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Article
Published: 2021-06-03

Herpetological higher taxa nomina. 6. Gymnophiona Rafinesque, 1814

Institut de Systématique, Évolution, Biodiversité, ISYEB—UMR 7205—CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 30, 75005, Paris, France
Association RACINE, 5 allée des Cygnes, 35750 Saint Maugan, France
Availability authorship allocation priority validity hoplonym anoplonym eikomorph distagmomorph distagmonym sozomorph sozodiaphonym nothosozonym ectomorph ectonym caecilians Amphibia Apoda Apodacea Apodes Apodi Apodia Apodida Apodif

Abstract

A taxon, traditionally referred to the rank order, encompassing all recent taxa of caecilians and their close fossil relatives, is highly supported as holophyletic in all recent cladistic analyses of Amphibia. Under the Duplostensional Nomenclatural System, among 12 nomina available in the literature, only one, Gymnophiona Rafinesque, 1814, qualifies as a sozodiaphonym and should be used for this amphibian taxon. We show here that the authorship of this nomen, as well as of 32 other nomina published in 1814, is ‘Rafinesque’, not ‘Rafinesque-Schmaltz’.

            The nomen Apoda Oppel, 1811, that has often been used for this taxon, proves to be a distagmonym and a junior homonym of Apodes Linnaeus, 1758 and five other distagmonyms, and as such it is invalid under DONS.

            In total, we identified 34 distinct class-series nomina derived from the stems ἄπους (apous) or Apus in 13 distinct zoological groups, only two of which are valid under DONS Criteria: Apodiformia Wetmore, 1947, for the suborder of Aves currently known as Apodi Wetmore, 1947, and Apodomorpha Sibley, Ahlquist & Monroe, 1988 for the order of Aves currently known as Apodiformes Peters, 1940.

                Several other nomenclatural findings were made in the course of this study: [1] in the Teleostei, the nomina "Eupercaria" and "Syngnatharia", like all the other new nomina proposed in the paper by Betancur-R. et al. (2017) are anoplonyms and cannot be used as valid; [2] in the Mammalia, the nomina Cetus and Cetacea should be credited to Brisson (1759); [3] in the Holothuroidea, the nomen Apoda Claus, 1868, currently considered valid under the apograph Apodida, is invalid and should be replaced by the nomen Paractinopoda Ludwig, 1891; [4] in the Isopoda, the subordinal nomen Cymothoida Wägele, 1989 should be replaced by its senior synonym Darwinida Lakshminarayana & Rama Rao, 1977.

References

  1. Anonymous [International Commission on Zoological Nomenclature] (1998) Opinion 1894. Regnum Animale ..., Ed. 2 (M. J. Brisson, 1762): rejected for nomenclatural purposes, with the conservation of the mammalian generic names Philander (Marsupialia), Pteropus (Chiroptera), Glis, Cuniculus and Hydrochoerus (Rodentia), Meles, Lutra and Hyaena (Carnivora), Tapirus (Perissodactyla), Tragulus and Giraffa (Artiodactyla). Bulletin of zoological Nomenclature, 55 (1): 64‒71.

    Anonymous [International Commission on Zoological Nomenclature] (2005) Opinion 2104 (Case 3226). Lacepède, B. G. E. de la V., 1788, Histoire naturelle des Quadrupèdes Ovipares: rejected as a non-binominal work. Bulletin of zoological Nomenclature, 62 (1): 55.

    Acosta-Galvis, A. R., Torres, M. & Pulido-Santacruz, P. (2019) A new species of Caecilia (Gymnophiona, Caeciliidae) from the Magdalena valley region of Colombia. Zookeys, 884: 135–157. <https://doi.org/10.3897/zookeys.884.35776>.

    Adams, A., Baikie, W. B. & Barron, C. (1854) A manual of natural history for the use of travellers; being a description of the families of the animal and vegetable kingdoms: with remarks on the practical study of geology and meteorology to which are appended directions for collecting and preserving. London (John van Voorst): i–viii + 1–749. <https://doi.org/10.5962/bhl.title.63983>.

    Betancur-R., R., Wiley, E. O., Arratia, G., Acero, A., Bailly, N., Miya, M., Lecointre, G. & Ortí, G. (2017) Phylogenetic classification of bony fishes. BMC evolutionary Biology, 17 (162): 1‒40 + Additional files 1‒6. <https://doi.org/10.1186/s12862-017-0958-3>.

    Billberg, G. J. (1827) Synopsis faunae scandinaviae. Tom. I. Pars 1. Mammalia. Holmiae (Officina typogr. Ordinum Equestrium): i–viii + 1–55 + i–xv + [i–iii], 1 pl. <https://doi.org/10.5962/bhl.title.69294>.

    Blainville, H. de (1816) Prodrome d’une nouvelle distribution systématique du règne animal. Bulletin des Sciences de la Société philomatique de Paris, juillet 1816: ‘105–112’ [113–120] + 121–124.

    Bocquet-Védrine, J. (1972) Suppression de l’ordre des Apodes (Crustacés Cirripèdes) et rattachement de son unique représentant, Proteolepas bivincta, à la famille des Crinoniscidae (Crustacés Isopodes, Cryptonisciens). Comptes rendus de l’Académie des Sciences, (D), 275: 2145–2148, pl. 1.

    Bonnier, J. (1900) Contribution à l’étude des Épicarides. Les Bopyridae. Travaux de la Station zoologique de Wimereux, 8: 1–476, pl. 1–41. <https://doi.org/10.5962/bhl.title.10522>.

    Brisson A. D. (1756) Regnum animale in classes IX distributum sive synopsis methodica sistens generale m animalium distributionem in Classes IX, & duarum primarum Classium, Quadrupedum scilicet & Cetaceorum, particularem divisionem in ordines, sectiones, genera & species. Cum brevi cujusque speciei descriptione , citationibus auctorum de iis tractantium, nominibus eis ab ipsis & nationibus impositis, nominibusque vulgaribus. Le règne animal divisé en IX classes, ou méthode contenant la division générale des animaux en ix classes , & la division particulière des deux premières Classes, sçavoir de celle des Quadrupèdes & de celle des Cétacées , en ordres, sections, genres & espèces. Auxquelles on a joint une courte description de chaque espéce, avec les citations des auteurs qui en ont traité, les noms quʼils leurs ont donnés, ceux que leurs ont donnés les différentes nations, & les noms vulgaires. Paris (Jean-Baptiste Bauche): i–vii + 1–382, 1 tab., 1 pl. <https://doi.org/10.5962/bhl.title.70840>.

    Brisson [A. D.] (1759) Division générale du règne animal. In: Anonymous [M. D. L. C. D. B.], Dictionnaire raisonné et universel des animaux ou le règne animal …, Tome quatrième, Paris (Bauche): 625‒632.

    Brisson, A. D. (1762) Regnum animale in classes IX. Distributum, sive synopsis methodica Siftens generalem Animalium distributionem in Classes IX, & duarum primarum Classium, Quadrupedum scilicet & Cetaceorum, particularem divisionem in Ordines, Sectiones, Genera & Species. Cum brevi cujusque speciei descriptione, citationibus auctorum de iis tractantium, nominibus eis ab ipsis & nationibus impositis, nominibusque vulgaribus. Lugduni Batavorum (Theodorum Haak): [i–vi] + 1–296. <https://doi.org/10.5962/bhl.title.40361>.

    Brodkorb, P. (1940) Ornithological literature. Check-list of birds of the world, Volume 4. By James Lee Peters. Harvard. University Press, Cambridge, Mass., 1940: 6 × 9 in., xii + 291 pp. The Wilson Bulletin, 52 (3): 214–215.

    Brown, T. (1832) The zoologist’s text-book, embracing the characters of the classes, orders, and genera, of almost the whole Animal kingdom; together with an example of a species of nearly each genus, and a complete glossary of technical terms. Glasgow (Fullarton & Co.), 1: i–xii + 13–578. <https://doi.org/10.5962/bhl.title.11641>.

    Cannatella, D. C. (2007) An integrative phylogeny of Amphibia. In: P. M. Narins, A. S. Feng, R. R. Fay & A. N. Popper (ed.), Hearing and sound communication in amphibians, New York (Springer): 12–43. <https://doi.org/10.1007/978-0-387-47796-1_2>.

    Cannatella, D. C. & Hillis, D. M. (2004) Amphibians: leading a life of slime. In: J. Cracraft & M. J. Donoghue (ed.), Assembling the tree of life, New York (Columbia University Press): 430‒450.

    Cantino, P. D. & Queiroz, K. de (2020) International code of phylogenetic nomenclature (PhyloCode). Boca Raton, London & New York (CRC Press): i‒xl + 1‒149. <https://doi.org/10.1201/9780429446320>.

    Claus, C. (1868) Grundzüge der Zoologie zum Gebrauche an Universitäten und höhern Lehranstalten. Leitfaden zur Einführung in das wissenschaftliche Studium der Zoologie. Marburg und Leipzig (N. G. Elwertʼsche Universitäts-Buchhandlung): i–viii + 1–840. <https://doi.org/10.5962/bhl.title.110152>.

    Cope, E. D. (1889) Synopsis of the families Vertebrata. American Naturalist, 23 (274): 849–877. <https://doi.org/10.1086/275018>.

    Cuénot, L. (1891) Études morphologiques sur les Échinodermes. Archives de Biologie, 11 (3–4): 313–680, pl. 24–31.

    Cuvier, G. (1816) Le règne animal distribué d’après son organisation, pour servir de base à l’histoire naturelle des animaux et d’introduction à l’anatomie comparée. Paris (Deterville), ‘1817’, 2: i–xviii + 1–532.

    Darwin, C. (1854) A·monograph on the subclass Cirripedia of all the species. London (Ray Society): i–viii + 1–684.

    Desvaux, N. A. (1815) Sur les ouvrages de M. Rafinesque-Schmaltz. Journal de Botanique, appliquée à l’Agriculture, à la Pharmacie, à la Médecine et aux Arts, ‘1814’, 4: 268‒279.

    Dubois, A. (1984) Miscellanea nomenclatorica batrachologica (V). Alytes, 3 (3): 111–116.

    Dubois, A. (2009) Incorporation of nomina of higher-ranked taxa into the International Code of Zoological Nomenclature: the nomenclatural status of class-series zoological nomina published in a non-latinized form. Zootaxa, 2106: 1–12. <https://doi.org/10.11646/zootaxa.2106.1.1>.

    Dubois, A. (2015) The Duplostensional Nomenclatural System for higher zoological nomenclature. Dumerilia, 5: 1–108.

    Dubois, A. & Frétey, T. (2020) Herpetological higher taxa nomina. 1. Introduction and methodology. Bionomina, 19: 1–56. <https://doi.org/10.11646/bionomina.19.1.1>.

    Dubois, A. & Ohler, A. (2019) The nomina Anura, Urodela, Ecaudata and Caudata, credited to ‘Fischer von Waldheim, 1813’, do not exist, with comments on the nomenclature of higher zoological taxa and on the authorships and dates of other amphibian nomina. Bionomina, 14: 1‒68. <https://doi.org/10.11646/bionomina.14.1.1>.

    Dubois, A., Ohler, A. & Pyron, R. A. (2021) New concepts and methods for phylogenetic taxonomy and nomenclature in zoology, exemplified by a new ranked cladonomy of recent amphibians (Lissamphibia). Megataxa, 5: 1‒738.

    Dubois, A. & Raffaëlli, J. (2012) A new ergotaxonomy of the order Urodela Duméril, 1805 (Amphibia, Batrachia). Alytes, 28 (3–4): 77–161.

    Ducrotay de Blainville, H. M. (1822) De l’organisation des animaux, ou Principes d’anatomie comparée. Tome premier, contenant la morphologie et l’aistésologie. Paris (Levrault): [i–iii] + i–lix + 1–574, 10 tab. <https://doi.org/10.5962/bhl.title.110421>.

    Duméril A. M. C. (1805) Zoologie analytique, ou méthode naturelle de classification des animaux, rendue plus facile à l’aide de tableaux synoptiques. Paris (Allais), ‘1806’: i–xxxiii + 1–544. <https://doi.org/10.5962/bhl.title.44835>.

    Duméril (1837) Rapport sur un ouvrage manuscrit de M. le docteur Cocteau, ayant pour titre: Tabulæ synopticæ Scincoideorum. Comptes rendus hebdomadaires des Séances de lʼAcadémie des Sciences, 4: 14–17.

    Dundee, H. A. (1989) Higher category name usage for Amphibians and Reptiles. Systematic Zoology, 38: 398‒406. <https://doi.org/10.2307/2992405>.

    Dupuis, C. (1995) Contre-propositions pour inscrire les ouvrages de Brisson (1759) et (1762) dans la Liste officielle. In: Comments on the proposed conservation of some mammal generic names first published in Brisson’s (1762) Regnum Animale, Bulletin of zoological Nomenclature, 52 (3): 273–275. <https://doi.org/10.5962/bhl.part.6795>.

    Eaton, A. (1826) Zoological text-book comprising Cuvierʼs four grand divisions of animals: also, Shawʼs improved Linnean genera, arranged according to the classes and orders of Cuvier and Latreille. Short descriptions of some of the most common species are given for studentʼs exercises. Albany (Websters & Skinners): i–vi + 7–288. <https://doi.org/10.5962/bhl.title.42226>.

    Ellsworth Call, R. (1895) The life and writings of Rafinesque. Louisville (John P. Morton): [i] + 1‒227.

    Erxleben, I. C. (1767) Dissertatio inauguralis physica sistens diiudicationem systematum animalium mammalium. Gottingae (Schulzianus): [i–ii] + 1–15.

    Fischer, G. (1808) Tableaux synoptiques de zoognosie. Moscou (Imprimerie de l’Université Impériale): [1–60] + 1–186, 6 pl.

    Frost D. R., Grant, T., Faivovich, J., Bazin, R. H., Haas, A., Haddad, C. F. B., Sá, R. O. de, Channing, A., Wilkinson, M., Donnellan, S. C., Raxworthy, C. J., Campbell, J. A., Blotto, B. L., Moler, P., Drewes, R. C., Nussbaum, R. A., Lynch, J. D., Green, D. M., Wheeler, W. C. (2006) The amphibian tree of life. Bulletin of the American Museum of natural History, 297: 1–370. <https://doi.org/10.1206/0003-0090(2006)297[0001:TATOL]2.0.CO;2>.

    Garrod, A. H. (1874) On certain muscles of birds and their value in classification. Part II. Proceedings of the zoological Society of London, 1874: 111–123, pl. 17. <https://doi.org/10.1111/j.1096-3642.1874.tb02459.x>.

    Gill, T. (1893) Families and subfamilies of fishes. Memoirs of the national Academy of Sciences, 6: 125–138. <https://doi.org/10.5962/bhl.part.6303>.

    Goldfuss, G. A. (1820) Handbuch der Zoologie. Dritter Theil, erste Abtheilung. Nürnberg (Johann Leonhard Schrag): i–xlvi + 1–696, pl. 1–2.

    Hartert, E. (1897) Aves. Podargidae, Caprimulgidae and Macropterygidae. Das Tierreich, 1: i–viii + i–iv + 1–98.

    Haworth, A. H. (1809) Lepidoptera britannica, sistens digestionem novam Insectorum Lepidopterorum quae in magna Britannia reperiuntur, larvarum pabulo, temporeque pascendi; expansione a/arum, mensibusque vo/andi, synonymis atque locis observationibusque variis. [Part 2]. London (J. Murray): 137–376.

    Hyžný, M., Pasini, G. & Garassino, A. (2019) Supergiants in Europe: on the cirolanid isopod Bathynomus A. Milne Edwards, 1879 (Malacostraca, Peracarida) from the Plio-Pleistocene of Italy. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen, 291 (3): 283–298. <https://doi.org/10.1127/njgpa/2019/0802>.

    Illiger, C. (1811) Prodromus systematis Mammalium et Avium additis terminis zoographicis utriusque classis, sorumque versione germanica. Berolini (C. Salfeld): i–xviii + 1–301, errata. <https://doi.org/10.5962/bhl.title.106965>.

    Irisarri, I., San Mauro, D., Abascal, F., Ohler, A., Vences, M. & Zardoya, R. (2012) The origin of modern frogs (Neobatrachia) was accompanied by acceleration in mitochondrial and nuclear substitution rates. BMC Genomics, 13 (626): 1–19. <https://doi.org/10.1186/1471-2164-13-626>.

    Kaup, [J. J.] (1856) Catalogue of Apodal fish in the collection of the British Museum. London (Trustees): i–viii + 1–163, pl. 1–19. [Probably published in 1857, as the preface of J. E. Gray is dated 30 December 1856].

    Lakshminarayana, K. V. & Rama Rao, K. V. (1977) On the problem of homonymy between supra-ordinal, ordinal, infra-ordinal names in zoology. II. (Apoda in Amphibia, Echinodermata, Crustacea; and Apodes in Crustacea and Pisces). Newsletter, Zoological Survey India, 3 (4): 160–165.

    Lamarck, [J.-B.] de (1818) Histoire naturelle des Animaux sans Vertèbres, présentant les caractères généraux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales espèces qui sʼy rapportent; précédée dʼune introduction offrant la determination des caractères essentiels de lʼAnimal, sa distinction du végétal et des autres corps naturels, enfin, lʼexposition des principes fondamentaux de la Zoologie. Tome 5. Paris (Déterville & Verdière): 1–612.

    Latreille, P. A. (1804) Tableau méthodique des poissons. In: Tableaux méthodiques d’histoire naturelle; in: Nouveau dictionnaire d’histoire naturelle, appliqué aux arts, principalement à l’agriculture et à l’économie rurale et domestique, Tome 24, Paris (Deterville): 71–105.

    Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Tomus 1. Holmiae (Laurentii Salvii): [i–iv] + 1–824. <https://doi.org/10.5962/bhl.title.542>.

    Livezey, B. C. & Zusi, R. L. (2007) Higher-order phylogeny of modern birds (Theropoda, Aves: Neornithes) based on comparative anatomy. II. Analysis and discussion. Zoological Journal of the Linnean Society, 149 (1): 1–95. <https://doi.org/10.1111/j.1096-3642.2006.00293.x>.

    Ludwig, H. (1891) Die Seewalzen. In: H. G., Bronn (ed.) Klassen und Ordnungen des Thier-Reichs, wissenschaftlich dargestellt in Wort und Bild, Zweiter Band, Dritte Abtheilung: Echinodermen (Stachelhäuter), I. Buch, Leipzig (C. F. Winter’sche Verlagshandlung): 240–376, pl. 13–17.

    Lütken, C. F. (1857) Oversigt over Grönlands Echinodermata samt over denne Dyreklasses geographiske og bathymetriske Udbredningsforhold i de nordiske Have. Kjöbenhavn (C. A. Reitzel): i + 1–109, 1 pl. <https://doi.org/10.5962/bhl.title.14858>.

    MacLeay, W. S. (1840) Note on the Annelida. Annals and Magazine of natural History, (1), 4: 385–388. <https://doi.org/10.1080/00222934009512510>.

    Marjanović, D. & Laurin, M. (2008) A reevaluation of the evidence supporting an unorthodox hypothesis on the origin of extant amphibians. Contributions to Zoology, 77 (3): 149–199. <https://doi.org/10.1163/18759866-07703002>.

    Merrem, B. (1820) Versuch eines Systems der Amphibien. Tentamen systematis Amphibiorum. Marburg (Iohann Christian Krieger): [i–xv + 1–191] × 2, 1 pl. <https://doi.org/10.5962/bhl.title.5037>.

    Meyer & Wolf (1810) Taschenbuch der deutschen Vögelkunde oder kurze Beschreibung aller Vögel Deutschlands. Esrter Theil die Landvögel enthaltend. Frankfurt am Main (Friedrich Wilmans): [i–iv] + i–xviii + 1–310, 40 pl. <https://doi.org/10.5962/bhl.title.66969>.

    Müller, J. (1832) Beiträge zur Anatomie und Naturgeschichte der Amphibien. Zeitschrift für Physiologie, 4 (2): 190–275, pl. 18–22.

    Oppel, M. (1811) Second mémoire sur la classification des Reptiles. Annales du Muséum d’Histoire naturelle, 16: 394–418.

    Pearse, A. S. (ed.) (1936) Zoological names. A list of phyla, classes, and orders. Durham, North Carolina (Duke University Press): 1‒24.

    Peters, J. L. (1940) Check-list of birds of the world. Cambridge (Harvard University Press), 4: i–xii + 1–291.

    Pyron, R. A. & Wiens, J. J. (2011) A large-scale phylogeny of Amphibia including over 2800 species, and a revised classification of extant frogs, salamanders and caecilians. Molecular Phylogenetics and Evolution, 61: 543–583. <https://doi.org/10.1016/j.ympev.2011.06.012>.

    Queiroz, K. de, Cantino, P. D. & Gauthier, J. A. (2020) Phylonyms. A companion to the PhyloCode. Boca Raton, London & New York (CRC Press): i‒xxvii + 1‒1324. <https://doi.org/10.1201/9780429446276>.

    Queiroz, K. de & Gauthier, J. (1992) Phylogenetic taxonomy. Annual Review of Ecology and Systematics, 23: 449–480. <https://doi.org/10.1146/annurev.es.23.110192.002313>.

    Rafinesque, C. S. (1814d) Prodrono [sic] di erpetologia siciliana. Specchio delle Scienze o Giornale enciclopedico di Sicilia, 2 (9): 65‒67.

    Rafinesque, C. S. (1814e) Fine del prodromo d’erpetologia siciliana. Specchio delle Scienze o Giornale enciclopedico di Sicilia, 2 (10): 102‒104.

    Rafinesque, C. S. (1815) Analyse de la nature ou tableau de l’univers et des corps organisés. Palerme (Jean Barravecchia): 1–124, 1 pl. <https://doi.org/10.5962/bhl.title.106607>.

    Rafinesque, [C. S.] (1845) Specchio delle Scienze o Giornale enciclopedico di Sicilia; deposito letterario delle moderne cognizioni, scoperte ed osservazioni sopra le scienze ed arti. Isis von Oken, 38: 222–230.

    Rafinesque-Schmaltz, C. S. (1814a) O quadro del metodo sinottico di somiologia. Specchio delle Scienze o Giornale enciclopedico di Sicilia, 1 (1): 11‒15.

    Rafinesque-Schmaltz, C. S. (1814b) Précis des découvertes et travaux somiologiques de Mr C. S. Rafinesque-Schmaltz entre 1800 et 1814, ou choix raisonné de ses principales découvertes en zoologie et en botanique, pour servir d’introduction à ses ouvrages futurs. Palerme (Royale Typographie Militaire, aux dépens de l’auteur): 1–56. <https://doi.org/10.5962/bhl.title.6135>.

    Rafinesque-Schmaltz, C. S. (1814c) Principes fondamentaux de somiologie ou les loix [sic] de la nomenclature et de la classification de l’empire organique, ou des animaux et des végétaux, contenant les règles essentielles de l’art de leur imposer des noms immuables et de les classer méthodiquement. Palerme (Franc. Abate): 1–52.

    Roelants, K. & Bossuyt, F. (2005) Archaeobatrachian paraphyly and pangaean diversification of crown-group frogs. Systematic Biology, 54 (1): 111–126. <https://doi.org/10.1080/10635150590905894>.

    Ruggiero, M. A., Gordon, D. P, Orrell, T. M., Bailly, N., Bourgoin, T., Brusca, R. C., Cavalier-Smith, T., Guiry, M. D. & Kirk, P. M. (2015) A higher level classification of all living organisms. PloS One, 10 (4) [e0119248]: 1‒60 + S1 Appendix (1‒12) + S1 Table. <https://doi.org/10.1371/journal.pone.0119248>.

    San Mauro, D. (2010) A multilocus timescale for the origin of extant amphibians. Molecular Phylogenetics & Evolution, 56: 554‒561. <https://doi.org/10.1016/j.ympev.2010.04.019>.

    Scopoli, I. A. (1777) Introductio ad historiam naturalem, sistens genera lapidum, plantarum, edt animalium hactenus detecta, caracteribus essentialibus donata, in tribus divisa, subinde ad leges naturae. Pragae (Gerle): [i–x] + 1–506 + i–xxxvi. <https://doi.org/10.5962/bhl.title.10827>.

    Selenka, E. (1867) Beiträge zur Anatomie und Systematik der Holothurien, Zeitschrift für wissenschaftliche Zoologie, 17: 291–374, pl. 17–20.

    Semper, C. (1868) Reisen im Archipel der Philippinen. Zweiter Theil. Wissenschafliche Resultate. Erster Band. Holothurien. Leipzig (Wilhelm Engelmann): i–iv + 1–288. <https://doi.org/10.5962/bhl.title.11687>.

    Shipunov, A. (2011) The problem of hemihomonyms and the on-line hemihomonyms database (HHDB). Bionomina, 4: 65–72. <https://doi.org/10.11646/bionomina.4.1.3>.

    Sibley, C. G. & Ahlquist, J. E. (1990) Phylogeny and classification of birds. A study in molecular evolution. New Haven (Yale University Press): i–xiii + 1–976. <https://doi.org/10.2307/j.ctt1xp3v3r>.

    Sibley, C. G., Ahlquist, J. E. & Monroe, B. L., Jr. (1988) Classification of the living birds of the world based on DNA-DNA hybridization studies. The Auk, 105 (3): 409–423. <https://doi.org/10.1093/auk/105.3.409>.

    Spagnesi, M. & Serra, L. (ed.) (2003) Uccelli d’Italia: Gruiformes, Charadriiformes, Pterocliformes, Columbiformes, Cuculiformes, Strigiformes, Caprimulgiformes, Apodiformes, Coraciiformes, Piciformes. Istituto nazionale per la fauna selvatica Alessandro Ghigi: 1–265.

    Starobogatov Y. I. (1984) O problemakh nomenklatury vysshikh taksonomicheskikh kategoriy. In: L. P. Tatarinov & V. N. Shimanskiy (ed.), Spravochnik po sistematike iskopayemykh organbizmov (taksony otryadnoy i vyshchikh grupp), Moscow (Izdatel’svo Nauka): 174–187. [In Russian; not seen; quoted after Starobogatov 1991).

    Starobogatov, Y. I. (1991) Problems in the nomenclature of higher taxonomic categories. Bulletin of zoological Nomenclature, 48 (1): 6–18. [English translation of Starobogatov 1984]. <https://doi.org/10.5962/bhl.part.663>.

    Théel, H. (1886) Report on the Holothurioidea dredged by H.M.S. Challenger, during the years 1873-1876. Part II. In: C. W. Thomson & J. Murray, Report on the scientific results of the voyage of H.M.S. Challenger during the years 1873–76, Zoology, London (Longmans & Co.), 14 (39): 1–290, pl. 1–16.

    Vigors, N. A. (1825) Observations on the natural affinities that connect the orders and families of birds. Transactions of the Linnean Society of London, 14 (3): 395–517. <https://doi.org/10.1111/j.1095-8339.1823.tb00098.x>.

    Wägele, J. W. (1989) Evolution und phylogenetisches System der Isopoda. Zoologica, 140: 1–262.

    Wetmore, A. (1934) A systematic classification for the birds of the world, revised and amended. Smithsonian miscellaneous Collections, 89 (13): 1–11.

    Wetmore, A. (1947) Nomenclature of the higher groups in swifts. The Wilson Bulletin, 59 (4): 211–212.

    Wetmore, A. & Miller, W. D. (1926) The revised classification for the fourth edition of the A. O. U. check-list. The Auk, 43 (3): 337–346. <https://doi.org/10.2307/4075427>.

    Wilkinson, M. & Nussbaum, R. A. (2006) Caecilian phylogeny and classification. In: J.-M. Exbrayat (ed.), Reproductive biology and phylogeny of Gymnophiona (caecilians); in: Reproductive biology and phylogeny, Volume 5, Enfield, Jersey & Plymouth (Science Publisher): 39–78.

    Wilkinson, M., San Mauro, D., Sherratt, E. & Gower, D. J. (2011) A nine-family classification of caecilians (Amphibia: Gymnophiona). Zootaxa, 1874: 41–64. <https://doi.org/10.11646/zootaxa.2874.1.3>.

    Wilson, D. E. & Reeder, D. M. (ed.) (2005) Mammal species of the world. A taxonomic and geographic reference. Third edition. Baltimore (Johns Hopkins University Press), 1: i–xxxviii + 1–744; 2: i–xx + 745–2142.

    Zhang, P., Liang, D., Mao, R.-L., Hillis, D. M., Wake, D. B. & Cannatella, D. C. (2013) Efficient sequencing of anuran mtDNAs and a mitogenomic exploration of the phylogeny and evolution of frogs. Molecular Biology and Evolution, 30 (8): 1899–1915. <https://doi.org/10.1093/molbev/mst091>.