Skip to main content Skip to main navigation menu Skip to site footer
Type: Articles
Published: 2011-12-08
Page range: 1–62
Abstract views: 313
PDF downloaded: 4

Systematics, morphology, and ecology of pigeons and doves (Aves: Columbidae) of the Mascarene Islands, with three new species

Bird Group, The Department of Zoology, Natural History Museum, Akeman St, Tring, Herts HP23 6AP
Aves Mascarene pigeons and doves Nesoenas Alectroenas Columba extinction affinities ecology

Abstract

The original diversity of the pigeons and doves (Columbidae: Nesoenas, Columba, Alectroenas) of the Mascarene Islands (Mauritius, Réunion, Rodrigues) has been poorly understood. Only two of perhaps as many as ten species are known from skin specimens, whereas the rest are known from old accounts and subfossil remains only. Most accounts, however, do not distinguish between species, so accurate identification is difficult to determine. The introduction of non-native pigeons has further exacerbated the problem and has led to erroneous interpretation. This paper provides a detailed re-analysis of the Mascarene columbid fauna (excluding the large, terrestrial “didines”, the Dodo Raphus cucullatus and Solitaire Pezophaps solitaria), based partly on newly discovered subfossil remains. Key findings include: a new species of Alectroenas from Rodrigues and new species of Nesoenas and Columba from Mauritius; referral of the problematic species 'Columba' rodericana of Rodrigues to the genus Nesoenas; and documentation of new morphological and historical information concerning the extant Pink Pigeon Nesoenas mayeri and the extinct Mauritius Blue Pigeon Alectroenas nitidissima. The Columbidae comprises the largest avian radiation in the Mascarenes and probably colonised the islands at least four times from Madagascar and SE Asia during low sea level stands.

References

  1. Alby, J. & Serviable, M (Eds.) (1993) Abbé Alexandre-Gui Pingré: Courser Venus, voyage scientifique à Rodrigue en 1761. M. Ste.Clotilde, Réunion, Ars Terres Créoles & Rose-Hill, Mauritius: Editions Ocean Indien, 123pp. [extracts from Pingré 1763, q.v., with a historical introduction].

    Baptista, L.F., Trail, P.W., & Horblit, H.M. (1997) Family Columbidae (Pigeons). In: del Hoyo, T., Elliott, A., & Sargatal, J. (Eds.) (1997). Handbook of the birds of the world volume 4, sandgrouse to cuckoos. Lynx Edicions, Barcelona, pp. 60–243.

    Baptista, L.F., Martinez Gómez, J. E., & Horblit, M. (2009) Darwin’s pigeons and the evolution of the Columbiforms: recapitulation of ancient genes. Acta Zoológica Mexicana new series, 25, 3, 719–741.

    Barnwell, P.J. (1948) Visits and despatches 1598-1948. Mauritius Standard Printing Establishment, Port Louis, 306 pp.

    Barré, N., Barau, A. & Jouanin, C. (1996) Oiseaux de la Réunion. Les Éditions du Pacifique, Paris, 208 pp.

    Baumel, J.J., & Witmer, L.M. (1993) Osteologia. In: Baumel, J.J., King, A.S., Breazile, J.E., Evans, H. E., Vanden Berge, J. C. (Eds.), Handbook of avian anatomy: nomina anatomica avium, second edition. Cambridge, Massachusetts, USA: Publications of the Nuttall Ornithological Club, 23, 45–132.

    Benson, C.W. (1967) The birds of Aldabra and their status. Atoll Research Bulletin, 118, 63–111.

    Benson, C.W. (1970) An introduction of Streptopelia picturata into the Amirantes. Atoll Research Bulletin, 136, 195–196.

    Benson, C.W. (1999) Type specimens of bird skins in the University Museum of Zoology, Cambridge, United Kingdom. British Ornithological Club Occasional Publications 4, 221 pp.

    Benson, C. W. & Penny, M.J. (1971) The land birds of Aldabra. Philosophical Transactions of the Royal Society London B, 260, 417–527.

    Bernardin de St.Pierre, J.H. (1773) Voyage à l'Isle de France, à l'Isle de Bourbon, au Cap de Bonne Espérance; &c. par un officier du Roi. Neuchâtel: Société Typographique [reprinted 2002 Oxford: Signal Books – with new tr. & intro. by Jason Wilson as Journey to Mauritius, 290pp.]

    Boie, S. (1828) Bermerfungen uber mehrere neue Vogelgattungen. Isis. Band XXI, heft III, p.327.

    Bonaparte, C.L. (1854) Comptes rendus hebdomadaires des séances de l'Académie des Sciences, Paris, volume 36. Mallet-Bachelier, Imprimeur-Librairie, Paris, 1277 pp.

    Bonaparte, C.L. (1854) Conspectus genera avium, volume I. E J Brill, Lugduni Batavorum.

    Bonaparte, C.L. (1857) Conspectus genera avium, volume II. E J Brill, Lugduni Batavorum. 252 pp.

    Bonnaterre, A. (1790) Tableau encyclopédique et méthodique, Des trois régnes de la nature. Panckoucke, Paris, 1460 pp.

    Bontekoe, W.Y. (1646) Journal ofte Gedenckwaerdige beschrijvinghe vande Oost-Indische Reyse van Willem Ijbrantz Bontekoe van Hoorn. Jan Jansz Deutel, Hoorn, 60 pp.

    Boucher, A. (1710) Mémoire d'Antoine Boucher sur l'Ile Bourbon en 1710 [Ed. A.Lougnon]. Recueil trimestriel de documents et travaux inédits pour servir à l'histoire des Mascareignes françaises, 5, 279–355 (1941).

    Carié, P. (1916) L'acclimatation à l'île Maurice. Bulletin de la Société Nationale d'Acclimatation de France, Paris, 63: 10–18, 37–46, 72–79, 107–10, 152–59, 191–98, 245–50, 355–63 & 401–4 [also reprinted by the society with new pagination, 62pp.]

    Cauche, F. (1651) Relation du voyage que François Cauche de Rouen à fait à Madagascae, îles adjacentes et coste d'Afrique. Recueilly par le sieur Morisot, avec notes en marge. Augustin Courbé, Paris, 194pp.

    Cheke, A.S (1987) An ecological history of the Mascarene Islands, with special reference to extinctions and introductions of land vertebrates. In: Diamond, A. W. (Ed.), Studies of Mascarene Island birds. Cambridge: Cambridge University Press, pp. 5–89.

    Cheke, A.S. (2005) Naming segregates from the Columba-Streptopelia group of pigeons following DNA studies on phylogeny. Bulletin of the British Ornithologists Club, 125, 4, 293–295.

    Cheke, A.S., & Hume, J.P. (2008) Lost land of the dodo: the ecological history of the Mascarene Islands. A. & C. Black publishers (Poyser imprint), London, 540pp.

    Cossigny [Charpentier de Cossigny], J.F. (1732) Trieze letters de Cossigny à Réaumer [Lougnon 1939-40]. Recueil trimestriel de documents et travaux inédits pour servir à l'histoire des Mascareignes françaises 4, 168–96,205–82, 305–16.

    Cowles, G. (1987) The fossil record. In: Diamond, A. W. (Ed.) Studies of Mascarene Island birds. Cambridge: Cambridge University Press, pp. 90–100.

    Cristidis, L., & Boles, W. (2008) Systematics and taxonomy of Australian birds. CSIRO Publishing, Australian Museum, 277 pp.

    Dapper, O. (1668) (Ed.) van Meurs, J. Naukeurige beschrijvinge der Afrikaensche gewesten van -Egypten, Barbaryen, Libyen, Biledulgerid, Negroslant, Guinea, Ethiopiën, Abyssinie: vertoont in de Benamingen, Grenspalen, Steden, Revieren, Gewassen, Dieren, Zeeden, Talen, Rijkdommen, Godsdiensten en Heerschappyen.

    Desjardins, J. (1832) Troisième rapport annual de la travaux société d’ histoire naturelle de I’Ile Maurice. [MS published pp.53–112 in Ly-Tio-Fane (1972), q.v.]

    Des Murs, O. (1854) Encyclopédie d'histoire naturelle; Oiseaux, volume VI. Marescq and Compagnie, Paris.

    Dickinson, E.C., David, N., Overstreet, L.K., Steinheimer, F.D. & Jansen, J. (2010) Histoire naturelle des pigeons or Les Pigeons: Coenraad Jacob Temminck versus Pauline Knip. Archives of Natural History 37, 2, 203–220.

    Dubois, [Sieur]. (1674) Les voyages faits par le sieur D.B. aux isles Dauphine ou Madagascar et Bourbon ou Macarenne, ès années 1669, 70, 71, et 72. Claude Barbin, Paris, 234 pp.

    Ducros, P. (1725) Lettre du Père Ducros, missionaire de la Compagnie des Indes, à M l'abbé Raguet, directeur de la Compagnie des Indes. Letrres édifiantes, Mémoires des Indes, 18, 1–26 (1728 ed.), 13, 320–340 (1781 Ed.) & 7, 461–473 (1819 Ed.) [parts also in Barnwell (1948) & Lougnon (1970), qq.v.]

    Edmunds, K., Bunbury, N., Sawmy, S., Jones, C.G. & Bell, D.J. (2008) Restoring avian island endemics: use of supplementary food by the endangered Pink Pigeon (Columba mayeri). Emu, 108, 74–80.

    Feare, C. & Gill, E.L. (1995) The turtle doves of Bird Island, Seychelles. Bulletin of the British Ornithologists Club, 115, 4, 206–210.

    Feuilley (1705) Mission à I’lle Bourbon du Sieur Feuilley en 1704. [Ed. A. Lougnon]. Recueil trimestriel de documents et travaux inédits pour servir à l'histoire des Mascareignes françaises, 4, 3–56, 101–67 (1939).

    Frith, C.B. (1979) Feeding ecology of land birds on West Island, Aldabra Atoll, Indian Ocean: a preliminary survey. Philisophical Transactions of the Royal Society of London, B. 286, 195–210.

    Fuller, E. (1987) Extinct birds. Rainbow Penguin, London & New York, 256 pp.

    Fuller, E. (2000) Extinct birds. Oxford University Press, Oxford, 398 pp.

    Gelderland Journal. (1601-2) Algemeen Rijksarchief, Den Haag. Archive compagnieën op Oost-Indië 1594-1603 (1.04.01), inv. Nos. 135, viz. 136, fol. 66r and 65v.

    Gibbs, D., Barnes, E., & Cox, J. (2001) Pigeons and doves: A guide to the pigeons and doves of the world. Pica Press, Sussex, 614 pp.

    Gmelin, J.F. (1788) Systema naturae per regna tria naturae secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. 13th edition 1, part II. Georg Emanuel Beer, Lipsiae [Leipzig].

    Goodwin, D. (1967) Pigeons and doves of the world. Trustees of the British Museum (Natural History): Cornell University Press, Ithaca, New York, 446 pp.

    Goodwin, D. (1983) Pigeons and doves of the world, third edition. Trustees of the British Museum (Natural History): Cornell University Press, Ithaca, New York, 363 pp.

    Grandidier, A. & Grandidier, G. (1905) Collection des ouvrages anciens concernant Madagascar, publiée sous la direction de MM. A. Grandidier, de l’Institut, Charles-Roux, Cl. Delhorbe, H. Froidevaux et G. Grandidier. T. III. Ouvrages ou extraits d’ouvrages anglais, hollandais, portugais, espagnols et allemands relatifs à Madagascar (1640 à 1716), pp. 368-380. Comité de Madagascar, Paris.

    Grant, C. (1801) The History of Mauritius, or the Isle of France, and the neighbouring islands; from their first discovery to the present time. W. Bulmer and Co., London.

    Gray. G.R. (1840) A list of the genera of birds. Richard & John. E. Taylor, London, 80 pp.

    Gray, G.R. (1849) The genera of birds comprising their generic characters, a notice of the habits of each genus, and an extensive list of species referred to their several genera, volume 3. Longman, Brown, Green & Longmans, London, 117 pp.

    Gray, G.R. (1870) Hand-list of genera and species of birds, volume 2, Conirostres, Scansores, Columbae, & Gallinae. Trustees of the British Museum, London.

    Greenway, J.C. (1958) Extinct and vanishing birds. Dover Books, New York, 518 pp.

    Greenway, J.C. (1967) Extinct and vanishing birds of the world. 2nd edition. Dover Publications Inc., New York, 520 pp.

    Griffiths, O.L. & Florens, V.F.B. (2006) A field guide to the non-marine molluscs of the Mascarene islands (Mauritius, Rodrigues and Réunion) and the northern dependencies of Mauritius. Owen Griffiths: Bioculture Press, Mauritius, 185 pp.

    Guérin, R. (1940-53) Faune ornithologique ancienne et actuelle des îles Mascareignes, Seychelles, Comores et des îles avoisinantes. General Printing & Stationery Co., Port Louis, Mauritius, 3 volumes.

    Günther, A., & Newton, E. (1879) The extinct birds of Rodriguez. Philosophical Transactions of the Royal Society of London, 168, 423–437.

    Hachisuka, M. (1953) The dodo and kindred birds, or the extinct birds of the Mascarene Islands. H. F. & G. Witherby, London, 250 pp.

    Herman, J.S., McGowan, R.Y. & Swinney, G.N. (1990) Catalogue of the type specimens of recent vertebrates in the National Museums of Scotland, Series 4. National Museums of Scotland, Edinburgh.

    Het Tweede Boeck. (1601) Journal oft Dagh-register / inhoudende een warachtig verhael ende historische vertellinghe van de Reyse / gedaen door de acht schepen van Amstelredame / gheseylt in den Maert Martij 1598 onder ‘t beleydt van den Admirael Jacob Cornelisz. Neck ende Wybrandt van Warwijck als vice admirael –van hare zeylagie ende gedenwaerdighe zaken ende geschiedenissen haer op de voortz-reys bejeghent. Ghedruct tot Amstelredam by Cornelis Claesz. Opt Water in ‘t Schrijf-boeck. Amsterdam.

    Hoffman, J.C. (1680) (See Grandidier & Grandidier 1905).

    Hume, J.P. (2003) The journal of the flagship Gelderland – dodo and other birds on Mauritius 1601. Archives of Natural History, 30, 1, 13–27.

    Hume, J.P. (2005) Contrasting taphofacies in ocean island settings: the fossil record of Mascarene vertebrates. Monografies de la Societat d’Història Natural de les Balears, 12, 129–144.

    Hume, J.P. (2006) The history of the dodo Raphus cucullatus and the penguin of Mauritius. Historical Biology, 2, 65–89.

    Hume, J.P., Ribes, S., Mourer-Chauviré, C. & Bour, R. (2006) Les animaux disparus. Muséum d'Histoire Naturelle, St Denis, Réunion, 32 pp.

    Hume, J.P. (2007) Reappraisal of the parrots (Aves: Psittacidae) from the Mascarene Islands, with comments on their ecology, morphology and affinities. Zootaxa, 1513, 1–76.

    International Commision on Zoological Nomenclature. (1999) International Code of Zoological Nomenclature 4th ed.. Natural History Museum, London, 306 pp.

    Jobling, J.A. (1991) A dictionary of scientific names. Oxford University Press, Oxford, 272 pp.

    Jobling, J.A. (2009) Helm dictionary of scientific bird names. A & C. Black Publishers Ltd, London, 432 pp.

    Johnson, K.P. & Clayton, D.H. (2000) Nuclear and mitochondrial genes contain similar phylogenetic signal for pigeons and doves (Aves: Columbiformes). Molecular Phylogenetics and Evolution 14, 1, 141 – 151.

    Johnson, K.P., Kort, S.de., Dinwoodey, K., Mateman, A.C., Cate, C.t., Lessells, C.M., & Clayton, D.H. (2001) A molecular phylogeny of the dove genera Streptopelia and Columba. Auk, 118, 4, 874–887.

    Jones, C. (1987) The larger land-birds of Mauritius. In: Diamond, A. W (Ed.), Studies of Mascarene Island birds. Cambridge: Cambridge University Press, pp. 208–300.

    Jouanin, C. (1962) Inventaire des oiseaux éteints ou en voie d’extinction conservés au Muséum de Paris. Terre et Vie, 109, 257–301.

    Knip [de Courcelles], P., & Temminck, J.C. 1809–1811. Les pigeons par Madame Knip, née Pauline de Courcelles ; texte par J. C. Temminck. Bellizard, Dufour et Cie, Paris, 1 volume.

    Knip [de Courcelles], P. (1838–43) Les pigeons. Volume 2. Paris.

    Latham, J. (1783) General synopsis of birds, volume 2, Leigh and Sotheby, London.

    Latham, J. (1790) Indicis ornithologici sive systematis ornithologiae. Sotherby, Londres, 920 pp.

    Leguat, F. (1708) Voyage et avantures de François Leguat et de ses compagnons en deux isles désertes des Indes Orientales. 2 volumes, J.L.de Lorme, Amsterdam.

    Levaillant, F. (1808) Histoire naturelle des oiseaux d' Afrique. Volume 6. Delachaussée, Paris.

    Linnaeus, C (Linné, Karl von). (1758) Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, edition 12, volume 1. Holmiæ, Salvius.

    Louette, M. (1988) Les oiseaux des Comores. Annales du Musée Royal de l'Afrique Centrale (Zool.), 255, 1–192.

    Lougnon, A. 1970. Sous la signe de la tortue. Voyages anciens à l'Ile Bourbon (1611-1725), 3rd edition. Saint-Denis, Réunion, [author], 284 pp.

    Ly-Tio-Fane, M. (Ed.). (1972) Société d'histoire naturelle de l'Ile Maurice. Rapports annuels I-IV, 1830-1834. Royal Society of Arts & Sciences of Mauritius, Port Louis, Mauritius, 200 pp.

    Marchel. (1843) [Prévost, in Knip, Les Pigeons, ed. 2, 2 [1838-1843], 1843: 113, pl. 60].

    McKelvey, S.D. (1976) A preliminary study of the Mauritian Pink Pigeon. Mauritius Institute Bulletin, 3, 2, 145–175.

    McKelvey, S.D. (1977) Observations on the Mauritian Pink Pigeon. Birds International, 2, 36–38.

    Meinertzhagen, R. (1912) On the birds of Mauritius. Ibis, 9, 6, 82–108.

    Melet, J de. (c1672) Relation de mon voyage aux Indes Orientales par mer... [Publ. 1999, Ed.A.Sauvaget (with intro., pp.95-102). Études Océan Indien, 25/26, 103–289].

    Milbert, J.G. (1812) Voyage pittoresque à l'Ile de France, au Cap de Bonne Espérance et à l'Ile de Ténériffe. 2 volumes + atlas. A.Nepveu, Paris.

    Milne-Edwards, A. (1874 [1873]) Recherches sur la faune ancienne des Iles Mascareignes. Annales des Sciences Naturelles – Zoologie et Paléontologie, (5) 19, Art. 3.

    Milne-Edwards, A. & Oustalet, E. (1893) Notice sur quelques espéces d’oiseaux actuellement éteintes qui se trouvent représentées dans les collections du Muséum d’Histoire Naturelle. In: Centenaire de la fondation du Muséum d’Histoire Naturelle. Muséum d’Histoire Naturelle, Paris, 190–252.

    Moree, P. J. (1998) A concise history of Dutch Mauritius, 1598–1710. Kegan Paul International, London and New York, 127 pp.

    Moree, P.J. 2001. Dodo's en galjoenen. Die reis van het schip Gelderland naar Oost-Indië, 1601–1603. Walburg Pers, Zutphen, Netherlands, 348 pp.

    Mourer-Chauviré, C., Bour, R., Ribes, S. & Moutou, F. (1999) The avifauna of Réunion Island (Mascarene Islands) at the time of the arrival of the first Europeans. In: Olson, S. (Ed.), Avian Paleontology at the Close of the 20th Century: Proceedings of the 4th International Meeting of the Society of Avian Paleontology and Evolution, Washington, D.C., 4-7 June 1996, pp. 1–38.

    Mourer-Chauviré, C., Bour, R., & Ribes, S. (2006) Recent avian extinctions on Réunion (Mascarene Islands) from paleontological and historical sources. Bulletin British Ornithologists Club, 126, Supplement, 40–48.

    Newton, E. (1861) Ornithological notes from Mauritius. No.II. A ten days’ sojourn at Savanne. Ibis, 1, 3, 270–277.

    Newton, A. (1879) Remarks upon the specimen of Alectroenas nitidissima exhibited by Dr. Traquair. Proceedings of the Zoolological Society of London, pp. 2-4.

    Newton, A. (1882) Scopoli's ornithological papers from his Deliciae Florae et Faunae Insubricae (Ticini: 17861788). (I-IV), 1–20. London, The Willughby Society.

    Newton, E., & Gadow, H. (1893) On additional bones of the dodo and other extinct birds of Mauritius obtained by Mr. Théodore Sauzier. Transactions of the Zoolological Society of London, 13, 281–302.

    North-Coombes, [G.] A. (1971) The island of Rodrigues. [author], Port Louis, Mauritius, 338pp. [reprinted 2002, with new introduction by M. & P.North-Coombes, Mauritius: [editors]]

    Oustalet, E. (1897) Notice sur la faune ornithologique ancienne et moderne des Iles Mascareignes et en particulier de I’lle Maurice. Annales des Sciences Naturelles; Zoologie, 8, 3, 1–128.

    Pereira, S.L., Johnson, K.P., Clayton, D.H., & Baker, A.J. (2007) Mitochondrial and nuclear DNA sequences support a Cretaceous origin of Columbiformes and a dispersal-driven radiation in the Paleogene. Systematic Biology, 56, 4, 656–672.

    Peters, J.L. 1937. Check-list of the birds of the world (III). Museum of Comparative Zoology, Cambridge Masschusetts, 311 pp.

    Pollen, F.P.L. 1863. N.T.D., Volume I. M. Westerman & Zoon, Amsterdam, 383 pp.

    Prévost, F. (1843) (in Knip de Courcelles, P). Les pigeons, Volume 2, [1838–1843].

    Prys-Jones, R.P. & Diamond, A.W. (1984) Ecology of the land birds on the granitic and coralline islands of the Seychelles, with particular reference to Cousin Island and Aldabra Atoll. in: Stoddart, D.R. (Ed.), Biogeography and ecology of the Seychelles Islands. Dr. W. Junk Publishers, The Hague, pp. 529–558.

    Rand, A.L. (1936) The distribution and habitats of Madagascar birds. Bulletin of the American Museum of Natural History, 72, 142–499.

    Reichenbach, H.G.L. (1855) [text] Die vollstandigste naturgeschichte der tauben und taubenartigen vogel. Wallnister, Erdtauber, Baumtauben, Hocco’s. Columbariae, Megapodinae, Persisterinae, Columbninae, Alectorinae [Handwritten date of 1862] p.70, item no, 160. [Illustration] Die vollstandigste naturgeschichte der Tauben und taubenartigen vogel, Columbariae. Les Pigeons, Les Penelopes et Les Hoccos. Plate CXXXIII, figure 1260 (plate also numbered 224).

    Renshaw, G. (1906) The pigeon Hollandais. The Zoologist, 76, 49–52.

    Renshaw, G. (1939) Some extinct birds. Bird Notes and News, 18, 5, 109–110.

    Ridgeway, R. (1896) On the birds collected by Dr. W. L. Abbott in the Seychelles, Amirantes, Gloriosa, Assumption, Aldabra and adjacent islands, with notes on habits etc., by the collector. Proceedings of the United States National Museum, 18, 509–546.

    Rothschild, W. (1907) Extinct birds. Hutchinson, London, 244 pp.

    Rountree, F.R.G. 1952. Catalogue of the birds of Mauritius. Mauritius Institute Bulletin, 3, 155–217.

    Safford, R.J. & Evans, M.I. (1992) Birds on the Comoro Islands, April 1990. Scopus, 15, 93–101.

    Salvadori, T. 1893. Catalogue of the birds in the British Museum, volume 21. Trustees of the British Museum, London.

    Schlegel, H., & Pollen, F.P.L. (1868) Mammifères et oiseaux. Recherches sur la faune de Madagascar et de ses dépendances, d'après les découvertes de François P.L.Pollen et D.C. van Dam. Volume 2. J.K.Steenhoff, Leiden.

    Scopoli, J.A. (1786) Deliciae florae et faunae insubricae seu novae, aut minus cognitae species plantarum et animalium quas insubria Austriaca. Fasc. 2, 93. Ticini, Monastierii S. Salvatoris.

    Shapiro, B., Sibthorpe, D., Rambaut, A., Austin, J., Wragg, G.M., Bininda-Emonds, O.R.P., Lee, P.L.M. & Cooper, A. (2002) Flight of the dodo. Science, 295, 1683.

    Shaw, G. (1819) General zoology. Volume 11, Part 1, Aves by James Francis Stephens, 1819. G. Kearsley, London.

    Shelley, G.E. (1883) On the Columbidae of the Ethiopian region. Ibis, 5, 1, 258–331.

    Siddall, M., Rohling, E.J., Almogi-Labin, A., Hemleben, C., Meischner, D, Schmelzer, I.& Smeed, D.A. (2003.)Sea-level fluctuations during the last glacial cycle. Nature, 423, 853–858.

    Skerrett, A., Bullock, I. & Disley, T. (2001) Birds of the Seychelles. Christopher Helm (Publishers) Ltd, London, 320 pp.

    Snouckaert, van S.R. (1935) Les pigeons de genre Alectroenas. Alauda, 7, 328–335.

    Sonnerat, P. (1782) Voyage aux Indes Orientales et à la Chine : fait par ordre du roi, depuis 1774 jusqu’en1781 ; suivi d’observations sur le Cap de Bonne-Espérance, les îles de France et de Bourbon, les Maldives, Ceylan, Malacca, les Philippines et les Moluques et de recherches sur l’histoire naturelle de ces pays. Dentu, Paris, 3 volumes.

    Staub, F. (1993) Fauna of Mauritius and associated flora. Précigraph Ltd, Mauritius, 97 pp.

    Steadman, D.W. (1992) New species of Gallicolumba and Macropygia (Aves: Columbidae) from archeological sites in Polynesia, in: Papers in Avian Paleontology Honoring Pierce Brodkorb, (Ed.) Kenneth. E. Campbell Jr. Los Angeles County Museum of Natural History, Science Series, 36, 329–348.

    Steadman, D.W. (1997) The historic biogeography and community ecology of Pacific island pigeons and doves. Journal of Biogeography, 24, 737–753.

    Steadman, D.W. (2006) Extinction and biogeography in tropical Pacific birds. Chicago & London: University of Chicago Press. 594 pp.

    Stoddart, D.R., & Benson, C.W. (1970) An old record of a blue pigeon Alectroenas species and sea‑birds on Farquhar and Providence, Atoll Research Bulletin, 136, 35–6.

    Strickland, H.E. & Melville, A.G. (1848) The dodo and its kindred. Reeve, Benham & Reeve, London, 141 pp.

    Sweet J.M. 1970. The collection of Louis Dufresne (1752-1832). Annals of Science, 26, 33–71.

    Swinnerton, K.J., Greenwood, A.G., Chapman, R.E. & Jones, C.G. (2005a) The incidence of the parasitic disease trichomoniasis and its treatment in reintroduced and wild Pink Pigeons Columba mayeri. Ibis, 147, 772–782.

    Swinnerton. K.J., Pierce, M.A., Greenwood, A., Chapman, R.E. & Jones, C.G. (2005b) Prevalence of Leucocytozoon marchouxi in the endangered Pink Pigeon Columba mayeri. Ibis, 147, 725–737.

    Tafforet, J [attrib.]. ca.1726. Relation de L’ile Rodrigue. MS in the Archives Nationales in Paris. [published in full, in original orthography, in Proceedings of the Royal Society of Arts and Science, Mauritius, 4, 1–16].

    Temminck, C.J. & Knip, P. (1811) Les pigeons par Madame Knip née Pauline de Courcelles, premier peintre d'histoire naturelle de S.M. Marie-Louis, le texte par Coenraad C.J. Themminck [sic], directeur de l'Académie des Sciences et des Arts de Harlem [sic] etc. Knip & Garnery, Paris.

    Temminck, C.J. (1813–15) Histoire naturelle des pigeons et des gallinacées. J.C.Sepp & fils, Amsterdam & G.Dufour, Paris, volume 1 [1813], 500 pp.

    Toussaint, A. (1972) Histoire des Iles Mascareignes. Berger-Levrault, Paris, 345 pp.

    Tuijn, P. (1969) Notes on the extinct pigeon from Mauritius Alectroenas nitidissima (Scopoli, 1786). Beaufortia, 16, 218, 163–170.

    Vaughan, R.E. & Wiehe, P.O. (1937) Studies on the vegetation of Mauritius. 1. A preliminary survey of the plant communities. Journal of Ecology, 25, 289–343.

    Vigors, N.A. (1825) Sketches in ornithology; or observations on the leading affinities of some of the more extensive groups of birds. Journal of Ecology, 1, 308–346.

    Voisin, C., Voisin. J.-F, Jouanin. J. & Bour, R. (2004) Liste des types d’oiseaux des collections du Muséum national d’Histoire naturelle de Paris. 13 : gangas et pigeons (Pteroclididae et Columbidae), première partie. Zoosystema, 26, 1, 107–128.

    Wagler, J. (1827) Systema avium. Stuttgartiae, Cottæ, 411 pp.

    Warren, B.H., Strasberg, D., Bruggemann, J.H., Prys-Jones, R. & Thébaud, C. (2010) Why does the biota of the Madagascar region have such a strong Asiatic flavor? Cladistics, 26, 526–538.

    Worthy, T.H., & Holdaway, R.N. (2002) The lost world of the moa: prehistoric life of New Zealand. Indiana University Press, Bloomington and Indianapolis, 718 pp.

    Worthy, T.H., Hand, S.J., Worthy, J.P., Tennyson, A.J.D. & Scofield, R.P. (2009) A large fruit pigeon (Columbidae) from the early Miocene of New Zealand. Auk, 126, 3, 649–656.