Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2023-11-29
Page range: 57-67
Abstract views: 134
PDF downloaded: 14

Taxonomic notes on the leafhopper genus Balclutha Kirkaldy (Hemiptera: Cicadellidae) from Pakistan with a new country record

Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A&F University, 666 Wusu street, Linan, Hangzhou, Zhejiang 311300, P.R. China
Institute of Entomology, The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University, Guiyang 550025, P.R. China
School of Life Sciences, Jiangsu University, Zhenjiang 212013, P.R. China
Provincial Key Laboratory for Agricultural Pest Management of Mountainous Regions, Institute of Entomology, Guizhou University, Guiyang 550025, P.R. China
Department of Entomology, Faculty of Crop Protection Sciences, The University of Agriculture, Peshawar-Pakistan
Anhui Province Key Laboratory of Integrated Pest Management on Crops, Key Laboratory of Biology and Sustainable Management of Plant Diseases and Pests of Anhui Higher Education Institutes, School of Plant Protection, Anhui Agricultural University, Hefei, Anhui 230036, P.R. China
Auchenorrhyncha taxonomy new record Pakistan


The leafhopper genus Balclutha Kirkaldy (Deltocephalinae: Macrostelini) is reviewed from Pakistan. Balclutha sinuata Webb & Vilbaste is recorded as a new faunal record for Pakistan. Herein we provide an updated checklist, distribution map, an identification key to the males of known Pakistani species, along with the digital illustrations of the species.


  1. Abdollahi T., Jalalizand A. R., Mozaffarian F. & Wilson M. 2015. A faunistic study on the leafhoppers of northwestern Iran (Hemiptera, Cicadellidae). ZooKeys 496: 27–51.
  2. Ahmed M. 1986. Some investigations of leafhoppers of grasslands and allied crops in Pakistan. Proceedings of the Sixth Pakistan Congress of Zoology, pp. 51–62.
  3. Ahmed M., Murtaza B. & Malik K. F. 1988. Some new Aphrodine leafhoppers from grasslands of Karachi, Pakistan. Pakistan Journal of Zoology 20: 409–421.
  4. Baloch R. A., Khatri I., Baloch A., Nizam-u-Din Menghwar S. & Dad S. 2018. Records of leafhoppers (Cicadellidae) of Mekran division, Balochistan, Pakistan. Journal of Entomology and Zoology Studies 6: 108–110.
  5. Blocker H. D. 1967. Classification of the Western Hemisphere Balclutha (Homoptera: Cicadellidae). Proceedings of the United States National Museum 122–155.
  6. Caldwell J. S. & Martorell L. F. 1952. New leafhoppers from Puerto Rico (Cicadellidae: Homoptera). The Journal of Agriculture of the University of Puerto Rico. Rio Piedras 35: 88–89.
  7. Chiang C. C. 1996. Studies on the Genus Balclutha (Homoptera: Cicadellidae) of Taiwan. Journal of Taiwan Museum 49 (1): 61–71.
  8. Dai R. H., Li Z. Z. & Chen X. X. 2004. Notes on Chinese species of Balclutha with descriptions of three new species (Homoptera, Cicadellidae, Euscelinae). Acta Zootaxonomica Sinica 29 (4): 749–755.
  9. Dietrich C. H. 2005. Keys to the families of Cicadomorpha and subfamilies and tribes of Cicadellidae (Hemiptera: Auchenorrhyncha). Florida Entomologist 88(4): 502–517.[502:KTTFOC]2.0.CO;2
  10. Distant W. L. 1908. Rhynchota-Homoptera. In: Bingham CT (Eds.), London. The Fauna of British India, including Ceylon and Burma 4: 1–501.
  11. Distant W. L. 1918. Rhynchota-Homoptera. In: Shipley AE, Marshall AK (Eds.), London. The Fauna of British India, including Ceylon and Burma 7: 1–210.
  12. Evans J. W. 1977. The leafhoppers and froghoppers of Australia and New Zealand. (Homoptera: Cicadelloidea and Cercopoidea). Part 2. Records of the Australian Museum 31: 83–129.
  13. Fabricius J. C. 1775. Ryngota. In: Systema entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. In Officina Libraria Kortii, Flensbvrgi et Lipsiae 1–816.
  14. Fieber F. X. 1866. Neue Gattungen und Arten in Homopteren (Cicadina Bur.). Verhandlungen Zoologisch-Botanischen Gesellschaft in Wien 16: 497–516.
  15. Khatri I. & Webb M. D. 2010. The Deltocephalinae leafhoppers of Pakistan (Hemiptera: Cicadellidae). Zootaxa 2365: 1–47.
  16. Kirkaldy G. W. 1900. Bibliographical and nomenclatorial notes on the Rhynchota. No. 1. Entomologist 33: 238–243.
  17. Kirkaldy G. W. 1906. Leaf-hoppers and their natural enemies, (pt. IX Leaf-hoppers, Hemiptera). Report of work of the experiment station of the Hawaiian Sugar Planters’ Association. Division of Entomology Bulletin 1(9): 271–479.
  18. Knight W. J. 1965. Techniques for use in the identification of leafhoppers (Homoptera: Cicadellidae). Entomologist’s Gazette 16: 129–36.
  19. Knight W. J. 1987. Leafhoppers of the grass-feeding genus Balclutha (Homoptera, Cicadellidae) in the Pacific region. Journal of Natural History 21: 1173–1224.
  20. Linnavuori R. E. 1959. Revision of the Neotropical Deltocephalinae and some related subfamilies (Homoptera). Annales Zoologici Societatis Zoologicae Botanicae Fennicae ‘Vanamo’ 20 (l): 1–370.
  21. Linnavuori R. E. 1960. Homoptera: Cicadellidae. Insects of Micronesia 6(5): 225–344.
  22. Linnavuori R. E. 1975. Homoptera: Cicadellidae, Supplement. Insects of Micronesia 6(9): 611–632.
  23. Lu L., Zhang Y. L. & Webb M. D. 2013. Review of the grass feeding leafhopper genus Balclutha Kirkaldy (Hemiptera: Cicadellidae: Deltocephalinae) in China. Zootaxa 3691(5): 501–537.
  24. Mahmood S. H. 1979. A revision of the leafhoppers (Cicadellidae: Homoptera) of Pakistan and adjoining countries of the Oriental region. Department of Zoology (Entomology), University of Karachi 10: 103.
  25. Matsumura S. 1902. Monographie der Jassinen Japans. Természetrajzi Füzetek. Kiadja a Magyar Nemzeti Museum Budapest 25: 353–404.
  26. Matsumura S. 1914. Die Jassinen und einige neue Acocephalinen Japans. Journal of the College of Agriculture, Tohoku Imperial University, Sapporo 5: 165–240.
  27. Melichar L. 1903. Homopteren-Fauna von Ceylon. Pp. 248 in: F.L. Dames, Berlin.
  28. Melichar L. 1905. Beitrag zur Kenntnis der Homopteren fauna Deutsch-Ost-Afrikas. Wiener Entomologische Zeitung 24: 279–304.
  29. Merino G. 1936. Philippine Cicadellidae (Homoptera). Philippine Journal of Science 61: 307–400.
  30. Metcalf Z. P. 1946. Homoptera. Fulgoroidea and Jassoidea of Guam. Bulletin of the Bernice P. Bishop Museum 189: 105–148.
  31. Mozaffarian F. & Wilson M. R. 2016. A checklist of the leafhoppers of Iran (Hemiptera: Auchenorrhyncha: Cicadellidae). Zootaxa 4062(1): 1–63.
  32. Naveed H. & Zhang Y. L. 2018. Review of the leafhopper tribe Macrostelini Kirkaldy (Cicadellidae: Deltocephalinae) in Pakistan with description of a new species. Zootaxa 4462(2): 257–273.
  33. Naveed H., Shah B., Khan B. S., Cao C., Webb M. & Zhang Y. L. 2021. Checklist and keys to Deltocephalinae leafhoppers (Hemiptera: Cicadellidae) from Pakistan. ZooKeys 1078: 135–188.
  34. Oman P. W. 1949. The Nearctic leafhoppers (Homoptera: Cicadellidae). A generic classification and checklist. Memoirs of Entomological Society of Washington 3: 1–253.
  35. Pander C. H. 1856. Monographie der fossilen Fische des Silu-rischen Systems der russisch-baltischen Gouvernements. Pp. 91, In: Akademie der Wissenschaften, St. Petersburg.
  36. Singh-Pruthi H. 1930. Studies on Indian Jassidae (Homoptera). Part 1. Introduction and description of some new genera and species. Memoirs of Indian Museum 11: 1–68.
  37. Rama Subba Rao V. & Ramakrishnan U. 1990. Two new species and some new records of the genus Balclutha Kirk. from India (Insecta, Homoptera, Auchenorrhyncha, Cicadellidae: Balcluthini). Reichenbachia 27(2): 105–108.
  38. Ramachandra Rao K. 1986. On a Collection of Leafhoppers (Cicadellidae Hemiptera) from the Silent Valley. Records of the Zoological Survey of India 84(1–4): 49–58.
  39. Rao V. & Ramakrishnan U. 1990. New species of Aconurella and Paramesodes (Homoptera: Cicadellidae) from India. Journal of the Bombay Natural History Society 87(2): 268–271.
  40. Sun J., Zhang Y. L., Lu L. & Webb M. D. 2019. A new species and new record of the leafhopper genus Balclutha (Hemiptera: Cicadellidae: Deltocephalinae) from China with key to species. Zootaxa 4551(4): 494–500.
  41. Webb M. D. & Vilbaste J. 1994. Review of the leafhopper genus Balclutha Kirkaldy in the Oriental Region (Insecta: Homoptera: Auchenorrhyncha: Cicadellidae). Entomologische Abhandlungen Staatliches Museum für Tierkunde Dresden 56: 56–86.