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Type: Articles
Published: 2005-08-08
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A supplement to the checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae) with notes on their tribal classification

Research Division, Canadian Museum of Nature, P.O. Box 3443, Station D, Ottawa, ON, K1P 6P4, CANADA
Department of Entomology, National Museum of Natural History, Smithsonian Institution, Washington, DC and Department of Recent Invertebrates, Virginia Museum of Natural History, Martinsville, VA, c/o1600 Nottoway Ave., Richmond, VA 23227, USA.
Coleoptera Scarabaeidae Melolonthinae chafers New World

Abstract

The checklist of the New World Melolonthinae published in 2003 by Evans is updated to 30 June 2005. Corrections and omissions to the previous checklist are also noted and necessary taxonomic changes are made. Melolontha elongata Fabricius, 1792 is designated as the type species of Philochloenia Dejean, 1833, syn. nov. thereby placing this genus as a junior synonym of Dichelonyx Harris, 1827. Phyllophaga guatemalica (Moser, 1918), syn. nov. and Phyllophaga longiclava (Moser, 1918), syn. nov. are placed in synonymy with Phyllophaga ravida (Blanchard, 1851). The Australian genus Deuterocaulobius Dalla Torre, 1912, stat. nov. is brought out of synonymy and considered the valid name for “Phyllochlaenia Blanchard, 1846.” Warwickia, nom. nov. is proposed as a replacement name for the junior homonym Benedictia Sanderson, 1939 (non Dybowski, 1875) and consequently, Warwickia pilosa (Sanderson, 1939), comb. nov. is a new combination for the single species in this genus. A brief overview of the tribal classification of Melolonthinae is presented.

References

  1. Ahrens, D. (2004) Revisional notes on Sericini: The taxonomic status of the “Neotropical” genus Rhynchosymmela Frey, 1974. Beiträge zur Entomologie, 53, 31–35.

    Alcázar-Ruiz, J.A., Morón-Rios, A. & Morón, M.A. (2003) Fauna de Coleoptera Melolonthidae de Villa Las Rosas, Chiapas, México. Acta Zoologica Mexicana (n.s.), 88, 59–86.

    Aragón, A. & Morón, M.A. (2003) Two new species of Phyllophaga Harris (Coleoptera: Scarabaeidae, Melolonthinae) from south central Mexico. Proceedings of the Entomological Society of Washington, 105(3), 559–567.

    Arce-Pérez, R. & Morón, M.A. (2005) New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactylini) from Honduras and Nicaragua. Zootaxa, 1012, 23–37.

    Arnett, R.H. (1971) The beetles of the United States. A manual for identification. The American Entomological Institute, Ann Arbor, USA, 1112 pp.

    Arrow, G.J. (1912) Scarabaeidae; Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Coleopterorum Catalogus, 43, 1–66.

    Baraud, J. (1985) Coléoptères Scarabaeoidea. Faune du Nord de l’Afrique du Maroc au Sinaï. Ëditions Lechevalier, Paris, 651 pp.

    Baraud, J. & Branco, T. (1991) Revision des Chasmatopterus Latreille, 1825 (Coleoptera: Melolonthidae). Coleopterological Monographs, 1 [1990], 1–55.

    Berthold, A.A. (1827) Latreille’s Naturliche Familien des Thierreichs. Aus dem Französischen. Mit Anmerkungen und Zusätzen. Landes Industrie Comptoirs, Weimar, 606 pp. [Not seen].

    Bezdek, A. (2004) Catalogue of Diplotaxini (Coleoptera: Scarabaeidae: Melolonthinae) of the Old World. Zootaxa, 463, 1–90.

    Blackwelder, R.E. (1939) Fourth supplement, 1933 to 1938 (inclusive) to the Leng catalogue of Coleoptera of America, north of Mexico. John D. Sherman, Mt. Vernon, USA, 146 pp.

    Blackwelder, R.E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Bulletin of the United States National Museum, 185, 189–341.

    Blackwelder, R.E. & Arnett, R.H. (1974) Checklist of the beetles of Canada, United States, Mexico, Central America and the West Indies. North American Beetle Fauna Project (red version). Volume 1, part 3. The scarab beetles, ant-loving beetles, clown beetles and related groups. The Biological Research Institute of America, Latham, USA, 120 pp.

    Blackwelder, R.E. & Blackwelder, R.M. (1948) Fifth supplement, 1939 to 1947 (inclusive) to the Leng catalogue of Coleoptera of America, north of Mexico. John D. Sherman, Mt. Vernon, USA, 87 pp.

    Blanchard, C.É. (1842) Insectes de l’Amérique Méridionale. In: d’Orbigny, A., Blanchard, C.É., & Brullé, A. (Ed), Voyage dans l’Amérique Méridionale, volume 6, part 2. P. Bertrand, Paris, plates 11–12.

    Blanchard, C.É. (1845) Histoire Naturelle des Insectes, Leurs Moeurs, Leurs Métamorphoses et Leur Classification ou Traité Élémentaire d’Entomologie, volume 1. Librairie F. Savy, Paris, 398 pp.

    Blanchard, C.É. (1846) Insectes coléoptères. In: Hombron, J. B. & Jacquinot, H. (Ed), Atlas d’histoire Naturelle Zoologie. Voyage au Pole Sud et dans l’Océanie sur les Corvettes l’Astrolabe et la Zélée; Exécuté par Ordre du Roi pendant les Années 1837–1838–1839–1840. Gide et J. Baudry, Paris, plate 8.

    Blanchard, C.É. (1847) Insectes de l’Amérique Méridionale. In: d’Orbigny, A., Blanchard, C.É., & Brullé, A. (Ed), Voyage dans l’Amérique Méridionale, volume 6, part 2. P. Bertrand, Paris, 448 pp.

    Blanchard, C.É. (1850) Ordre des Coléoptéres. In: Milne-Edwards, H., Blanchard, C.É., & Lucas, H. (Ed) Museum d’Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des insectes, volume 1, part 1. Gide & Baudry, Paris, pp. 1–128.

    Blanchard, C.É. (1851) Ordre des Coléoptéres. In: Milne-Edwards, H., Blanchard, C.É., & Lucas, H. (Ed) Museum d’Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des insectes, volume 1, part 2. Gide & Baudry, Paris, pp. 129–240.

    Blanchard, C.É. (1854) Description des Insectes. In: Hombron, J. B. & Jacquinot, H. (ed), Voyage au Pole Sud et dans l’Océanie sur les Corvettes l’Astrolabe et la Zélée; Exécuté par Ordre du Roi pendant les Années 1837–1838–1839–1840. Zoologie, volume 4. Gide et J. Baudry, Paris, 422 pp.

    Branco, T. (2005). Description of a new species of Chasmatopterus Dejean, 1821, from Portugal, with a note on the subfamily Chasmatopterinae (Coleoptera: Melolonthidae). Elytron (2003–2004), 17–18, 27–36. [Dated 2003–2004]

    Brenske, E. (1900) Coléoptères. Scarabaeidae. In: Diagnoses d’insectes recuellis par l’expédition anatarctiques belge. Annales de la Société Entomologique de Belgique, 44, 109–110.

    Britton, E.B. (1957) A revision of the Australian chafers (Coleoptera: Scarabaeidae, Melolonthinae). Volume I. British Museum (Natural History), London, 185 pp.

    Britton, E.B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2 Tribe Melolonthini. Australian Journal of Zoology Supplementary Series, 60, 1–150.

    Britton, E.B. (1988) Synopsis of the genera of Australian Heteronycini (Coleoptera: Scarabaeidae: Melolonthinae). Journal of the Australian Entomological Society, 27, 27–36.

    Britton, E.B. (1990) A synopsis of the Australian genera of Liparetrini (Coleoptera: Scarabaeidae: Melolonthinae). Invertebrate Taxonomy, 4, 159–195.

    Britton, E.B. (2000) A review of Heteronyx Guérin-Méneville (Coleoptera: Scarabaeidae; Melolonthinae). Invertebrate Taxonomy, 14, 465–589.

    Burmeister, H. (1855) Handbuch der Entomologie. Fierter Band. Besondere Entomologie. Fortfessung. Zweite Abtheilung. Coleoptera Lamellicornia Phyllophaga chaenochela. Enslin, Berlin, 569 pp.

    Casey, T.L. (1914) A review of the genus Thyce and of the North American species of Polyphylla. Memoirs on the Coleoptera, 5, 306–453.

    Chalumeau, F. & Gruner, L. (1976) Scarabaeoidea des Antilles Françaises 2e partie: Melolonthinae et Rutelinae. Annales de la Société Entomologique France, (N.S.), 12(10), 83–112.

    von Dalla Torre, K.W. (1912) Scarabaeidae; Melolonthinae I. Coleopterorum Catalogus, 20, (45, 47, 49), 1–450.

    von Dalla Torre, K.W. (1913) Scarabaeidae; Melolonthinae IV. Coleopterorum Catalogus, 20, (50), 291–450.

    Dejean, P.F.M.A. (1833) Catalogue des Coléoptères de la collection de M. le Comte Dejean, Fascicles 1–2. Méquignon-Marvis Père et Fils, Paris, 176 pp.

    Delgado, L. (2001) A new Mexican species of Diplotaxis Kirby (Coleoptera, Melolonthidae, Melolonthinae) of the puberula group. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 74(1–2), 139–142.

    Delgado, L. & Navarrete-Heredia, J.L. (2004) Ceraspis jaliscoensis, a new species from México (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 787, 1–7.

    Erichson, W.F. (1847) Naturgeschichte der Insecten Deutschlands. Erste Abtheilung. Coleoptera. Dritter Band. Nicolaischen Buchhandlung, Berlin, pp. 481–800.

    Evans, A.V. (1988) Systematics of the Subsaharan Pachydemini (Coleoptera: Melolonthidae: Melonthinae). University of Pretoria, South Africa. [Unpublished PhD dissertation].

    Evans, A.V. (2002) Melolonthinae. In: Arnett, R.H., Thomas, M.C., Skelley, P.E., & Frank, J.H. (Ed), American Beetles. Volume 2. Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press, Boca Raton, USA, pp. 51–60.

    Evans, A.V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211, 1–458.

    Fairmaire, M.L. & Germain, P. (1863) Revision des Coléoptères du Chili. Suite (1). Fam. Scarabaeidae. Genres Maypa et Listronyx. Annales de la Société Entomologique de France, (4)2, 723–746.

    Frey, G. (1967) Die Gattung Plectris (Philochlaenia) (Coleoptera, Melolonthinae). Entomologischen Arbeiten aus dem Museum G. Frey, 18, 1–136.

    Frey, G. (1975) Eine neue südamerikanische Gattung und Tribus der Melolonthiden. Entomologischen Arbeiten aus dem Museum G. Frey, 26, 84–86.

    Gutiérrez, R. (1952) Notas sobre Scarabaeidae neotrópicos (III). Revista Chilena de Entomologia, 2, 207–227.

    Hardy, A.R. (1978) Placement of the genus Benedictia Sanderson (Coleoptera: Scarabaeidae). The Coleopterists Bulletin, 32, 67–70.

    von Harold, E. (1869) Scarabaeidae. In: Gemminger, M. & Harold, E. (Ed), Catalogus Coleopterorum. bucusque descriptorum, synonymicus et systematics, autotribus. Tome IV. Scarabaeidae. E.H. Gummi, Münich, pp. 976–1346.

    Hatch, M.H. (1971) The beetles of the Pacific northwest. Part V. Rhipiceroidea, Sternoxi, Phytophaga, Rhynchophora, and Lamellicornia. University of Washington Publications in Biology, 16, 1–662.

    Hope, F.W. (1837) The Coleopterist’s Manual containing the lamellicorn insects of Linnaeus and Fabricius. Henry G. Bohn, London, 121 pp.

    Horn, G.H. (1887) Revision of the species of Lachnosterna of America North of Mexico. Transactions of the American Entomological Society, 14, 209–296.

    Houston, W.W.K. & Weir, T.A. (1992) Melolonthinae. In: Houston, W.W.K. (Ed), Zoological catalogue of Australia. Coleoptera: Scarabaeoidea. Australian Government Printing Service, Canberra, pp. 174–358.

    Howden, H.F. (1954) A review of the genus Podolasia Harold (Coleoptera: Scarabaeidae). American Museum Novitates, 1661, 1–11.

    Howden, H.F. (1958) Species of Acoma Casey having a three-segmented antennal club (Coleoptera: Scarabaeidae). The Canadian Entomologist, 90(7), 377–401.

    Howden, H.F. (1968) Generic relationships of Thyce, Plectrodes, Dinacoma, and Hypotrichia, with a description of a new genus and species from eastern Texas (Coleoptera: Scarabaeidae: Melolonthini). The Canadian Entomologist, 100(5), 542–548.

    Howden, H.F. (1997) Podolasiini Howden, new tribe, and a revision of the included genera, Podolasia Harold and Podostena Howden, new genus (Coleoptera: Scarabaeidae: Melolonthinae). The Coleopterists Bulletin, 51(3), 223–255.

    Howden, H.F. & Hardy, A.R. (1971) Generic placement and adult behavior of the genus Leptohoplia Saylor. Proceedings of the Entomological Society of Washington, 73, 337–341.

    Iablokoff-Khnzorian, S.M (1977) Über die Phylogenie der Lamellicornia (Insecta, Coleoptera). Entomologische Abhandlungen, 41(5), 135–200.

    Janssens, A. (1949) Contribution a l'etude des Coléoptères Lamellicornes. XIII – Table synoptique et essai de classification pratique des Coléoptères Scarabaeidae. Bulletin de l’Institute royal des Sciences naturelles de Belgique, 25, 1–30.

    Katovich, K. (2002) A phylogenetic world revision of the Macrodactylini (Coleoptera: Scarabaeidae: Melolonthinae). University of Wisconsin-Madison. 754 pp. [unpublished PhD dissertation]

    Keith, D. (2004) Note homonymique sur Liogenys excisus Moser, 1919 (Col. Melolonthidae). Bulletin de la Société entomologique de France, 109(2), 195.

    Keith, D. & Lacroix M. (2003) Un nouveau Tanyproctus Ménétriés, 1832 de Chine et remarques sur les genres Pachnessa Brenske, 1894 et Peritryssus Reitter, 1918 (Coleoptera, Scarabaeoidea, Melolonthidae). Coléoptères, 9(4), 37–50.

    Kohlmann, B. & Morón M.A. (2003) Análisis histórico de la clasificación de los Coeloptera, Scarabaeoidea, o Lamellicornia. Acta Zoológica Mexicana (n.s.), 90, 175–280.

    Lacordaire, J.T. (1856) Histoire naturelle des insectes. Genera des Coléoptères, or exposé méthodique et criticque de tous genres proposés jusquici dans cet odre d’insectes. Tome Troiséme. Contenant les familles des Pecticornes et Lamellicornes. Librarie Encyclopedique de Roret, Paris, 594 pp.

    Lacroix, M. (1993) Fauna de Madagascar. Insectes Coléoptères Melolonthidae (2 partie). Muséum National d’Histoire Naturelle, Paris, pp. 303–875.

    Lacroix, M. (2001) Pachydeminae de l’Est africain (Kenya et Tanzanie) (Coleoptera, Melolonthidae). Coléoptères, 7(13), 173–225.

    LeConte, J.L. (1856) Synopsis of the Melolonthidae of the United States. Journal of the Academy of Natural Sciences of Philadelphia, (2)3, 225–288.

    LeConte, J.L. (1861) Classification of the Coleoptera of North America. Part I. Smithsonian Miscellaneous Collections, 136, 1–208.

    LeConte, J. L. & Horn, G.H. (1883) Classification of the Coleoptera of North America. Prepared for the Smithsonian Institution. Smithsonian Miscellaneous Collections, 26(4), i–xxxviii + 1–567.

    Leng, C.W. (1920) Catalogue of the Coleoptera of America, north of Mexico. John D. Sherman, Jr., Mount Vernon, USA, 470 pp.

    Leng, C.W. & Mutchler, A.J (1927) Supplement 1919 to 1924 (inclusive) to catalogue of the Coleoptera of America, north of Mexico. John D. Sherman, Jr., Mount Vernon, USA, 52 pp.

    Leng, C.W. & Mutchler, A.J (1933) Second and third supplements, 1925 to 1932 (inclusive) to catalogue of the Coleoptera of America, north of Mexico. John D. Sherman, Jr., Mt. Vernon, USA, 112 pp.

    Machatschke, J.W. (1959) Phylogenetische Untersuchungen über die Sericini (sensu Dalla Torre 1912) (Coleoptera: Lamellicornia, Melolonthidae). Beiträge zur Entomologie, 9(7/8), 730–746.

    Martínez, A. (1967) El género Athlia Erichson (Col. Scarabaeidae, Sericinae). Entomologischen Arbeiten aus dem Museum G. Frey, 18, 327–372.

    Martínez, A. (1971) Un nueva especie de Sericoides Guérin de la Argentina (Col. Scarab. Sericinae, Liparetrini). Revista Sociedad Entomologica Argentina, 33, 115–120.

    Martínez, A. (1972) Algunas consideraciones sobre los generos Apterodema Fairmaire y Sericoides Guerin, con descripcion de dos nuevas especies (Scarab. Sericinae, Liparetrini). Comunicaciones del Museo Argentino de Ciencias Naturales Bernardino Rivadavia, 1(3), 37–62.

    Martínez, A. (1975) Contribución al conocimiento de los Pachydemini neotropicales (Col. Scarabaeidae, Melolonthinae). Entomologischen Arbeiten aus dem Museum G. Frey, 26, 227–251.

    Martínez, A. (1982) Un nuevo genero de Pachydemini Argentino (Col. Scarab. Melolonthinae). Revista Sociedad Entomologica Argentina, 41(1–4), 89–91.

    Morón, M.A. (2003a) Revision of the Phyllophaga s.s. schizorhina species group (Coleoptera: Melolonthidae: Melolonthinae). Canadian Entomologist, 135(2), 213–302.

    Morón, M.A. (2003b) Diversidad, distribución e importancia de las especies de Phyllophaga Harris en México (Coleoptera: Melolonthidae). In: Aragón, G.A., Morón M.A., & Marín A. (Ed), Estudios Sobre Coleópteros del Suelo en América. Benemérita Universidad Autónoma de Puebla, México, pp 1–27.

    Morón, M.A. (2004) Revision of the cavata group of Phyllophaga (Listrochelus) Blanchard (Coleoptera: Melolonthidae: Melolonthinae). Annals of the Entomological Society of America, 97(1), 77–96.

    Morón, M.A., Ratcliffe, B.C., & Deloya, C. (1997) Atlas de los escarabajos de Mexico (Coleoptera: Lamellicornia). Vol.1. Familia Melolonthidae (subfamilias Rutelinae, Dynastinae, Cetoniinae, Trichiinae, Valginae, Melolonthinae). Sociedad Mexicana de Entomologica, Mexico, 280 pp.

    Morón, M. A. & Robbins, P.S. (2004) Especie nueva de Phyllophaga (Chlaenobia) (Coleoptera: Melolonthidae: Melolonthinae) de Honduras, América Central. Anales del Instituto de Biologia, Universidad Nacional Autónoma de México (serie Zoologia), 75(2), 331–339.

    Moser, J. (1918) Neue Arten der Gattungen Lachnosterna Hope und Phytalus Er. (Col.). Stettiner Entomologische Zeitung, 79, 19–74.

    Muslera Ramos, L. & Fernandez Garcia, I. (1998). Estado actual de conocimiento del genero Phyllophaga (Coleoptera: Scarabaeidae) en Cuba. Poeyana, 461, 1–22

    Philippi, F. (1887) Catálogo de los Coléopteros de Chile. Anales de la Universidad Republica de Chile, 71, 619–806.

    Reitter, E. (1902) Bestimmungs-Tabelle der Melolonthidae aus der europäischen Fauna und den angrenzenden Ländern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brünn, 40 [1901], 93–303.

    Reitter, E. (1918) Eine neue Lamellicornien-Gattung aus Sizilien. Wiener Entomologische Zeitung, 37, 77–78.

    Restrepo-Giraldo, H., Morón, M.A., Vallejo, F., Pardo-Locarno, L.C., & López-Aviola, A. (2003) Catálogo de Coleoptera Melolonthidae (Scarabaeidae Pleurosticti) de Colombia. Folia Entomologica Mexicana, 42(2), 47–54.

    Riley, E.G. & Wolfe, C.S. (2003) An annotated checklist of the Scarabaeoidea of Texas (Coleoptera). Southwestern Entomologist Supplement, 26, 1–37.

    Ritcher, P.O. (1966) White grubs and their allies. A study of North America scarabaeoid larvae. Oregon State University Press, Corvallis, USA, 219 pp.

    Ritcher, P.O. (1969a) Spiracles of adult Scarabaeoidea (Coleoptera) and their phylogenetic significance. I. The abdominal spiracles. Annals of the Entomological Society of America, 62(4), 869–880.

    Ritcher, P.O. (1969b) Spiracles of adult Scarabaeoidea (Coleoptera) and their phylogenetic significance. II. Thoracic spiracles and adjacent sclerites. Annals of the Entomological Society of America, 62(6), 1388–1398.

    Sanderson, M.W. (1939) A new genus of Scarabaeidae with descriptions and notes on Phyllophaga. Journal of the Kansas Entomological Society, 12(1), 1–15.

    Sanderson, M.W. (1942) Descriptions and records of distribution of Phyllophaga (Coleoptera; Scarabaeidae). Journal of the Kansas Entomological Society, 15(2), 49–55.

    Sanderson, M.W. (1958) Faunal affinities of Arizona Phyllophaga, with notes and descriptions of new species. Journal of the Kansas Entomological Society, 13, 158–173.

    Saylor, L.W. (1935) A new genus and two new species of Coleoptera from California (Scarabaeidae). Pan-Pacific Entomologist, 11, 132–134

    Saylor, L.W. (1937) The beetles of the subfamily Chasmatopterinae in the New World. Journal of the Washington Academy of Sciences, 27(12), 531–535.

    Saylor, L.W. (1938) Revision of the subfamily Oncerinae with description of a new genus (Coleoptera: Scarabaeidae). Proceedings of the Entomological Society of Washington, 40, 99–103.

    Skelley, P.E. (2005) Review of the tribe Melolonthini in the southeastern United States (Coleoptera: Scarabaeidae: Melolonthinae). Insecta Mundi, 17 [2003], 129–156. [mailing date January 2005, P. E. Skelley, personal communication].

    Smith, A.B.T. (2003) Checklist of the Scarabaeoidea of the Nearctic realm (includes Canada, the continental United States, and the following states of northern Mexico: Baja California, Baja California Sur, Chihuahua, Coahuila de Zaragoza, Durango, Neuvo Leon, Sinaloa, Sonora, Tamaulipas, and Zacatecas). Lincoln, Nebraska, USA, 74 pp. Available from: http://www.museum.unl.edu/research/entomology/nearctic.htm (accessed 28 June 2005).

    Woodruff, R.E. (2005) [Species descriptions] In: Woodruff, R.E. & Sanderson, M.W. (authors), Revision of the Phyllophaga of Hispaniola (Coleoptera: Scarabaeidae: Melolonthinae). Insecta Mundi, 18(1–4), 1–154. [mailing date January 2005, P. E. Skelley, personal communication].

    Woodruff, R.E. & Sanderson, M.W. (2005) Revision of the Phyllophaga of Hispaniola (Coleoptera: Scarabaeidae: Melolonthinae). Insecta Mundi, 18(1–4) [2004], 1–154. [mailing date January 2005, P. E. Skelley, personal communication].