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Type: Correspondence
Published: 2020-02-28
Page range: 22-25
Abstract views: 239
PDF downloaded: 2

First country record of Pseudoathyreus orientalis (Coleoptera: Bolboceratidae: Athyreinae) from Pakistan

Department of zoology, Faculty of Science, Charles University, Vinicna 7, Praha 2, 128 44 CZ
Department of zoology, Faculty of Science, Charles University, Vinicna 7, Praha 2, 128 44 CZ
Department of zoology, Faculty of Science, Charles University, Vinicna 7, Praha 2, 128 44 CZ


Bolboceratidae is a large family of about 600 species (e.g., Schoolmeesters 2019), in the superfamily Scarabaeoidea. Currently, it includes two subfamilies, Athyreinae and Bolboceratinae (e.g., Scholtz & Grebennikov 2016; Nikolajev et al. 2016). The Athyreinae includes five genera and about 130 species from the Afrotropical, Neotropical, Oriental and Palearctic regions (e.g., Schoolmeesters 2019). The genus Pseudoathyreus Howden & Martínez, 1963 contains nine species occurring in Africa and Asia (e.g., Carpaneto et al. 2000; Schoolmeesters 2019). Little is known about the biology of Athyreinae (Scholtz & Grebennikov 2016). Immature stages of this group remain unknown (Grebennikov & Scholtz 2004).


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