Abstract
Alpha taxonomy of Vietnamese ants was initiated by European and American authors in the early 20th century, and approximately 160 species and infraspecific taxa were described or recorded in this early period. From 1965 to 1966 an inventory project of insects in northern Vietnam was conducted by the Agriculture Ministry of Vietnam, and 36 ant species were recorded. The identity of those taxa, however, needs to be revised based on the modern taxonomy of ants. Since theend of the 1980's dozens of ant species have been newly recorded or described from Vietnam. Regional revisions dealing with Vietnamese species have also been published for several genera including Myrmica, Probolomyrmex, Pheidole, Acanthomyrmex and Anillomyrma. Furthermore, two new genera, Parvimyrma (Myrmicinae) and Opamyrma (Amblyoponinae), were recently described from the country. However, there has been no comprehensive systematic treatment of the Vietnamese ants to date. In order to develop the taxonomy of ants in Vietnam, we have begun a program to: (1) explore local ant faunas; (2) give species codes to all species; (3) overview ant genera known from Vietnam; (4) describe unnamed taxa and review higher taxa known from Vietnam and adjacent areas. Ninety-two genera in twelve subfamilies have so far been recognized by us: DOLICHODERINAE: Chronoxenus, Dolichoderus, Iridomyrmex, Liometopum, Ochetellus, Philidris, Tapinoma, Technomyrmex; FORMICINAE: Acropyga, Anoplolepis, Camponotus, Cladomyrma, Echinopla, Gesomyrmex, Lasius, Lepisiota, Myrmoteras, Nylanderia, Oecophylla, Paratrechina, Paraparatrechina, Plagiolepis, Polyrhachis, Prenolepis, Pseudolasius, Undescribed genus "eg-2"; PSEUDOMYRMECINAE: Tetraponera; CERAPACHYINAE: Cerapachys, Simopone; AENICTINAE: Aenictus; DORYLINAE: Dorylus; LEPTANILLINAE: Leptanilla, Protanilla; AMBLYOPONINAE: Amblyopone, Myopopone, Mystrium, Opamyrma, Prionopelta; PONERINAE: Anochetus, Centromyrmex, Cryptopone, Diacamma, Harpegnathos, Hypoponera, Leptogenys, Odontomachus, Odontoponera, Pachycondyla, Platythyrea, Ponera; ECTATOMMINAE: Gnamptogenys; PROCERATIINAE: Discothyrea, Probolomyrmex, Proceratium; MYRMICINAE: Acanthomyrmex, Anillomyrma, Aphaenogaster, Calyptomyrmex, Cardiocondyla, Cataulacus, Crematogaster, Dacatria, Dilobocondyla, Gauromyrmex, Kartidris, Lasiomyrma, Liomyrmex, Lophomyrmex, Lordomyrma, Mayeriella, Meranoplus, Monomorium, Myrmecina, Myrmica, Myrmicaria, Oligomyrmex, Paratopula, Parvimyrma, Pheidole, Pheidologeton, Pristomyrmex, Proatta, Pyramica, Recurvidris, Rhopalomastix, Rhoptromyrmex, Solenopsis, Strumigenys, Temnothorax, Tetramorium, Vollenhovia, Vombisidris. As the first major contribution to the third goal of the program, here we provide: (1) a key to subfamilies, (2) a key to myrmicine genera, and (3) a synopsis of myrmicine and pseudomyrmecine genera known from Vietnam. A second paper will deal with the Aenictinae, Cerapachyinae, Dorylinae, Leptanillinae Amblyoponinae, Ponerinae, Ectatomminae and Proceratiinae, and a third with the Dolichoderinae and Formicinae.References
Baroni Urbani, C. & De Andrade, M.L. (2007) The ant tribe Dacetini: limits and constituent genera, with descriptions of new species. Annali del Museo Civico di Storia Naturale “G. Doria”, 28, 1–191.
Bingham, C.T. (1903) The fauna of British India, including Ceylon and Burma. Hymenoptera 2. Ants and Cuckoo-Wasps. London. 506pp.
Bolton, B. (1974) A revision of the Palaeotropical arboreal ant genus Cataulacus F. Smith. Bulletin of the British Museum (Natural History) (Entomology), 3, 1–105.
Bolton, B. (1976) The ant tribe Tetramoriini. Constituent genera, review of smaller genera and revision of Triglyphothrix Forel. Bulletin of the British Museum (Natural History) (Entomology), 34, 281–379.
Bolton, B. (1977) The ant tribe Tetramoriini. The genus Tetramorium Mayr in the Oriental and Indo-Australian regions, and in Australia. Bulletin of the British Museum (Natural History) (Entomology), 36, 67–151.
Bolton, B. (1981) A revision of the ant genera Meranoplus F. Smith, Dicroaspis Emery and Calyptomyrmex Emery (Hymenoptera: Formicidae) in the Ethiopian zoogeographical region. Bulletin of the British Museum (Natural History), 42, 43–81.
Bolton, B. (1982) Afrotropical species of the myrmicine ant genera Cardiocondyla, Leptothorax, Melissotarsus, Messor and Cataulacus (Formicidae). Bulletin of the British Museum (Natural History) (Entomology), 45, 307–370.
Bolton, B. (1986) A taxonomic and biological review of the tetramoriine ant genus Rhoptromyrmex. Systematic Entomology, 11, 1–17.
Bolton, B. (1988b) A review of Paratopula Wheeler, a forgotten genus of myrmicine ants. Entomologist’s Monthly Magazine, 124, 125–143.
Bolton, B. (1991) New myrmicine ant genera from the Oriental Region (Hymenoptera: Formicidae). Systematic Entomology, 16, 1–13.
Bolton, B. (1992) A review of the ant genus Recurvidris (Hym.: Formicidae), a new name for Trigonogaster Forel. Psyche, 99, 35–48.
Bolton, B. (1994) Identification Guide to the Ant Genera of the World: 222 pp. Cambridge, Massachusetts.
Bolton, B. (2000) The ant tribe Dacetini. Memoirs of the American Entomological Institute, 65, 1–1028.
Bolton, B. (2003) Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute, 71, 1–370.
Bolton, B., Alpert, G., Ward, P.S. & Naskrecki, P. (2007) Bolton’s Catalogue of Ants of the World: 1758–2005. Harvard University Press, Cabridge, Massachusetts, London, England.
Borowiec, M.L. (2009) New ant species related to Cerapachys sexspinus and discussion of the status of Yunodorylus (Hymenoptera: Formicidae). Zootaxa, 2069, 43–58.
Brady, S.G., Schultz, T.R., Fisher, B.L. & Ward, P.S. (2006) Evaluating alternative hypotheses for the early evolution and diversification of ants. Proceedings of the National Academy of Sciences, USA, 103, 18172–18177.
Branstetter, M.G. (2009) The ant genus Stenamma Westwood (Hymenoptera: Formicidae) redefined, with a description of a new genus Propodilobus. Zootaxa, 2221, 41–57.
Brown, W.L.Jr. (1953) Characters and synonymies among the genera of ants. Part 1. Breviora, 11, 1–13.
Brown, W.L.Jr. (1973) A comparison of the Hylean and Congo-West African rain forest ant faunas. Pp 161–185 in: Meggers, B.J., Ayensu, E.S. & Duckworth, W.D. (eds.). Tropical Forest Ecosystems in Africa and South America: a Comparative Review: 350 pp. Washington D.C.
Brühl, C. (2003) Vollenhovia sp. aff. acanthina: an ant species that builds nests using resin. ANeT Newsletter, No. 6, 21–22.
Bui, T.V. & Eguchi, K. (2003) Ant Survey in Hoang Lien Son Nature Reserve, Lao Cai, N. Vietnam. ANeT Newsletter, No. 5, 4–11.
Curtis, J. (1829) British Entomology; being illustrations and descriptions of the genera of insects found in Great Britain and Ireland, 6, 242–288. London.
Dubovikoff, D.A. (2004) A new species of the genus Gesomyrmex Mayr, 1868 (Hymenoptera: Formicidae) from Vietnam. Proceedings of the Russian Entomological Society, 75, 219–221.
Eguchi, K., Bui, T.V. & Yamane, Sk. (2004) A preliminary study on foraging distance and nesting sites of ants in Indo-Chinese lowland vegetation (Insecta, Hymenoptera, Formicidae). Sociobiology, 43, 445–457.
Eguchi, K. & Bui, T.V. (2005) A new South Vietnamese species of the genus Pheidole with a truncated head in the major and queen (Insecta, Hymenoptera, Formicidae). Sociobiology, 45, 721–730.
Eguchi, K., Bui, T.V., Yamane, Sk., Okido, H. & Ogata, K. (2005) Ant fauna of Ba Vi and Tam Dao, N. Vietnam (Insecta: Hymenoptera: Formicidae). Bulletin of the Institute of Tropical Agriculture, Kyushu University, 27, 77–98.
Eguchi, K. (2006) Six new species of Pheidole Westwood from North Vietnam (Insecta, Hymenoptera, Formicidae). Revue Suisse de Zoologie, 113, 115–131.
Eguchi, K. & Bui, T.V. (2006) Cladomyrma scopulosa new species (Hymenoptera: Formicidae: Formicinae) from Vietnam. Sociobiology, 47, 305–314.
Eguchi, K., Yoshimura, M. & Yamane, Sk. (2006) The Oriental species of the ant genus Probolomyrmex (Insecta: Hymenoptera: Formicidae: Proceratiinae). Zootaxa, 1376, 1–35.
Eguchi, K. & Bui, T.V. (2007) Parvimyrma gen. nov. belonging to the Solenopsis genus group from Vietnam (Hymenoptera: Formicidae: Myrmicinae: Solenopsidini). Zootaxa, 1461, 39–47.
Eguchi, K. & Bui, T.V. (2007) Ecological notes on a plant ant, Cladomyrma scopulosa Eguchi & Bui (Hymenoptera, Formicidae, Formicinae) associating with a tree species Saraca dives Pierre (Leguminosae). Asian Myrmecology, 1, 51–58.
Eguchi, K., Yamane, Sk. & Zhou, S.Y. (2007) Taxonomic revision of the Pheidole rinae Emery complex. Sociobiology, 50, 257–284.
Eguchi, K. (2008) A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae). Zootaxa, 1902, 1–118.
Eguchi, K., Bui, T.V. & Yamane, Sk. (2008) Vietnamese species of the genus Acanthomyrmex Emery, 1893 – A. humilis sp.n. and A. glabfemoralis Zhou & Zheng 1997 (Hymenoptera: Formicidae: Myrmicinae). Myrmecological News, 11, 231–241.
Eguchi, K. & Bui, T.V. (2009) Exploration of subterranean ant fauna by underground bait-trapping – A case study in Binh Chau-Phuoc Buu Nature Reserve, Vietnam (Insecta: Hymenoptera: Formicidae). Ari, 32, 21–25.
Eguchi, K., Bui, T.V., General, D.M. & Alpert, G.D. (2010) Revision of the ant genus Anillomyrma Emery, 1913 (Hymenoptera: Formicidae: Myrmicinae: Solenopsidini). Myrmecological News, 13, 31–36.
Emery, C. (1869) Enumerazione dei formicidi che rinvengonsi nei contorni di Napoli. Annali dell’Accademia degli Aspiranti Naturalisti, (2) 2, 1–26.
Emery, C. (1892) Sopra alcune formiche raccolte dall’Ingegnere L. Bricchetti Robecchi nel paese dei Somali. Annali del Museo Civico di Storia Naturale di Genova, (2) 12, 110–122.
Emery, C. (1893 [1892]) [Untitled. Introduced by: "M. C. Emery, de Bologne, envoie les diagnoses de cinq nouveaux genres de Formicides".] Annales de la Société Entomologique de France (Bulletin), 61, cclxxv–cclxxvii.
Emery, C. (1895) Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden. Zoologische Jahrbücher. Abtheilung für Systematik, Geographie und Biologie der Thiere, 8, 685–778.
Emery, C. (1897) Formicidarum species novae vel minus cognitae in collectionae Musaei Nationalis Hungarici, quas in Nova-Guinea, Colonia Germanica, collecit L. Biró. Természetrajzi Füzetek, 20, 571–599.
Emery, C. (1913) Etudes sur les Myrmicinae. Annales de la Societe Entomologique de Belgique, 57, 250–262.
Emery, C. (1914) Intorno alla classificazione dei Myrmicinae. Rendiconto delle Sessioni della R. Accademia delle Scienze dell’Istituto di Bologna (N.S.), 18, 29–42.
Ettershank, G. (1966) A generic revision of the world Myrmicinae related to Solenopsis and Pheidologeton (Hymenoptera: Formicidae). Australian Journal of Zoology, 14, 73–171.
Fernández, F. (2004) The American species of the myrmicine ant genus Carebara Westwood (Hymenoptera: Formicidae). Caldasia, 26, 191–238.
Fernández, F. (2007) Two new South American species of Monomorium Mayr with taxonomic notes on the genus. Memoirs of the American Entomological Institute, 80, 128–145.
Forel, A. (1890) Aenictus-Typhlatta découverte de M. Wroughton. Nouveaux genres de formicides. Annales de la Société Entomologique de Belgique, Comptes-rendus, 34, cii–cxiv.
Forel, A. (1900) Un nouveau genre et une nouvelle espèce de myrmicide. Annales de la Société Entomologique de Belgique, 44, 24–26.
Forel, A. (1902) Fourmis nouvelles d’Australie. Revue Suisse de Zoologie, 10, 405–548.
Forel, A. (1912) Descriptions provisoires de genres, sous-genres et espèces de formicides des Indes orientales. Revue Suisse de Zoologie, 20, 761–774.
Heterick, B. (2003) Two new Australian Monomorium Mayr (Hymenoptera: Formicidae), including a highly distinctive species. Australian Journal of Entomology, 42, 249–253.
Jaitrong, W. (2010) A new species of the ant genus Lasiomyrma (Hymenoptera, Formicidae, Myrmicinae) from Thailand. Sociobiology, 55, 427–431.
Japanese Ant Database Group (2003) Ants of Japan. 224 pp. Gakken, Tokyo, Japan.
Karavaiev, V. (1935) Neue Ameisen aus dem Indo-Australischen Gebiet, nebst Revision einiger Formen. Treubia, 15, 57–117.
Lapolla, J.S., Brady, S.G. & Shattuck, S.O. (2010) Phylogeny and taxonomy of the Prenolepis genus-group of ants (Hymenoptera: Formicidae). Systematic Entomology, 35, 118–131.
Latreille, P.A. (1804) Tableau méthodique des insectes. Classe huitième. Insectes, Insecta. Nouveau Dictionnaire d’Histoire Naturelle, 24, 129–200.
Lund, P.W. (1831) Lettre sur les habitudes de quelques fourmis du Brésil, adressée à M. Audouin. Annales des Sciences Naturelles, 23, 113–138.
Mayr, G. (1853) Beiträge zur Kenntniss der Ameisen. Verhandlungen des Zoologisch-Botanischen Vereins in Wien, 3, 101–114.
Mayr, G. (1855) Formicina austriaca. Beschreibung der bisher im österreichischen Kaiserstaate aufgefundenen Ameisen nebst Hinzufügung jener in Deutschland, in der Schweiz und in Italien vorkommenden Ameisen. Verhandlungen des Zoologisch-Botanischen Vereins in Wien, 5, 273–478.
Mayr, G. (1861) Die Europäischen Formiciden. (Ameisen.): 80 pp. Wien.
Mayr, G. (1865) Reise der Österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859, unter den befehlen des Commodore B. von Wüllerstrorf-Urbair. Zoologischer Theil. Formicidae: 119pp. Wien.
Mayr, G. (1866) Diagnosen neuer und wenig gekannter Formiciden. Verhandlungen der k.k. Zoologisch-Botanischen Gesellschaft in Wien, 16, 885–908.
Mayr, G. (1901) Südafrikanische Formiciden, gesammelt von Dr. Hans Brauns. Annalen des k.k. Naturhistorischen Hofmuseums Wien, 16, 1–30.
Menozzi, C. (1933) Description préliminaire d'une espèce nouvelle de fourmi constituant un genre nouveau. Natuurhistorisch Maandblad, 22, 146–147.
Moffett, M.W. (1986) Revision of the Myrmicine Genus Acanthomyrmex (Hymenoptera: Formicidae). Bulletin of the Museum of Comparative Zoology, 151, 55–89.
Moreau, C.S., Bell, C.D., Vila, R., Archibald, S.B. & Pierce, N.E. (2006) Phylogeny of the ants: diversification in the age of angiosperms. Science, 312, 101–104.
National Institute of Plant Protection (1976) Result of insect surveys 1967–1968. NXB Nong Thon. Pp. 399–404 (part). (In Vietnamese.)
Nguyen, K.V., Nguyen, T.H., Phan, K.L. & Nguyen, T.H. (2000) Bioclimatic Diagrams of Vietnam. Vietnam National University Publishing House (Hanoi). 129 pp.
Radchenko A.G. (1993a) New ants of the subfamily Cerapachyinae from Vietnam. Journal of the Ukranian Entomological Society, 1(1), 43–47.
Radchenko, A.G. (1993b) Ants from Vietnam in the collection of the Institute of Zoology, PAS, Warsaw. I. Pseudomyrmicinae, Dorylinae, Ponerinae. Annales Zoologici, 44, 75–82.
Radchenko, A. & Elmes, G.W. (2001) First record of the genus Myrmica (Hymenoptera: Formicidae) from northern Vietnam, with a description of two new species. Annales Zoologici (Warszawa), 51, 221–225.
Radchenko, A., Elmes, G.W. & Bui, T.V. (2006) Ants of the genus Myrmica (Hymenoptera: Formicidae) from Vietnam, with a description of a new species. Myrmecologische Nachrichten, 8, 35–44.
Rigato, F. (1994a) Revision of the myrmicine ant genus Lophomyrmex, with a review of its taxonomic position (Hymenoptera: Formicidae). Systematic Entomology, 19, 47–60.
Rigato, F. (1994b) Dacatria templaris gen.n., sp.n. A new myrmicine ant from the Republic of Korea. Deutsche Entomologische Zeitschrift (N.F.), 1, 155–162.
Rigato, F. & Bolton, B. (2001) The ant genus Liomyrme: a review (Hymenoptera: Formicidae). Bollettino della Società Entomologica Italiana, 133, 247–256.
Roger, J. (1862) Einige neue exotische Ameisen-Gattungen und Arten. Berliner Entomologische Zeitschrift, 6, 233–254.
Roger, J. (1863) Verzeichniss der Formiciden-Gattungen und Arten. Berliner Entomologische Zeitschrift 7 (Beilage), 1–65.
Roncin, E. (2002) Two new Tetramorium species (Hymenoptera: Formicidae) from Vietnam with a discussion of the mixtum, tonganum, and scabrosum groups. Sociobiology, 40, 281–292.
Santschi, F. (1910) Deux nouvelles fourmis du Tonkin. Naturaliste (Revue Illustrée des Sciences Naturelles), (2) 32, 283–284.
Santschi, F. (1920a) Cinq nouvelles notes sur les fourmis. Bulletin de la Société Vaudoise des Sciences Naturelles, 53, 163–186.
Santschi, F. (1920b) Fourmis d’Indo-Chine. Annales de la Société Entomologique de Belgique, 60, 158–176.
Santschi, F. (1924) Fourmis d’Indochine. Opuscules de l’Institut Scientifique de l’Indochine (Saigon), 3, 95–117 [= Faune Entomologique de l’Indochine Francaise, 8, 95–117].
Saunders, W.W. (1842) Descriptions of two hymenopterous insects from northern India. Transactions of the Entomological Society of London, 3 (1841), 57–58.
Schödl, S. (1998) Taxonomic revision of Oriental Meranoplus F. Smith, 1853 (Insecta: Hymenoptera: Formicidae: Myrmicinae). Annalen des Naturhistorischen Museums in Wien, 100B, 361–394.
Seifert, B. (2003) The ant genus Cardiocondyla – a taxonomic revision of the C. elegans, C. bulgarica, C. batesii, C. nuda, C. shuckardi, C. stambuloffii, C. wroughtonii, C. emeryi and minutior species groups. Annalen des Naturhistorischen Museums in Wien, 104B, 203–338.
Shattuck, S.O. (1999) Australian ants. Their biology and identification. Monographs on Invertebrate Taxonomy, 3, 226pp.
Shattuck, S.O. & Barnett, N.J. (2007) Revision of the ant genus Mayriella. Memoirs of the American Entomological Institute, 80, 437–458.
Smith, F. (1852) Descriptions of some hymenopterous insects captured in India, with notes on their economy, by Ezra T. Downes, Esq., who presented them to the Honourable the East India Company. Annals and Magazine of Natural History, (2) 9, 44–50.
Smith, F. (1853) Monograph of the genus Cryptocerus, belonging to the group Cryptoceridae – family Myrmicidae – division Hymenoptera Heterogyna. Transactions of the Entomological Society of London, (2) 2 (1854), 213–228.
Smith, F. (1863) Descriptions of new genera and species of exotic Hymenoptera. Journal of Entomology, 1, 65–84.
Sterling, E.J., Hurley, M.M. & Le, D.M. (2006) Vietnam: A Natural History. Yale University Press, New Haven and London. 423pp. ISBN: 978-0-300-10608-4.
Taber, S.W. (2000) Fire Ants. Texas A&M University Press. 308 pp. ISBN: 0-89096-945-0.
Taylor, R.W. (2009) Ants of the genus Lordomyrma Emery (1) Generic synonymy, composition and distribution, with notes on Ancyridris Wheeler and Cyphoidris Weber (Hymenoptera: Formicidae: Myrmicinae). Zootaxa, 1979, 16–28.
Terayama, M. (1999) Family Formicidae, pp. 138–317, in Identification guide to the aculeata of the Nansei Islands, Japan, Eds. Sk. Yamane, S. Ikudome and M. Terayama. 831pp. Hokkaido University Press, Sapporo.
Terayama, M. (2009) A synopsis of the family Formicidae of Taiwan (Insecta, Hymenoptera). Bulletin of Kanto Gakuen University, 17, 81–266.
Terayama, M. & Kihara, A. (1994) Distribution Maps of Japanese Ants. The Myrmecological Society of Japan. 63 pp.
Terayama, M. & Yamane, Sk. (2000) Lasiomyrma, a new stenammine ant genus from Southeast Asia (Hymenoptera: Formicidae). Entomological Science, 3, 523–527.
The Myrmecological Society of Japan (ed.) (1989) A Guide for the Identification of Japanese Ants (I) — Ponerinae, Cerapachyinae, Pseudomyrmecinae, Dorylinae and Leptanillinae (Hymenoptera: Formicidae). The Myrmecological Society of Japan. 42 pp.
The Myrmecological Society of Japan (ed.) (1991) A Guide for the Identification of Japanese Ants (II) — Dolichoderinae and Formicinae (Hymenoptera: Formicidae). The Myrmecological Society of Japan. 56 pp.
The Myrmecological Society of Japan (ed.) (1992) A Guide for the Identification of Japanese Ants (III) — Myrmicinae and supplement to Leptanillinae (Hymenoptera: Formicidae). The Myrmecological Society of Japan. 94 pp.
Wang, M. (2003) A monographic revision of the ant genus Pristomyrmex. Bulletin of the Museum of Comparative Zoology, 157, 383–542.
Ward, P.S. (2001) Taxonomy, phylogeny and biogeography of the ant genus Tetraponera (Hymenoptera: Formicidae) in the Oriental and Australian regions. Invertebrate Taxonomy, 15, 589–665.
Westwood, J.O. (1839) An introduction to the modern classification of insects; founded on the natural habits and corresponding organisation of the different families 2 (part 11), 193–224. London.
Wheeler, W.M. (1919) The ants of Borneo. Bulletin of the Museum of Comparative Zoology at Harvard College, 63, 43–147.
Wheeler, W.M. (1922) Ants of the American Museum Congo Expedition. A contribution to the myrmecology of Africa. VII. Keys to the genera and subgenera of ants. Bulletin of the American Museum of Natural History, 45, 631–710.
Wheeler, W.M. (1924) Ants of Krakatau and other islands in the Sunda strait. Treubia, 5, 239–258.
Wheeler, W.M. (1927) Ants collected by Professor F. Silvestri in Indochina. Bolletino del laboratorio di Zoologia generale e agraria della R. Scuola Superiore di Agricoltura di Portici, 20, 83–106.
Xu, Z.H. (2000) Two new genera of ant subfamilies Dorylinae and Ponerinae (Hymenoptera: Formicidae) from Ynnan, China. Zoological Research, 21, 297–302.
Yamane, Sk. (2000) Preliminary survey on the distribution pattern of Southeast Asian Pheidologeton species (Hymenoptera: Formicidae). In Maryati M., Fellowes, J.R. & Yamane, Sk. (eds): Proceedings of the 2nd AneT Workshop and Seminar. 118 pp. Universiti Malaysia Sabah, Kota Kinabalu, Malaysia.
Yamane, Sk., Bui, T.V., Ogata, K., Ôkido, H. & Eguchi, K. (2003) Ant fauna of Cuc Phuong National Park, North Vietnam (Hymenoptera: Formicidae). Bulletin of the Institute of Tropical Agriculture, Kyushu University, 25 (2002), 51–62.
Yamane, Sk., Bui, T.V. & Eguchi, K. (2008) Opamyrma hungvuong, a new genus and species of ant related to Apomyrma (Hymenoptera: Formicidae: Amblyoponinae). Zootaxa, 1767, 55–63.