Abstract
Thirteen species of Helicopsyche, subgenus Feropsyche (Trichoptera: Helicopsychidae) are recorded from Costa Rica, including 3 new species: H. alajuela, n. sp., H. dorsocurvata, n. sp., and H. golfitoensis, n. sp., as well as 10 previously described species: H. borealis (Hagen), H. chiriquensis Johanson & Malm, H. dampfi Ross, H. incisa Ross, H. lewalleni Denning & Blickle, H. mexicana Banks, H. rentzi Denning & Blickle, H. selanderi Ross, H. truncata Ross, and H. vergelana Ross. The male genitalia of the new species are illustrated and distribution maps are provided for all species.References
Bueno-Soria, J. & Flint, O.S., Jr. (1978) Catálogo sistemático de los Tricópteros de México (Insecta: Trichoptera), con algunos registros de Norte, Centro y Sudamérica. Anales del Instituto de Biologia, Universidad Nacional Autónoma de México, Series de Zoologia, 57, 299–310.
Denning, D.G. & Blickle, R.L. (1979) New species of Helicopsyche (Trichoptera: Helicopsychidae). Pan-Pacific Entomologist, 55, 27–33.
Flint, O.S., Jr. (1986) Studies of Neotropical caddisflies, XXXVI: the genus Cochliopsyche in Middle America (Trichoptera: Helicopsychidae). Anales del Instituto de Biología, Universidad Nacional Autónoma de México, Serie Zoología, 56, 213–216.
Flint, O.S., Jr. Holzenthal, R.W. & Harris, S.C. (1999) Catalog of the Neotropical Caddisflies (Trichoptera). Ohio Biological Survey, Columbus, Ohio, 1–239 pp.
Flint, O.S., Jr. & Sykora, J.L. (2004) Caddisflies of Hispaniola, with special reference to the Dominican Republic (Insecta: Trichoptera). Annals of the Carnegie Museum, 73, 1–60.
Hagen, H.A. (1861) Synopsis of the Neuroptera of North America with a list of the South American species. Smithsonian Miscellaneous Collections, 4 (1), xx+347 pp.
Holzenthal, R.W. (1988) Catalogo systematico de los Trichopteros de Costa Rica (Insecta: Trichoptera). Brenesia, 29, 51–82.
Holzenthal, R.W., Blahnik, R.J., Prather, A.L. & Kjer, K.M. (2007) Order Trichoptera Kirby 1813 (Insecta), Caddisflies. In, Z.-Q. Zhang & W.A. Shear (editors). Linneaus Tercentenary: Progress in Invertebrate Taxonomy, Zootaxa, 1668, 639–701.
Jackson, J.K. & Sweeney, B.W. (1995) Egg and larval development times for 35 species of tropical stream insects from Costa Rica. Journal of the North American Benthological Society, 14, 115–130.
Johanson, K.A. (1997) Zoogeography and diversity of the snail case caddisflies (Trichoptera: Helicopsychidae). In: Holzenthal, R.W. & Flint, O.S., Jr. (Eds.), Proceedings of the 8th International Symposium on Trichoptera. Ohio Biological Survey, pp. 205–212.
Johanson, K.A. (1998) Phylogenetic and biogeographic analysis of the family Helicopsychidae (Insecta: Trichoptera). Entomologica scandinavica, Supplement, 53, 1–172.
Johanson, K.A. (2002) Systematic revision of American Helicopsyche of the subgenus Feropsyche (Trichoptera: Helicopsychidae). Insect Systematics and Evolution, Supplement, 60, 1–146.
Johanson, K.A. (2003a) The Helicopsyche (Feropsyche) (Insecta, Trichoptera, Helicopsychidae) from Barro Colorado Island, Panama. Zootaxa, 283, 1–12.
Johanson, K.A. (2003b) Revision of the Neotropical caddisfly subgenus Helicopsyche (Cochliopsyche) (Trichoptera, Helicopsychidae), with descriptions of twelve new species. Insect Systematics and Evolution, 34, 381–414.
Johanson, K.A. (2003c) The sister species of Jamaican Helicopsyche kingstona sp.n., is Mexican H. villegasi Denning & Blickle (Trichoptera, Helicopsychidae). Tijdschrift voor Entomologie, 146, 33–37.
Johanson, K.A. (2006) Check-list of New Caledonian Trichoptera. http://www2.nrm.se/en/caledonia/trichopteranc.html [Accessed 26 September 2008.]
Johanson, K.A. & Holzenthal, R.W. (2004) Thirteen new species and new distribution records of Helicopsyche (Feropsyche) Johanson from Venezuela (Trichoptera: Helicopsychidae). Zootaxa, 711, 1–40.
Johanson, K.A. & Malm, T. (2006) Seven new Helicopsyche (Feropsyche) Johanson, 2002 from the Neotropical region and Nearctic Mexico (Insecta: Trichoptera: Helicopsychidae). Zootaxa, 1208, 1–24.
Monson, M.P., Holzenthal, R.W. & Ahlstrand, G.G. (1988) The larvae and pupa of Cochiliopsyche vazquezae (Trichoptera: Helicopsychidae). Journal of the North American Benthological Society, 7, 152–159.
Morse, J.C. (2008) Trichoptera World Checklist. http://entweb.clemson.edu/database/trichopt/index.htm [Accessed 26 September 2008.]
Müller, F. (1885) Wie Ensteht die Gleiderung der Insektenfuhler? Kosmos, 17, 201–204.
Oláh, J. & Johanson, K.A. (2008) Reasoning an appendicular and functional caddisfly genital terminology. Braueria, 35, 29–40.
Ross, H.H. (1944) The caddisflies or Trichoptera of Illinois. Bulletin of the Illinois Natural History Survey, 23, 1–326.
Ross, H.H. (1956) New species of Helicopsyche from the western hemisphere (Trichoptera, Helicopsychidae). Journal of the Washington Academy of Sciences, 46, 397–401.