Notas batrachologicas. I. Boletim do Museu nacional do Rio de Janeiro, 2: 137–143.

    Oppel, M. (1811) Ordre II. Reptiles à écailles. Section 2. Ophidiens. Annales du Muséum d’Histoire naturelle, ‘1810’, 16: 254–295 + 376–393.

    Piveteau, J. (1937) Paléontologie de Madagascar. XXIII. Un Amphibien du Trias inférieur: essai sur l’origine et l’évolution des Amphibiens Anoures. Annales de Paléontologie, 26: 135–178, pl. 19–20.

    Přikryl, T. & Carnevale, G. (2017) An Oligocene toadfish (Teleostei, Percomorpha) from Moravia, Czech Republic: the earliest skeletal record for the order Batrachoidiformes. Bulletin of Geosciences, 92 (1): 123–131. <>.

    Queiroz, K. de, Cantino, P. D. & Gauthier, J. A. (2020) Phylonyms. A companion to the PhyloCode. Boca Raton, London & New York (CRC Press): i‒xxvii + 1‒1324. <>.

    Rafinesque, C. S. (1814) Fine del Prodromo d’Erpetologia siciliana. Specchio delle Scienze o Giornale enciclopedico di Sicilia, 2 (10): 102–104.

    Regan, C. T. (1912) The classification of the teleostean fishes of the order Pediculati. Annals and Magazine of natural History, (8), 9 (51): 277‒289. <>.

    Säve-Söderbergh, G. (1932) Preliminary note on Devonian stegocephalians from East Greenland. Meddelelser om Grønland, 94 (7): 1–109, pl. 1–22.

    Säve-Söderbergh, G. (1934) Some points of view concerning the evolution of the Vertebrates and the classification of this group. Arkiv för Zoologi, 26A (17): 1–20.

    Säve-Söderbergh, G. (1935) On the dermal bones of the head in labyrinthodont Stegocephalians and primitive Reptilia, with special reference to eotriassic Stegocephalians from East Greenland. Meddelelser om Grønland, 98 (3), 1–211: pl. 1–15.

    Schaeffer, I. C. (1760) De studii ichthyologici faciliori ac tutiori methodo. Ratisbonae (Typis Weissianis et impensis Montagii): 1–24, 1 pl.

    Taylor, E. H. & Hesse, C. J. (1943) A new salamander from the upper Miocene beds of San Jacinto County, Texas. American Journal of Science, 241: 185–193. <>.

    Van der Hoeven, J. (1855) Handboek der dierkunde. Tweede, vrbeterde en vermeerderde Uitgave. Tweede deel. Amsterdam (J. C. A. Sulpke): i–xxviii + 1–1068, pl. 1–24.

    Zittel, K. A. (1888) Handbuch der Palaeontologie. I Abtheilung. Palaeozoologie. III Band. Vertebrata (Pisces, Amphibia, Reptilia, Aves). Second instalment. München & Leipzig (R. Oldenbourg): 257–436.