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Issue: Vol. 36 No. 1: 29 Dec. 2023
Type: Article
Published: 2023-12-29
DOI: 10.11646/bionomina.36.1.2
Page range: 36-63
Abstract views: 94
PDF downloaded: 0

The nomenclatural status of Agama cristata Merrem, 1819 and Corytophanes Schlegel, 1826 (Squamata, Corytophanidae) and of other tropical lizard nomina, with comments on some tricky issues of zoological nomenclature

Society for Southeast Asian Herpetology; Calle Rio Segura 26; 30600 Archena/Murcia; Spain
Institut de Systématique; Évolution; Biodiversité (ISYEB); Muséum national d’Histoire naturelle; CNRS; Sorbonne Université; EPHE; Université des Antilles; Paris; France
Association RACINE; 5 allée des cygnes; 35750 Saint Maugan; France
Institut de Systématique; Évolution; Biodiversité (ISYEB); Muséum national d’Histoire naturelle; CNRS; Sorbonne Université; EPHE; Université des Antilles; Paris; France
Agama cristata Calotes (Agama) tigrina Iguana superciliosa Lacerta superciliosa Corythophanes Corytophanes Lophyrus Ophryessa Uranoscodon Corythophanae Corytophanidae original description new replacement nomen nomen-bearing types authorship publication date

Abstract

The nomenclatural status of the nomina Agama cristata and Corytophanes (Squamata, Corytophanidae) are examined. The former should be credited to Merrem (1819), not Merrem (1820), and the latter to Schlegel (1826a), not H. Boie in Schlegel (1826b). The nomen Iguana superciliosa Latreille in Sonnini & Latreille, 1801 is shown to be a senior synonym of Agama cristata Merrem, 1819. Article 23.9 is used here to protect the latter through reversal of precedence. Several other nomenclatural problems related to these nomina and related ones are explored. Updated synonymies and aponymies are provided for all the nomina of the genus Corytophanes currently considered valid and for the family, the valid nomen of which is Corytophanidae Fitzinger, 1843 [Frost & Etheridge, 1989] based on Article 35.4.1 and through reversal of precedence. Attention is drawn to several nomina and spellings present in the literature but forgotten by all or most authors until now.

References

  1. Anonymous (1863) Lichtenstein’s Preis-Verzeichniss mexicanischer Vögel etc. vom Jahre 1830. Journal für Ornithologie, 11 (61): 54–60. <https://doi.org/10.1007/BF02005194>.
  2. Anonymous (1962) Species of animals bred in zoos and aquaria during 1961. International Zoo Yearbook, 3: 253–288. <https://doi.org/10.1111/j.1748-1090.1962.tb03446.x>.
  3. Anonymous [International Commission on Zoological Nomenclature] (1999) International code of zoological nomenclature. ‘Fourth edition’. London (International Trust for zoological Nomenclature): i–xxix + 1–306. [For the use of the authorship ‘Anonymous’ for this reference, see Dubois 2015b].
  4. Agassiz, L. (1848) Nomenclatoris zoologici. Index universalis, continens nomina systematica classium, ordinum, familiarum et generum animalium omnium, tam viventium quam fossilium, secundum ordinem alphabeticum unicum disposita, adjectis homonymiis plantarum., Soloduri (Jent & Gassmann): i–x + 1‒1135. <https://doi.org/10.5962/bhl.title.1819>.
  5. Alvarez del Toro, M. (1982) Los Reptiles de Chiapas. Tercera Edición, corregida y aumentada. Tuxtla Gutiérrez, Chiapas (Publicaciones del Instituto de Historia Natural): 1–248.
  6. Andersson, L. G. (1900) Catalogue of Linnean type-specimens of Linnaeus’ Reptilia in the royal Museum in Stockholm. Bihang till Kungliga Svenska Vetenskaps-Akademiens Handlingar, 26 (4) [1]: 1–29.
  7. Arnold, E. N. (1994) Investigating the evolutionary effects of one feature on another: does muscle spread suppress caudal autotomy in lizards? Journal of Zoology, 232: 505–523. <https://doi.org/10.1111/j.1469-7998.1994.tb01591.x>.
  8. Bartlett, R. D. (1983) Notes on the iguanid lizard, Corytophanes cristata. Notes from Noah, 10 (11): 2–4.
  9. Bauer, A. M. (2003) On the identity of Lacerta punctata Linnaeus, 1758, the type species of the genus Euprepis Wagler, 1830, and the generic assignment of Afro-Malagasy skinks. African Journal of Herpetology. 52: 1–7. <https://doi.org/10.1080/21564574.2003.9635472>.
  10. Bauer, A. M. (2009) Linnaean names and pre-Linnaean sources in herpetology. Bibliotheca herpetologica, 9: 53–79.
  11. Bauer, A. M. & Günther, R. (2014) Die Amphibien und Reptilien aus der von Borcke-Sammlung des Berliner Museums für Naturkunde: eine Fundgrube für verschollen geglaubtes Seba-Material. Sekretär, 14 (1): 3–33.
  12. Blömeke, E. & Müller, J. (1890) Katalog der Bibliothek des Reichstags. Berlin (Reichstag): 1‒744.
  13. Bocourt, M. F. (1874) 3. Reptiles. In: Milne-Edwards (1870‒1909): 113‒192, pl. 16‒17, 17bis, 18, 18bis. [Date and authorship according to Flores-Villela et al. (2016)].
  14. Boettger, O. & Pechuel-Loesche, [M. E.] (1892) Die Kriechtiere und Lurche. In: Dr. Alfred E. Brehm, Brehms Tierleben, Dritte, gänzlich neubearbeitete Auflage, Leipzig (Bibliographisches Institut), Vol. 7: i–xiv + 1–825, 1 pl.
  15. Boie, [H.] (1825) Zu Kaup’s Aufsatz über Lurche. Isis von Oken, 16 (10): 1089‒1090.
  16. Cantino, P. D. & Queiroz, K. de (2020) International code of phylogenetic nomenclature (PhyloCode). Boca Raton, London & New York (CRC Press): i‒xl + 1‒149. <https://doi.org/10.1201/9780429446320>.
  17. Caup [Kaup], [J.] (1825) Einige Bemerkungen zu Merrems Handbuch. Isis von Oken, 16 (4): 589‒593.
  18. Conrad, J. L. (2015) A new Eocene casquehead lizard (Reptilia, Corytophanidae) from North America. PLoS One, 10 (7) [0127900]: 1‒17. <https://doi.org/10.1371/journal.pone.0127900>.
  19. Cope, E. D. (1865) Third contribution to the herpetology of tropical America. Proceedings of the Academy of natural Sciences of Philadelphia, 17: 185–198.
  20. Cope, E. D. (1900) The crocodilians, lizards and snakes of North America. Annual Report of the United States national Museum for 1898: 153–1270.
  21. Cuvier, [G.] (1816) Le règne animal distribué d’après son organisation, pour servir de base à l’histoire naturelle des animaux et d’introduction à l’anatomie comparée. Tome 2, contenant les reptiles, les poissons, les mollusques et les annélides. Paris (Deterville), ‘1817’: Paris (Deterville): i–xviii + 1–532.
  22. Daudin, F. M. (1802) [An. X] Histoire naturelle, générale et particulière, des Reptiles. Tome 3. Paris (Dufart): 1–452, pl. 29–45.
  23. Daudin, F. M. (1803) [An. XI] Histoire naturelle des rainettes, des grenouilles et des crapauds. Quarto version. Paris (Levrault): 1–108, pl. 1–38.
  24. Denzer, W. & Dondorp, E. (2023) Lyriocephalus scutatus (Linnaeus, 1758). Anmerkungen zur Nomenklatur und zu möglichem Typusmaterial. Sauria, 45 (1): 35–42.
  25. Deppe, W. (1830) Preis-Verzeichniss der Säugethiere, Vögel, Amphibien, Fische und Krebse, welche von den Herren Deppe und Schiede in Mexico gesammelt worden, und bei dem unterzeichneten Bevollmächtigten in Berlin gegen baare Zahlung in Preuss. Courant zu erhalten sind. Privately printed by the author, Berlin. [Not seen, reprinted in Anonymous (1863)].
  26. Dubois, A. (1982) Le statut nomenclatural des noms génériques d’Amphibiens créés par Kuhl & Van Hasselt (1822): Megophrys, Occidozyga et Rhacophorus. Bulletin du Muséum national d’Histoire naturelle, (4), 4 (A): 261–280. <https://doi.org/10.5962/p.286038>.
  27. Dubois, A. (1987) Again on the nomenclature of frogs. Alytes, 6 (1–2): 27–55.
  28. Dubois, A. (2000) Synonymies and related lists in zoology: general proposals, with examples in herpetology. Dumerilia, 4 (2): 33–98.
  29. Dubois, A. (2005) Proposed Rules for the incorporation of nomina of higher-ranked zoological taxa in the International Code of Zoological Nomenclature. 1. Some general questions, concepts and terms of biological nomenclature. Zoosystema, 27 (2): 365–426. <https://doi.org/10.11646/zootaxa.1337.1.1>.
  30. Dubois, A. (2010) Retroactive changes should be introduced in the Code only with great care: problems related to the spellings of nomina. Zootaxa, 2426: 1–42. <https://doi.org/10.11646/zootaxa.2426.1.1>.
  31. Dubois, A. (2011) The International Code of Zoological Nomenclature must be drastically improved before it is too late. Bionomina, 2: 1–104. <https://doi.org/10.11646/bionomina.2.1.1>.
  32. Dubois, A. (2013) Zygoidy, a new nomenclatural concept. Bionomina, 6: 1–25. <https://doi.org/10.11646/bionomina.6.1.1>.
  33. Dubois, A. (2015a) Zoological nomina in the century of extinctions: new proposals. Bionomina, 8: 11–53. <https://doi.org/10.11646/bionomina.8.1.2>.
  34. Dubois, A. (2015b) What is an anonymous publication? Is the International Code of Zoological Nomenclature anonymous? Bionomina, 9: 27–34. <https://doi.org/10.11646/bionomina.9.1.2>.
  35. Dubois, A. (2017a) A few problems in the generic nomenclature of insects and amphibians, with recommendations for the publication of new generic nomina in zootaxonomy and comments on taxonomic and nomenclatural databases and websites. Zootaxa, 4237 (1): 1–16. <https://doi.org/10.11646/zootaxa.4237.1.1>.
  36. Dubois, A. (2017b) The nomenclatural status of Hysaplesia, Hylaplesia, Dendrobates and related nomina (Amphibia, Anura), with general comments on zoological nomenclature and its governance, as well as on taxonomic databases and websites. Bionomina, 11: 1–48. <https://doi.org/10.11646/bionomina.11.1.1>.
  37. Dubois, A. (2021) Recommendations concerning the presentation of synonymic and related lists in zootaxonomy: the example of the family Cryptobranchidae Fitzinger, 1826 (Amphibia, Urodela). Bionomina, 25: 52–69. <https://doi.org/10.11646/bionomina.25.1.4>.
  38. Dubois, A. (2022) Type species of genera in zoological nomenclature. Bionomina, 26: 1‒33. <https://doi.org/10.11646/bionomina.31.1.1>.
  39. Dubois, A. & Aescht, E. (ed.) (2019) LZC Session 20. The Principles of the Zoocode. 6. The Principle of Neonymy. Dumerilia, 8: 48–52.
  40. Dubois, A., Bauer, A. M., Ceríaco, L. M. P., Dusoulier, F., Frétey, T., Löbl, I., Lorvelec, O., Ohler, A., Stopiglia, R. & Aescht, E. (2019) The Linz Zoocode project: a set of new proposals regarding the terminology, the Principles and Rules of zoological nomenclature. First report of activities (2014‒2019). Bionomina, 17: 1‒111. <https://doi.org/10.11646/bionomina.17.1.1>.
  41. Dubois, A., Frétey, T., Lorvelec, O. & Ohler, A. (2021b) The nomenclatural status of the amphibian and reptile nomina introduced by La Cepède in his Histoire Naturelle des Quadrupèdes Ovipares et des Serpens, with comments on various questions of zoological nomenclature. Bionomina, 23: 1‒180. <https://doi.org/10.11646/bionomina.23.1.1>.
  42. Dubois, A., Frétey, T. & Ohler, A. (2022) The nomenclature of Eurasian giant salamanders (Amphibia, Urodela, Andrias). Bionomina, 31: 70‒87. <https://doi.org/10.11646/bionomina.31.1.4>.
  43. Dubois, A., Nemésio, A. & Bour, R. (2014) Primary, secondary and tertiary syntypes and virtual lectotype designation in zoological nomenclature, with comments on the recent designation of a lectotype for Elephas maximus Linnaeus, 1758. Bionomina, 7: 45–64. <https://doi.org/10.11646/bionomina.7.1.2>.
  44. Dubois, A. & Ohler, A. (1997a) Early scientific names of Amphibia Anura. I. Introduction. Bulletin du Muséum national d’Histoire naturelle, (4), 18 (3–4): 297–320. <https://doi.org/10.5962/p.290335>.
  45. Dubois, A. & Ohler, A. (1997b) Early scientific names of Amphibia Anura. II. An exemplary case: Rana arborea Linnaeus, 1758. Bulletin du Muséum national d’Histoire naturelle, (4), 18 (3–4): 321–340. <https://doi.org/10.5962/p.290335>.
  46. Dubois, A., Ohler, A. & Pyron, R. A. (2021a) New concepts and methods for phylogenetic taxonomy and nomenclature in zoology, exemplified by a new ranked cladonomy of recent amphibians (Lissamphibia). Megataxa, 5 (1): 1‒738. <https://doi.org/10.11 646/megataxa.5.1.1>.
  47. Dugès, A. (1896). Reptiles y batracios de los E.U. mexicanos. La Naturaleza, (2), 2 (11): 479–485.
  48. Duméril, A. H. A. (1856) Description des reptiles nouveaux ou imparfaitement connus de la collection du Muséum d’Histoire Naturelle et remarques sur la classification et les caractères des reptiles. Deuxième mémoire. Troisième, quatrième et cinquième famille de l’ordre des Sauriens. Archives du Muséum d’Histoire naturelle, Paris, 8: 437‒588, pl. 17‒24.
  49. Duméril, A. M. C. (1805) Zoologie analytique, ou méthode naturelle de classification des animaux, rendue plus facile à l’aide de tableaux synoptiques. Paris (Allais) ‘1806’: i–xxxiii + 1–544. <https://doi.org/10.5962/bhl.title.44835>.
  50. Duméril, A.-M.-C. & Bibron, G. (1837) Erpétologie générale ou Histoire naturelle complète des Reptiles. Tome 4. Paris (Roret): [i–iii] + i–ii + 1–572.
  51. Duméril, A.-M.-C. & Bibron, G. (1841) Erpétologie générale ou Histoire naturelle complète des Reptiles. Tome 8. Paris (Roret): i–vii + 1–792.
  52. Elter, O. (1982) La collezione erpetologica del Museo di Zoologia dell’Università di Torino. Cataloghi, Museo regionale de Scienze naturale, Torino, 5: 1–116.
  53. Etheridge, R. (1970) A review of the South American iguanid genus Plica. Bulletin of the British Museum of natural History (Zoology), 19: 237–256. <https://doi.org/10.5962/bhl.part.24087>.
  54. Etheridge, R. & Queiroz, K. de (1988) A phylogeny of Iguanidae. In: R. Estes & G. Pregill (ed.), Phylogenetic relationships of the lizard families, Essays commemorating Charles L. Camp, Stanford, California (Stanford University Press): 283–368.
  55. Fitzinger, L. I. (1843) Systema Reptilium. Fasc. 1. Amblyglossae. Vindobonae (Braumüller & Seidel): 1–106 + i–ix. <https://doi.org/10.5962/bhl.title.4694>.
  56. Fitzinger, L. J. (1861) Die Ausbeute der österreichischen Naturforscher an Säugethieren und Reptilien während der Weltumsegelung Sr. Majestät Fregatte Novara. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe, ‘1860’, 42: 383–416.
  57. Flores-Villela, O., Bour, R. & Adler, K. (2016) Publication history of the Mission scientifique au Mexique et dans l’Amérique Centrale, reptiles and amphibians. Historia de la publicación de la Mission scientifique au Mexique et dans l’Amérique Centrale, reptiles y anfibios. Revista mexicana de Biodiversidad, 87: 1162‒1167. <https://doi.org/10.1016/j.rmb.2016.07.013>.
  58. Frétey, T. & Dubois, A. (2019) The authorship and date of five generic nomina of Squamata and Amphibia published by Cuvier (1816, 1829), with recourse to and comments on twenty Articles of the Code. Bionomina, 15: 1‒36. <https://doi.org/10.11646/bionomina.15.1.1>.
  59. Frétey, T. & Raffaëlli, J. (2021) Barton and the forgotten description of Salamandra horrida (Amphibia, Urodela): story of a rediscovery. Bionomina, 25: 35‒51. <https://doi.org/10.11646/bionomina.25.1.3>.
  60. Froriep, L F; (1806) C. Duméril’s Analytische Zoologie. Aus dem Französchen, mit Zusätzen. Weimar (Landes-Industrie Comptoirs): [i] + i‒vi + 1‒344.
  61. Frost, D. R. (1992) Phylogenetic analysis and taxonomy of the Tropidurus group of lizards (Iguania: Tropiduridae). American Museum Novitates, 3033: 1‒68.
  62. Frost, D. R. & Etheridge, R. (1989) A phylogenetic analysis and taxonomy of iguanian lizards (Reptilia: Squamata). Miscellaneous Publications of the Museum of natural History, University of Kansas, 81: 1–65. <https://doi.org/10.5962/bhl.title.16288>.
  63. Frost, D. R., Etheridge, R., Janies, D. & Titus, T. A. (2001). Total evidence, sequence alignment, evolution of polychrotid lizards and a reclassification of the Iguania (Squamata: Iguania). American Museum Novitates, 3343: 1–38. <https://doi.org/10.1206/0003-0082(2001)343<0001:TESAEO>2.0.CO;2>.
  64. Gravenhorst, J. L. C. (1833) Über Phrynosoma orbicularis, Trapelus hispidus, Phrynocephalus helioscopus, Corythophanes cristatus, und Chamaeleopsis hernandesii. Nova acta physico-medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosum, 16: 909–958.
  65. Gray, J. E. (1825) A synopsis of the genera of Reptiles and Amphibia, with a description of some new species. Annals of Philosophy, (2), 10: 193–217.
  66. Gray, J. E. (1827) A description of new species of saurian reptiles; with a revision of the species of chamaeleons. The Philosophical Magazine and Annals of Philosophy, (2), 2 (9): 207–214. <https://doi.org/10.1080/14786442708675648>.
  67. Gray, J. E. (1831) A synopsis of the species of the class Reptilia. In: E. Griffith & E. Pidgeon (ed.), The class Reptilia arranged by the Baron Cuvier, with specific descriptions; in: E. Griffith, & E. Pidgeon (ed.), The animal kingdom arranged in conformity with its organization, by the Baron Cuvier, member of the Institute of France, &c. &c. &c., with additional descriptions of all the species hitherto named, and of many not before noticed, vol. 9, London (Whittaker, Treacher & Co.): 1–110, pl. 1–55.
  68. Günther, A. (1864) The Reptiles of British India. London (Ray Society): i–xxvii + 1–452, pl. 1–26. <https://doi.org/10.5962/bhl.title.5012>.
  69. Günther, A. C. L. G. (1885) Reptilia and Batrachia. In: O. Salvin & F. D. Godman (ed.), Biologia centrali-americana; or, Contributions to the knowledge of the fauna and flora of Mexico and Central America, London (R. H. Porter & Dulau & Co.): 33–56.
  70. Herrera, A. L. (1899) Sinonimia vulgar y cientifíca de los principales vertebrados mexicanos. México, D.F. (Oficio Tipográfico de la Secretaría de Fomento): 1‒31. • Work rejected by the International Commission on Zoological Nomenclature (Melville 1984). See also Smith & Smith (1975).
  71. Johnson, J. D., Ely, C. A. & Webb, R. G. (1977) Biogeographical and taxonomic notes on some herpetozoa from the northern highlands of Chiapas, Mexico. Transactions of the Kansas Academy of Science, ‘1976’, 79: 131–139. <https://doi.org/10.2307/3627292>.
  72. Kästle, W. (1975) The iguanids. In: B. Grzimek, H. Hediger, K. Klemmer, O. Kuhn & H. Wermuth (ed.), Grzimek’s animal life encyclopedia, Vol. 6, Reptiles, New York (Van Nostrand Reinhold Co.): 177–203.
  73. Kaup, J. J. (1826) Monographie der Gattung Uraniscodon. Isis von Oken, 18 (1): 89–91.
  74. Kaup, J. (1827) Zoologische Monographien. Isis von Oken, 20 (6): 610‒625.
  75. Kaup, [J.] (1828) Ueber Hyaena, Uromastyx, Basiliscus, Corytheolus, Acontias. Isis von Oken, 21 (9): 1144‒1150.
  76. Kraus, F., O’Shea, M. & Kaiser, H. (2023) Why Toxicocalamus longhagen Roberts, Iova & Austin, 2022 (Serpentes, Elapidae) is a taxonomic nomen dubium. Bionomina, 32: 41‒51. <https://doi.org/10.11646/bionomina.32.1.4>.
  77. Kuhn, O. (1944) Weitere Lacertilier, insbesondere Iguanidae aus dem Eozän des Geiseltales. Palaeontologische Zeitschrift, 23: 360‒366. <https://doi.org/10.1007/BF03160444>.
  78. La Cepède, [B.-G.-É. de la Ville], de (1788) Histoire naturelle des quadrupèdes ovipares et des serpens. [In-quarto]. Tome 1. Paris (Hôtel de Thou): [i–v]+ 6–17 + 1–651, pl. 1–41, 2 tab.
  79. La Cépède, [B.-G.-É. de la Ville], de (1789) Histoire naturelle des serpens. [In-quarto]. Tome 2. Paris (Hôtel de Thou): [i] + 1–8 + 1–19 + [i] + 1–144 + 1–527, pl. 1–22.
  80. Lang, M. A. (1989) Phylogenetic and biogeographical patterns of basiliscine iguanians (Reptilia: Squamata: ‘Iguanidae’). Bonner Zoologische Monographien, 28: 1–172.
  81. Laurenti, J. N. (1768) Specimen medicum, exhibens synopsin Reptilium emendatam cum experimentis circa venena et antidota Reptilium austriacorum. Viennae (Joan. Thom. Nob. de Trattnern): i–ii + 1–215, pl. 1–5. <https://doi.org/10.5962/bhl.title.5108>.
  82. Lichtenstein, H., Weinland, D. & Martens, [E.] von (1856) Nomenclator Reptilium et Amphibiorum Musei Zoologici Berolinensis. Namenverzeichniss der in der zoologischen Sammlung der Königlichen Universität zu Berlin aufgestellten Arten von Reptilien und Amphibien nach ihren Ordnungen, Familien und Gattungen. Berlin (Königliche Academie der Wissenschaften): i–iv + 1‒48. <https://doi.org/10.5962/bhl.title.45245>.
  83. Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Tomus 1. Holmiae (Laurentii Salvii): [i–iv] + 1–824. <https://doi.org/10.5962/bhl.title.542>.
  84. Linné, C. a (1766) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus 1. Editio duodecima, Reformata. Holmiae (Laurentii Salvii): 1–532. <https://doi.org/10.5962/bhl.title.68927>.
  85. Macey, J. R., Larson, A., Ananjeva, N. B. & Papenfuss, T. J. (1997) Evolutionary shifts in three major structural features of the mitochondrial genome among iguanian lizards. Journal of molecular Evolution, 44: 660–674. <https://doi.org/10.1007/PL00006190>.
  86. Manthey, U. & Denzer, W. (1993) Die Echten Winkelkopfagamen der Gattung Gonocephalus Kaup (Sauria: Agamidae). VI. Die chamaeleontinus-Gruppe. Sauria, 15 (2): 23‒38.
  87. McCranie, J. R. (2018) The lizards, crocodiles, and turtles of Honduras. Systematics, distribution, and conservation. Bulletin of the Museum of comparative Zoology, Special Publications Series, 2: [i‒viii] + 1–666. <https://doi.org/10.3099/0027-4100-15.1.1>.
  88. Melville, R. V. (ed.) (1984) Direction 114. Herrera, 1899, ‘Sinonima vulgar y cientifica de los principales vertebrados mexicanos’ placed on the Official Index of Rejected and Invalid Works in Zoology (Direction supplementary to Direction 32). Bulletin of zoological Nomenclature, 41 (1): 39‒40.
  89. Mendelson, J. R., III (1990) Notas sobre una colleción [sic] de anfibios y reptiles de Pueblo Viejo, Alta Verapaz, Guatemala. Notes on a collection of amphibians and reptiles from Pueblo Viejo, Alta Verapaz. Publicaciόn ocasional, Fundaciόn interamericana de Investigaciόn tropical, Guatemala (Guatelmala City), 3: 1–18.
  90. Merrem, [B.] (1819) Agama. In: J. S. Ersch & J. G. Gruber, Allgemeine Encyclopädie der Wissenschaften und Künste in alphabetischer Folge von genannten Schriftstellern bearbeitet, Theil 2, Äga–Aldus: 162–168. [Publication date according to Blömeke & Müller (1890)].
  91. Merrem, B. (1820) Versuch eines Systems der Amphibien. Tentamen systematis Amphibiorum. Marburg (Iohann Christian Krieger): [i–xv] × 2 + 1 + [2–188] × 2 + 189–191, 1 pl.
  92. Milne-Edwards, H. (ed.) (1870–1909) Mission scientifique au Mexique et dans l’Amérique Centrale. Recherches zoologiques pour servir à l’histoire de la faune de l’Amérique Centrale et du Mexique. Paris (Imprimerie Impériale): i–xiv +[1‒6] + 1‒1012 + 1 map (text volume), 99 pl. (atlas).
  93. Myers, C. W. & Stothers, R. B. (2006) The myth of Hylas revisited: the frog name Hyla and other commentary on Specimen medicum (1768) of J. N. Laurenti, the ‘father of herpetology’. Archives of natural history, 33 (2): 241–266. <https://doi.org/10.3366/anh.2006.33.2.241>.
  94. Neave, S. A (ed.) (1939) Nomenclator zoologicus. Vol. 1. A-C. London (Zoological Society of London): i–xiv + 1–957.
  95. Oppel, M. (1811) Ordnungen, Familien und Gattungen der Reptilien als Prodrom einer Naturgeschichte derselben. München (Joseph Lindauer): i–xii + 1–86. <https://doi.org/10.5962/bhl.title.4911>.
  96. Peters, J. A. & Donoso-Barros, R. (1970) Catalogue of the Neotropical Squamata. Part II. Lizards and Amphisbaenians. Bulletin of the United States national Museum, 297: i–vii + 1–293. <https://doi.org/10.5479/si.03629236.297.1>.
  97. Poli, G. S. (1791) Testacea utruisque Siciliae corumque historia et anatome. Vol. 1. Parmae (ex Regio Typpographeio): [i‒vii] + i‒x + 1‒90 + 1‒51 + 1‒lxxiii. <https://doi.org/10.5962/bhl.title.79042>.
  98. Renous-Lécuru, S. & Jullien, R. (1972) Les grands troncs nerveux du zeugopode des lacertiliens (Reptiles, Squamates). Bulletin du Muséum d’Histoire naturelle, (3), 29, (Zoologie 23): 165–206.
  99. Russell, A. P. & Bauer, A. M. (1988) An early description of a member of the genus Phelsuma (Reptilia: Gekkonidae), with comments on names erroneously applied to Uroplatus fimbriatus. Amphibia-Reptilia, 9: 107‒116. <https://doi.org/10.1163/156853888X00521>.
  100. Savage, J. M. (1986) Book review. La nomenclature supragénérique des Amphibiens Anoures. Alain Dubois. 1984. Copeia, 1986: 259–262. <https://doi.org/10.2307/1444926>.
  101. Savage, J. M., Myers, C. W., Frost, D. R. & Grant, T. (2007) Case 3345. Dendrobatidae Cope, 1865 (1850) (Amphibia, Anura): proposed conservation. Bulletin of zoological Nomenclature, 64 (4): 255‒260.
  102. Schinz, H. R. (1822) Das Thierreich eingetheilt nach dem Bau der Thiere als Grundlage ihrer Naturgeschichte und der vergleichenden Anatomie von dem Herrn Ritter von Cuvier. Vol. 2. Stuttgart & Tübingen (J. G. Cotta): i‒xvi + 835.
  103. Schlegel, [H.] (1826a) [31 October] Notice sur l’erpétologie de l’île de Java; par M. Boie. (Ouvrage manuscrit.). Bulletin des Sciences naturelles et de Géologie, 9 (2): 233‒240.
  104. Schlegel, [H.] (1826b) [31 December] Erpetologische Nachrichten. Isis von Oken, 20 (3): 281‒294.
  105. Schreber, J. C. D. von (1782) Beytrag zur Naturgeschichte der Frösche. Der Naturforscher, Halle, 18: 182–193.
  106. Schulte, J. A., II., Valladares, J. P. & Larson, A. (2003) Phylogenetic relationships within Iguanidae inferred using molecular and morphological data and a phylogenetic taxonomy of iguanian lizards. Herpetologica, 59: 399–419. <https://doi.org/10.1655/02-48>.
  107. Seba, A. (1734) Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam. Opus, cui, in hoc rerum genere, nullum par exstitit. Ex toto terrarum orbe collegit, digessit, descripsit, et depingendum curavit. Tomus 1. Amstelaedami (J. Wetstenium, Gul. Smith & Janssonio-Waesbergios): [i–xxxii] + 1–178, 1 pl., pl. 1–111. <https://doi.org/10.5962/bhl.title.14110>.
  108. Sehnal, P. (1996) Esquinas-Nationalpark, “Der Regenwald der Österreicher”. Gebietsbeschreibung ‒ Regenwaldbiologie ‒ Naturbeobachtungen. In: P. Sehnal and H. Zettel (ed.), Esquinas-Nationalpark, Der Regenwald der Österreicher in Costa Rica, Wien (Naturhistorisches Museum, Wien): 80–111.
  109. Sherborn, C. D. (1932) Index animalium sive index nominum quae ab a.d. MDCCLVIII generibus et speciebus animalium imposita sunt. Sectio secunda. A kalendis ianuariis, MDCCCL usque ad finem decembris, MDCCCL. Part XXIX. Epilogue, additions to bibliography, additions and corrections, index to trivialia under genera, pp. 1‒208. 1801‒1850. London (Printed by order of the Trustees of the British Museum): [i] + i–vii + cxxxiii‒cxlviii + 1–208.
  110. Smith, H. M. (1947) Notas sobre una colección de reptiles y anfibios de Chiapas, Mex. Revista de la Sociedad méxicana de Historia natural, ‘1946’, 7 (1–4): 63–74.
  111. Smith, H. M. & Smith, R. B. (1973) Chresonymy ex synonymy. Systematic Zoology, ‘1972’, 21: 445. <https://doi.org/10.2307/2412440>.
  112. Smith, H. M. & Smith, R. B. (1975) The herpetological names of Herrera, 1899, and their status. Transactions of the Kansas Academy of Science, 78: 85–87. <https://doi.org/10.2307/3627227>.
  113. Smith, H. M. & Taylor, E. H. (1950) An annotated checklist and key to the reptiles of Mexico exclusive of the snakes. Bulletin of the United States National Museum, 199: i‒v + 1‒253. <https://doi.org/10.5479/si.03629236.199>.
  114. Smith, P. W. & Burger, W. L. (1950) Herpetological results of the University of Illinois Field Expedition, Spring 1949. III. Sauria. Transactions of the Kansas Academy of Science, 53: 165–175. <https://doi.org/10.2307/3625970>.
  115. Sonnini, C. S. & Latreille, P. A. (1801) Histoire naturelle des reptiles. Tome 1. Première partie. Quadrupèdes et Bipèdes Ovipares. Paris, Deterville: [i–iii] + i–xxii + 1–332, 14 pl.
  116. Spix, J. B. de (1825) Animalia nova sive species nova lacertarum, quas in itinere per Brasiliam annis MDCCCXVII-MDCCCXX jussu et auspicius Maximiliani Josephi I Bavariae Regis suscepto collegit et descripsit. Monachii (Typis Franc. Seraph. Hübschmann): [i‒vi] + 1‒26, pl. 1‒28. <https://doi.org/10.5962/bhl.title.5117>.
  117. Strickland, H. E., Henslow, J. S., Phillips, J., Shuckard, W. E., Richardson, J., Waterhouse, G. R., Owen, R., Yarrell, W., Jenyns, L., Darwin, C., Broderip, W. J. & Westwood, J. O. (1843) Series of propositions for rendering the nomenclature of zoology uniform and permanent, being the Report of a Committee for the consideration of the subject appointed by the British Association for the Advancement of Science. Annals & Magazine of natural History, (1), 11: 259–275. <https://doi.org/10.1080/03745484309445300>.
  118. Stuart, L. C. (1948) The amphibians and reptiles of Alta Verapaz Guatamala. Miscellaneous Publications, Museum of Zoology, University of Michigan, 69: [i‒iv] + 1‒109.
  119. Sumichrast, F. (1864) Note on the habits of some Mexican Reptiles. Annals & Magazine of natural History, (3), 13: 497‒507. <https://doi.org/10.1080/00222936408681641>.
  120. Sumichrast, F. (1870) Notas sobre las costumbres de algunos reptiles de México. Familia de los iguanidéos. La Naturaleza, 1: 176–180 + 203–206.
  121. Taylor, E. H. (1956) A review of the lizards of Costa Rica. The University of Kansas Sience Bulletin, 38 (1): 3‒322.
  122. Townsend, J. H., McCranie, J. R. & Wilson, L. D. (2004a) Corytophanes Boie. Catalogue of American Amphibians and Reptiles, 788: 788.1‒788.4.
  123. Townsend, J. H., McCranie, J. R. & Wilson, L. D. (2004b) Corytophanes cristatus (Merrem). Catalogue of American Amphibians and Reptiles, 789: 789.1‒789.6.
  124. Troschel, F. H. (1857) Bericht über die Leistungen in der Herpetologie während des Jahres 1856. Archiv für Naturgeschichte, 23 (2): 62–76.
  125. Troschel, F. H. (1865) Amphibia. In: J. W. von Müller, Reisen in den Vereinigten Staaten, Canada und Mexico, Beiträge zur Geschichte, Statistik und Zoologie von Mexico, Leipzig (Brodhaus): 595–619.
  126. Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. & Smith, G. F. (ed.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum vegetabile, 159. Glashütten (Koeltz Botanical Books). <https://doi.org/10.12705/Code.2018>.
  127. Uetz, P., Freed, P.,Aguilar, R., Reyes, F. & Hošek, J. (ed.) (2023) The Reptile Database. <http://www.reptile-database.org>. [Last accessed 20 October 2023].
  128. Velasco, A. L. (1895) Geografía y estadística del Estado de Campeche. Geografía y estadística de la República Mexicana, 16, México, D.F. (Tipográfica de la Secretaría de Fomento): 1‒140.
  129. Velasco, A. L. (1896) Geografía y estadística del Estado de Coliima. Geografía y estadística de la República Mexicana, 18, México, D.F. (Tipográfica de la Secretaría de Fomento): 1‒142.
  130. Vieira, G. H. C., Colli, G. R. & Báo, S. N. (2005) Phylogenetic relationships of corytophanid lizards (Iguania, Squamata, Reptilia) based on partitioned and total evidence analyses of sperm morphology, gross morphology, and DNA data. Zoologica scripta, 34: 605–625. <https://doi.org/10.1111/j.1463-6409.2005.00208.x>.
  131. Wagler, J. (1830) Natürliches System der Amphibien, mit vorangehender Classification der Säugethiere und Vögel. München, Stuttgart & Tübingen (J. G. Cotta): i–vi + 1–354. <https://doi.org/10.5962/bhl.title.108661>.
  132. Wagler, J. (1833) Deutung der in Seba’s Thesaura reum naturalium T.1 et 2. enthaltenen Abbildungen von Lurchen, mit critischen Bermerkungen. Isis von Oken, 26 (9): 885–905.
  133. Wied-Neuwied, M., Prinz zu (1821) Reise nach Brasilien in den Jahren 1815 bis 1817. Vol. 2. Frankfurt a. M. (Heinrich Ludwig Bronner): i‒xviii + 1‒346, 1 pl. <https://doi.org/10.5962/bhl.title.85967>.
  134. Wiegmann, A. F. (1828) Beyträge zur Amphibienkunde. Isis von Oken, 21 (3): 364‒381.
  135. Wiegmann, A.F.A. (1831) Zoologische Notizen. Isis von Oken, 24 (3): 282‒301.
  136. Wiegmann, A. F. A. (1834) Herpetologia mexicana, seu descriptio Amphibiorum Novae Hispaniae, quae itineribus Comitis de Sack, Ferdinandi Deppe et Chr. Guil. Schiede in Museum Berolinense pervenerunt. Pars prima, Saurorum species amplectens. Adiecto systematis Saurorum prodromo, additisque multis in hunc Amphibiorum ordinem observationibus. Berolini (C. G. Lüderitz): i‒vi + 1‒54, pl. 1‒10. <https://doi.org/10.5962/bhl.title.119131>.
  137. Zug, G. R. (1971) The distribution and patterns of the major arteries of the iguanids and comments on the intergeneric relationships of iguanids (Reptilia: Lacertilia). Smithsonian Contributions in Zoology, 83: [i‒iv] + 1–23. <https://doi.org/10.5479/si.00810282.83>.