Abstract
Twenty sponge species (totalling 190 individuals) were collected during the 1938, 1994 and 2004/5 expeditions to the remote island of Clipperton in the East Pacific Ocean. Seven species are widespread Indo-Pacific sponges; nine species comprise sponges new to science; four species were represented only by small thin patches insufficient for proper characterization and could be only determined to genus. The new species may not be necessarily endemic to the island, as several show similarities with species described from elsewhere in the East and West Pacific. Four species: Tethya sarai Desqueyroux-Faúndez & Van Soest (1997), Callyspongia (Callyspongia) roosevelti n.sp., Spongia (Spongia) sweeti (Kirkpatrick, 1900) and Suberea etiennei n.sp. were found commonly occurring in localities around the island in depths between 10 and 55 m, growing on dead corals, under overhangs and rubble stones. The remaining sponges were either rare or were thinly encrusting on coral fragments. The latter may be more abundant than appears from the present study as they are probably not easily observed. The sponge fauna of Clipperton Island shows strongest affinities with the Central and West Pacific regions and only two or three species are shared with the East Pacific region.References
Bakus, G.J. (1975) Marine zonation and ecology of Cocos Island, off Central America. Atoll Research Bulletin, 179, 1–8.
Béarez, P. & Séret, B. (2008) Les poissons de Clipperton. In: L. Charpy (ed.), Clipperton: environnement et biodiversité d’un microcosme océanique. Patrimoines Naturels, Muséum national d’Histoire naturelle & IRD Éditions, Paris.
Bergquist, P.R. (1965) The sponges of Micronesia, Part I. The Palau Archipelago. Pacific Science, 19 (2), 123–204.
Bergquist, P.R. (1968) The marine fauna of New Zealand: Porifera, Demospongiae, Part 1. (Tetractinomorpha and Lithistida). New Zealand Oceanographic Institute Memoir, 37, 1–105.
Bergquist, P.R. (1977) Porifera. In: Devaney D. M. & Eldredge L.G. (eds), Reef and shore fauna of Hawaii. Section I. Protozoa through Ctenophora. B.P. Bishop Museum Press, Honolulu, p. 53–69.
Bergquist, P.R. (1980) A revision of the supraspecific classification of the orders Dictyoceratida, Dendroceratida and Verongida (class Demospongiae). New Zealand Journal of Zoology, 7 (4), 443–503.
Bergquist, P.R. (1995) Dictyoceratida, Dendroceratida and Verongida from the New Caledonia Lagoon (Porifera: Demospongiae). Memoirs of the Queensland Museum, 38 (1), 1–51.
Bergquist, P.R. & Fromont, P.J. (1988) The marine fauna of New Zealand: Porifera, Demospongiae, Part 4 (Poecilosclerida). New Zealand Oceanographic Institute Memoir, 96, 1–197.
Bergquist, P.R. & Kelly, M. (2004) Taxonomy of some Halisarcida and Homosclerophorida (Porifera: Demospongiae) from the Indo—Pacific. New Zealand Journal of Marine and Freshwater Research, 38, 51—66.
Borojevic, R., Boury-Esnault, N., Manuel, M. & Vacelet, J. (2002) Order Leucosolenida Hartman, 1958. In: Hooper, J.N.A. & Van Soest, R.W.M. (eds), Systema Porifera: a guide to the classification of sponges. Kluwer Plenum Academic Press, London, New York, p 1157–1184.
Borojevic, R. & Klautau, M. (2000) Calcareous sponges from New Caledonia. Zoosystema, 22 (2), 187–201.
Boury-Esnault, N. (1973) Résultats scientifiques des campagnes de la ‘Calypso’. Campagne de la ‘Calypso’ au large des côtes atlantiques de l’Amérique du Sud (1961–1962). I. 29. Spongiaires. Annales de l’Institut océanographique, 49 (Supplément 10), 263–295.
Bowerbank, J.S. (1864) A monograph of the British Spongiadae.Volume 1. Ray Society, London, i–xx, 1–290, pls I–XXXVII.
Burton, M. (1930) The Porifera of the Siboga Expedition. III. Calcarea. In: Weber, M. (ed.), Siboga-Expeditie. Uitkomsten op zoologisch, botanisch, oceanographisch en geologisch gebied verzameld in Nederlandsch Oost-lndië 1899–1900 aan boord H.M. Siboga onder commando van Luitenant ter zee 1e kl. G. F. Tydeman, III (Monographie VIa2). E.J. Brill, Leiden, p 1–18.
Burton, M. (1932) Report on a collection of sponges made in South Saghalin by Mr. Tomoe Urita. Science Reports of the Tôhoku Imperial University, (ser. 4, Biology), 7 (2), 195—206.
Burton, M. (1934) Sponges. Scientific Reports of the Great Barrier Reef Expedition 1928–29, 4 (14), 513–621.
Burton, M. (1963) Revision of the classification of the calcareous sponges. British Museum (Natural History) London, p i–v, 1–693.
Carballo, J.L. & Cruz-Barraza, J.A. (2006) The genus Timea (Porifera: Timeidae) in the Eastern Pacific Ocean: revision of type material, new species and intraspecific variability. Cahiers de Biologie Marine, 47 (3), 301–320.
Carballo J.L. & Nava, H. (2007) Comparison of sponge assemblage patterns between two adjacent tropical rocky habitats (tropical Pacific Ocean, Mexico). Ecoscience, 14, 92–102.
Carter, H.J. (1874) Descriptions and figures of deep-sea sponges and their spicules from the Atlantic Ocean, dredged up on board H.M.S.‘Porcupine’, chiefly in 1869; with figures and descriptions of some remarkable spicules from the Agulhas Shoal and Colon, Panama. Annals and Magazine of Natural History, (4) 14 (79), 207–221, 245–257, pls. XIII–XV.
Cook, S.D. (2007) Clarification of dictyoceratid taxonomic characters, and the determination of genera. In: Custódio M.R., Lôbo-Hajdu G., Hajdu E. & Muricy G. (eds). Porifera research, biodiversity, innovation and sustainability. (Série Livros 28). Museu Nacional, Rio de Janeiro, p 265–274.
Cook, S.D. & Bergquist, P.R. (2002) Order Dictyoceratida Minchin, 1900. In: Hooper, J.N.A. & Van Soest, R.W.M. (eds), Systema Porifera: a guide to the classification of sponges. Kluwer Plenum Academic Press, London, New York, p 1021.
Cruz—Barraza, J.A. & Carballo, J.L. (2005) First record of Plakortis Schulze (Porifera: Homosclerophorida) from the Northeast Pacific coast, with the description of Plakortis albicans sp. nov. Zootaxa, 868, 1–12.
Cruz—Barraza, J.A. & Carballo, J.L. (2008) Taxonomy of sponges (Porifera) associated with corals from the Mexican Pacific Ocean. Zoological Studies 47 (6), 741-758.
Dendy, A. (1905) Report on the sponges collected by Professor Herdman, at Ceylon, in 1902. In: Herdman W.A. (ed.), Report to the government of Ceylon on the pearl oyster fisheries of the Gulf of Manaar, 3 (Supplement 18). Royal Society: London, p 57–246, pls I–XVI.
Dendy, A. (1913) The Percy Sladen trust expedition to the Indian Ocean in 1905 (V). I. Report on the Calcareous Sponges collected by HMS ‘Sealark’ in the Indian Ocean. Transactions of the Linnean Society of London, (2) 16 (1), 1–29.
Dendy, A. (1924) Porifera. Part I. Non—Antarctic sponges. Natural History Report British Antarctic (Terra Nova) Expedition, 1910, (Zoology) 6 (3), 269–392, pls I–XV.
Desqueyroux-Faúndez, R. (1990) Sponges (Demospongiae) from Easter Island (Isla de Pascua) (South Pacific Ocean). Revue Suisse de Zoologie, 97(2), 373–410.
Desqueyroux-Faúndez, R. & Van Soest, R.W.M. (1997) Shallow water Demosponges of the Galápagos Islands. Revue suisse de Zoologie, 104 (2), 379–467.
Díaz, M.C. & Van Soest, R.W.M. (1994) The Plakinidae: A systematic review. In: Van Soest R.W.M., van Kempen Th.M.G. & Braekman J.C. (eds) Sponges in time and space. Balkema, Rotterdam, p 93–109.
Dickinson, M.G. (1945) Sponges of the Gulf of California. Reports on the collections obtained by Alan Hancock Pacific Expeditions of Velero III off the coast of Mexico, Central America, South America, and Galapagos Islands in 1932, in 1933, in 1934, in 1935, in 1936, in 1937, in 1939, and in 1940. The University of Southern California Press, Los Angeles:, p 1–55, pls 1–97.
Duchassaing De Fonbressin, P. & Michelotti, G. (1864) Spongiaires de la mer Caraïbe. Natuurkundige verhandelingen van de Hollandsche maatschappij der wetenschappen te Haarlem, 21 (2). 1–124, pls I–XXV.
Ereskovsky, A.V. (2006) A new species of Oscarella (Demospongiae: Plakinidae) from the Western Sea of Japan. Zootaxa,1376, 37–51.
Glynn, P.W., Veron, J.E.N. & Wellington, G.M. (1996) Clipperton Atoll (Eastern Pacific): oceanography, geomorphology, reef—building coral ecology and biogeography. Coral Reefs, 15, 71–99.
Gómez, P. & Bakus, G. (1992) Aplysina gerardogreeni and Aplysina aztecus (Porifera: Demospongiae) new species from the Mexican Pacific. Anales del Instituto de Ciencias del Mar y Limnología, Universidad Autónoma de México, 19 (2), 175–180.
Grant, R.E. (1836) Animal Kingdom. In: Todd R.B. (ed.), The Cyclopaedia of Anatomy and Physiology. Volume 1. Sherwood, Gilbert, and Piper, London, p. 107–118.
Gray, J.E. (1867) Notes on the arrangement of sponges, with the descriptions of some new genera. Proceedings of the Zoological Society of London 1867, (2), 492–558, pls XXVII–XXVIII.
Green, G. & Gómez, P. (1986) Estudio taxonómico de las esponjas de la Bahía de Mazatlán, Sinaloa, México. Anales del Instituto de Ciencias del Mar y Limnología, Universidad Autónoma de México, 13 (3), 273–300.
Haeckel, E. (1872) Die Kalkschwämme. Eine Monographie in zwei Bänden Text und einem Atlas mit 60 Tafeln Abbildungen. G. Reimer, Berlin, 1: 1–484, 2: 1–418, 3: pls 1–60.
Hallmann, E.F. (1917) A revision of the genera with microscleres included, or provisionally included, in the family Axinellidae; with descriptions of some Australian species. Part III. Proceedings of the Linnean Society of New South Wales, 41 (164), 634–675.
Hentschel, E. (1909) Tetraxonida. 1. Teil. In: Michaelsen W. & Hartmeyer R. (eds), Die Fauna Südwest-Australiens. Ergebnisse der Hamburger südwest-australischen Forschungsreise 1905.Volume 2 (21). Fischer, Jena, p 347–402.
Hentschel, E. (1911) Tetraxonida. 2. Teil. In: Michaelsen W. & Hartmeyer R. (eds), Die Fauna Südwest-Australiens. Ergebnisse der Hamburger südwest-australischen Forschungsreise 1905. Volume 3 (10). Fischer, Jena, p 279–393.
Hentschel, E. (1912) Kiesel- und Hornschwämme der Aru- und Kei-Inseln. Abhandlungen herausgegeben von der Senckenbergischen naturforschenden Gesellschaft, 34 (3), 293–448.
Hertlein, L. G. & Emerson, W.K. (1957) Additional notes on the invertebrate fauna of Clipperton Island. American Museum of Natural History Novitates, 1859, 1–9.
Hooper, J.N.A. (1994) Coral reef sponges of the Sahul Shelf – a case for habitat preservation. Memoirs of the Queensland Museum, 36 (1), 93–106.
Hooper, J.N.A. (1996) Revision of Microcionidae (Porifera: Poecilosclerida: Demospongiae), with description of Australian species. Memoirs of the Queensland Museum, 40, 1–626.
Hooper, J.N.A. & Van Soest, R.W.M. (eds) (2002) Systema Porifera: a guide to the classification of sponges. Volumes 1 and 2. Kluwer Academic Plenum Publishers, London, New York, p i–xlviii, 1–1706.
Hoshino, T. (1977) Demosponges from the Kii channel and its environs, western Japan. Proceedings of the Japanese Society of Systematic Zoology, 13, 5–15.
Hoshino, T. (1981) Shallow-water Demosponges of Western Japan, 1. Journal of Science of the Hiroshima University, (B) 29 (1), 47–205.
Jinhe, L. (1986) Sponges as marine fouling organisms in China waters. I. Studia Marina Sinica, 26 (2), 76–116.
Kaiser, K.L. (2007) The Recent molluscan fauna of Île Clipperton (Tropical Eastern Pacific). The Festivus 39 (Supplement), i–iii, 1–162.
Kelly-Borges, M. & Bergquist, P.R. (1988) Sponges from Motupore Island, Papua New Guinea. Indian-Malayan Zoology, 5, 121–159.
Kelly-Borges, M. & Valentine, C. (1995) The sponges of the tropical island region of Oceania: A taxonomic status review. In: Maragos, J.E., Peterson, M.N.A., Eldredge, L.G., Bardach, J.E. & Takeuchi, H.F. (eds), Marine and coastal biodiversity in the tropical island Pacific region. Volume 1 - Species Systematics and Information Management Priorities. Chapter 6. Honolulu, Hawaii, p 83–120.
Kennedy, J.A. & Hooper, J.N.A. (2000) New species of toxic Tedania from northern Vanuatu (Porifera: Demospongiae: Poecilosclerida: Tedaniidae). Memoirs of the Queensland Museum, 45 (2), 445–451.
Kirkpatrick, R. (1900) Description of sponges from Funafuti. Annals and Magazine of Natural Histor, (7) 6 (34), 345–362.
Klautau, M. & Valentine, C. (2003) Revision of the genus Clathrina (Porifera, Calcarea). Zoological Journal of the Linnean Society, 139 (1), 1–62.
Lamarck, J.B.P. De (1814) Suite du mémoire intitulé: sur les polypiers empâtés. Suite des éponges. Annales du Muséum national d’Histoire naturelle, Paris, 20 (6), 294–312.
Lambe, L.M. (1895) Sponges from the Western Coast of North America. Transactions of the Royal Society of Canada, 12 (4), 113–138.
Laubenfels, M.W. De (1930) The sponges of California. (Abstracts of dissertations for the degree of doctor of philosophy). Stanford University Bulletin, (5) 5 (98), 24–29.
Laubenfels, M.W. De (1932) The marine and fresh-water sponges of California. Proceedings of the United States National Museum, 81 (2927), 1–140.
Laubenfels, M.W. De (1936a). A discussion of the sponge fauna of the Dry Tortugas in particular and the West Indies in general, with material for a revision of the families and orders of the Porifera. Carnegie Institute of Washington (Tortugas Laboratory Paper N° 467), 30, 1–225, pls 1–22.
Laubenfels, M.W. De (1936b) A comparison of the shallow-water sponges near the Pacific end of the Panama Canal with those at the Caribbean end. Proceedings of the United States National Museum, 83 (2993), 441–466.
Laubenfels, M.W. De (1939) Sponges collected on the presidential cruise of 1938. Smithsonian Miscellaneous Collections, 98 (15), 1–7.
Laubenfels, M.W. De (1950) The sponges of Kaneohe Bay, Oahu. Pacific Science, 4,3–36.
Laubenfels, M.W. De (1951) The sponges of the Islands of Hawaii. Pacific Science, 5, 256—271.
Laubenfels, M.W. De (1954a) The sponges of the West-Central Pacific. Oregon State Monographs. Studies in Zoology, 7, i–x, 1–306, pls I–XII.
Laubenfels, M.W. De (1954b) Occurrence of sponges in an aquarium. Pacific Science, 8, 337–340.
Laubenfels, M.W. De (1957) New species and records of Hawaiian sponges. Pacific Science, 11, 236–251.
Lendenfeld, R. Von (1888) Descriptive catalogue of the sponges in the Australian Museum, Sidney. Taylor & Francis, London, p i–xiv, 1–260, pls 1–12.
Lendenfeld, R. Von (1897) Spongien von Sansibar. Abhandlungen herausgegeben von der Senckenbergischen naturforschenden Gesellschaft, 21, 93–133.
Lendenfeld, R. Von (1907) Die Tetraxonia. Wissenschaftliche Ergebnisse der Deutschen Tiefsee — Expedition auf der Dampfer Valdivia 1898–1899, 11 (1–2), i–iv, 59–374, pls IX–XLVI.
Lévi, C. (1953) Description de Plakortis nigra nov. sp. et remarques sur les Plakinidae (Démosponges). Bulletin du Muséum national d’Histoire naturelle, (2) 25 (3), 320–328.
Lévi, C. (1961) Résultats scientifiques des Campagnes de la ‘Calypso’. Campagne 1954 dans l’Océan Indien (suite). 2. Les spongiaires de l’Ile Aldabra. Annales de l’Institut océanographique, 39 (1), 1–32.
Lévi, C. & Lévi P. (1983) Démosponges bathyales récoltées par le N/O‘Vauban’ au sud de la Nouvelle—Calédonie. Bulletin du Muséum national d’Histoire naturelle, (4, A) 5 (4), 931–997.
Lévi, C. & Lévi, P. (1989) Spongiaires (MUSORSTOM 1 & 2). In: Forest J. (ed.), Résultats des Campagnes MUSORSTOM, Volume 4. Mémoires du Muséum national d'Histoire naturelle, (A, Zoologie), 143, 25–103.
Lewis J.B. (1965) A preliminary description of some marine benthic communities from Barbados, West Indies. Canadian Journal of Zoology, 43, 1049–1063.
Linnæus, C. (1759) Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus II. Editio decima, reformata. L. Salvii, Holmiæ, p 825–1384.
Lundbeck, W. (1910) Porifera. (Part III.) Desmacidonidae (pars.). The Danish Ingolf-Expedition, 6 (3), 1–124, pls I–XI.
Maldonado, M. (1992) Demosponges of the red coral bottoms from the Alboran Sea. Journal of Natural History, 26, 1131–1161.
Marenzeller, E. Von (1877) Die Coelenteraten, Echinodermen und Würmer der K.K.Österreichisch-Ungarischen Nordpol-Expedition. Denkschriften der Mathematisch—naturwissenschaftlichen Classe der K. Akademie der Wissenschaften von Wien, 35, 357–398.
Moraes, F.C. & Muricy, G. (2003) Taxonomy of Plakortis and Plakinastrella (Demospongiae: Plakinidae) from oceanic islands off north-eastern Brazil, with description of three new species. Journal of the Marine Biological Association of the United Kingdom, 83 (2), 385–397.
Mothes, B., Hajdu, E. & Van Soest, R.W.M. (2000) Tedania brasiliensis new species (Demospongiae, Poecilosclerida, Tedaniidae) from Brazil, with some remarks about the genus Tedania in the tropical southwestern Atlantic. Bulletin of Marine Science, 66 (1), 1–11.
Muricy, G. & Díaz, M.C. (2002) Order Homosclerophorida Dendy, 1905. Family Plakinidae Schulze, 1880. In: Hooper, J.N.A. & Van Soest, R.W.M. (eds), Systema Porifera: a guide to the classification of sponges. Kluwer Plenum Academic Press, London, New York, p 71–82.
Muricy, G. & Pearse, J.S. (2004) A new species of Oscarella (Demospongiae: Plakinidae) from California. Proceedings of the California Academy of Sciences, 55 (33), 598–612.
Pulitzer—Finali, G. (1978) Report on a collection of sponges from the Bay of Naples. III. Hadromerida, Axinellida, Poecilosclerida, Halichondrida, Haplosclerida. Bollettino dei Musei e degli Istituti Biologici della (R.) Università di Genova, 45, 7–89.
Pulitzer—Finali, G. (1996) Sponges from the Bismarck Sea. Bollettino dei Musei e degli Istituti Biologici della (R.) Università di Genova, 60–61, 101–138.
Robertson, D.R. & Allen, G.R. (1996) Zoogeography of the shorefish fauna of Clipperton Atoll. Coral Reefs, 15, 121—131.
Rützler, K. (2002) Family Timeidae Topsent, 1928. In: Hooper, J.N.A. & Van Soest, R.W.M. (eds), Systema Porifera: a guide to the classification of sponges. Kluwer Plenum Academic Press, London, New York, p 266–267.
Salvat, B., Adjeroud, M. & Charpy, L. (2008) Les récifs coralliens de Clipperton. Revue d’Écologie (Terre Vie), 63:,179–187.
Sarà, M. & Bavestrello, G. (1995) Tethya omanensis, a remarkable new species from an Oman cave (Porifera, Demospongiae). Bolletino di Zoologia, 62, 23–27.
Sarà, M. & Bavestrello, G. (1998) Two new species of Tethya (Porifera, Demospongiae) from the Canary and Cape Verde Islands. Italian Journal of Zoology, 65, 371–376.
Sarà, M. & Siribelli, L. (1960) La fauna di Poriferi delle ‘secche’ del Golfo di Napoli. 1. La ‘secca’ della Gaiola. Annuario dell’Istituto e Museo de Zoologia dell’Università di Napoli, 12 (3), 1–93.
Sarà, M., Gómez, P. & Sarà, A. (2001) East Pacific Mexican Tethya (Porifera: Demospongiae) with descriptions of five new species. Proceedings of the Biological Society of Washington, 114 (3), 794–821.
Schmidt, O. (1862) Die Spongien des adriatischen Meeres. Engelmann, Leipzig, p i–viii, 1–88, pls I–VII.
Schmidt, O. (1868) Die Spongien der Küste von Algier. Mit Nachträgen zu den Spongien des Adriatischen Meeres (Drittes Supplement). Engelmann, Leipzig, p i–iv, 1–44, pls I–V.
Schmidt, O. (1870) Grundzüge einer Spongien—Fauna des atlantischen Gebietes. Engelmann, Leipzig, p iii–iv, 1–88, pls I–VI.
Schuffner, O. (1877) Beschreibung einiger neuer Kalkschwämme. Jenaische Zeitschrift für Naturwissenschaft, 11, 403–433.
Schulze, F.E. (1880) Untersuchungen über den Bau und die Entwicklung der Spongien. Neunte Mittheilung. Die Plakiniden. Zeitschrift für wissenschaftliche Zoologie, 34 (2) 407–451.
Sim, C.J. & Bakus, G.J. (1986) Marine sponges of Santa Catalina Island, California. Occasional Papers of the Allan Hancock Foundation, 5, 1–23.
Szymanski, J.M. (1904) Zur Anatomie und Systematik der Hornschwämme des Mittelmeeres. Zoologischen Anzeiger, 27 (6), 445–449.
Thiele, J. (1898) Studien über pazifische Spongien. I. Japanische Demospongien. Zoologica. Original — Abhandlungen aus dem Gesamtgebiete der Zoologie. Stuttgart, 24 (1), 1–72, pls I–VIII.
Thiele, J. (1899) Studien über pazifische Spongien. II. Ueber einige Spongien von Celebes. Zoologica. Original-Abhandlungen aus dem Gesamtgebiete der Zoologie Stuttgart, 24 (2), 1–33, pls I–V.
Thiele, J. (1903) Kieselschwämme von Ternate. II. Abhandlungen herausgegeben von der Senckenbergischen naturforschenden Gesellschaft, 25, 933–968, pl. XVIII.
Thiele, J. (1905) Die Kiesel- und Hornschwämme der Sammlung Plate. Zoologische Jahrbücher Supplement, 6 (Fauna Chilensis III), 407–496, pls 27–33.
Thomson, C.W. (1873) The depths of the sea. Macmillan and Co., London, p 1–527.
Topsent, E. (1890) Notice préliminaire sur les spongiaires recueillis durant les campagnes de l'Hirondelle. Bulletin de la Société zoologique de France, 15, 26–32, 65–71.
Topsent, E. (1892) Contribution à l’étude des Spongiaires de l’Atlantique Nord (Golfe de Gascogne, Terre—Neuve, Açores). Résultats des campagnes scientifiques accomplies par le Prince Albert I. Monaco, 2, 1–165, pls I–XI.
Topsent, E. (1906) Éponges recueillies par M. Ch. Gravier dans la Mer Rouge. Bulletin du Muséum national d’Histoire naturelle, (1) 7, 557–570.
Topsent, E. (1927) Diagnoses d'éponges nouvelles recueillies par le Prince Albert ler de Monaco. Bulletin de l'Institut Océanographique Monaco, 502, 1–19.
Topsent, E. (1931) Éponges de Lamarck conservées au Muséum de Paris. Archives du Muséum national d’Histoire naturelle Paris, (6) 5, 1–6, pls I–IV.
Topsent, E. (1934) Sur les Aplysinopsis Lend. Bulletin de l'Institut Océanographique Monaco, 655, 1–15.
Uliczka, E. (1929) Die tetraxonen Schwämme Westindiens (auf Grund der Ergebnisse der Reise Kükenthal—Hartmeyer). In: Kükenthal, W. & Hartmeyer, R. (eds), Ergebnisse einer zoologischen Forschungsreise nach Westindien. Zoologische Jahrbücher Abteilung für Systematik, Geographie und Biologie der Thiere, Supplement, 16, 35–62.
Van Soest, R.W.M. (1984) Marine sponges from Curaçao and other Caribbean localities. Part III. Poecilosclerida. Studies on the Fauna of Curaçao and other Caribbean Islands, 62 (191), 1–173.
Van Soest, R.W.M. (2009) New sciophilous sponges from the Caribbean (Porifera: Demospongiae). Zootaxa, 2107, 1–40.
Van Soest, R.W.M. & Beglinger, E.J. (2008) Tetractinellid and hadromerid sponges from the Sultanate of Oman. Zoölogische Mededelingen Leiden, 82 (53), 749–790.
Van Soest, R.W.M., De Kluijver, M.J., Van Bragt, P.H., Faasse, M., Nijland, R., Beglinger, E.J., De Weerdt, W.H. & De Voogd, N.J. (2007) Sponge invaders in Dutch coastal waters. Journal of the Marine Biological Association of the United Kingdom, 87 (6), 1733–1748.
Van Soest, R.W.M., Boury-Esnault, N., Hooper, J.N.A., Rützler, K., De Voogd, N.J., Alvarez, B., Hajdu, E., Pisera, A.B., Vacelet, J., Manconi, R., Schoenberg, C., Janussen, D., Tabachnick, K.R., Klautau, M. & Picton, B. (2008) World Porifera database. Available online at http://www.marinespecies.org/porifera. Consulted on 2010–01–07.
Vosmaer, G.C.J. (1887) Klassen und Ordnungen der Spongien (Porifera). In: Bronn, H.G. (ed.), Die Klassen und Ordnungen des Thierreichs. 2. Winter, Leipzig & Heidelberg, p i–xii, 1–496, pls I–XXXIV.
Wörheide, G. & Hooper, J.N.A. (1999) Calcarea from the Great Barrier Reef. 1: Cryptic Calcinea from Heron Island and Wistari Reef (Capricorn-Bunker Group). Memoirs of the Queensland Museum, 43 (2), 859–891.
Zullov, V.A. (1969) A new subspecies of Balanus tintinnabulum (Linnaeus, 1758) (Cirripedia, Thoracica) from Clipperton Island, Eastern Pacific. Proceedings of the California Academy of Sciences, (4) 36 (16), 501–510.