Abstract
Masked water snakes of the genus Homalopsis are abundant in the wetlands of Southeastern Asia. Currently, two speciesare recognized, the widespread H. buccata Linnaeus and the Mekong drainage endemic H. nigroventralis Deuve. On thebasis of morphology we resurrect H. hardwickii Gray and H. semizonata Blyth and describe a new species from Indochina.We establish a neotype for Coluber buccatus Linnaeus and examine the status of other names associated with this species.Homalopsis species can be distinguished from each other on the basis of dorsal scale row counts, the scales in the ocularring, and other head scale architecture. The new species, Homalopsis mereljcoxi, is distinguished from all others by itsone postocular and one postsubocular scale, and a higher dorsal scale row count (40–47 scale rows at midbody) that arereduced to more than 30 scale rows at the body’s posterior. The new species is heavily exploited for the skin trade at TonleSap, Cambodia. Homalopsis likely contains other cryptic species that have evolved in the changing aquatic habitats of Southeast Asia. A key to the species of Homalopsis is provided.References
Ahmed, M.F., Das, A. & Dutta, S.K. (2009) Amphibains and Reptiles of Northeast India, A Photographic Guide. Aaranyak, Guwah, India. 167 pp.
Alfaro, M.E., Karns, D.R., Voris, H.K., Brock, C.D. & Stuart, B.L. (2008) Phylogeny, evolutionary history, and biogeography of Oriental-Australian rear-fanged water snakes (Colubroidea: Homalopsidae) inferred from mitochondrial and nuclear DNA sequences. Molecular Phylogenetics and Evolution, 46, 76–593.
Andersson, L.G. (1899) Catalogue of Linnean type-specimens of snakes in the Royal Museum in Stockhlom, Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar, Stockholm, 24, 1–35.
Andersson, L.G. (1900) Catalogue of Linnaean type-specimens of Linnaeus’s reptilia in The Royal Museum in Stockholm. Bihang till Kongl. Svenska Vetenskaps-Akademiens Handlingar, Stockholm, 26, 1–29.
Auliya, M.A. (2006) Taxonomy, life history and conservation of giant reptiles in West Kalimantan (Indonesia Borneo). Natur und Tier – Verlag GmbH, Münster. 432 pp.
Baker, B.W. (2006) Forensic Implications of dorsal scale row counts on puff-faced water snakes (Colubridae: Homalopsinae: Homalopsis buccata). Herpetological Review, 37, 171–173.
Barbour, T. (1912) A contribution to the zoogeography of the East Indian Islands. Memoires of the Museum of Comparative Zoology, 44, 1–203
Bergman, R.A.M. (1951) The anatomy of Homalopsis buccata. Proceeding of the Koninklijke Nederlandse Akademie van Wetenschappen, 54C, 511–524.
Bergman, R.A.M. (1962) Die anatomie der Elapinae. Zeitschrift für Wissenschaftliche Zoologie, Leipzig, 167, 291–337.
Berry, P.Y. & Lim, G.S. (1967) The breeding pattern of the puff-faced water snake, Homalopsis buccata Boulenger. Copeia, 1967, 307–313.
Bezuijen, M.R. Vinn, B. & Send, L. (2009) A collection of amphibians and reptiles from the Mekong River, north-eastern Cambodia. Hamadryad, 34, 135–164.
Bickford, D., Lohman, D. J., Sodhi, N.S., Ng, P.K.L., Meier, R. Winker, K., Ingram, K.K. & Das, I. (2007) Cryptic species as a window on diversity and conservation. Trends in Ecology and Evolution, 22, 148–155.
Blanford, W. T. (1881) On a collection of reptiles and frogs, chiefly from Singapore. Proceedings of the Zoological Society of London, 1881, 215–227.
Bleeker, P. (1857) Opsomming der soorten van Reptiliën, tot dus verre van het eiland Java bekend geworden. Natuurkundig Tijdschrift voor Nederlandsch Indië, 14, 235–246.
Blyth, E. (1855) Collections presented by Capt. Beddome and Mr. Theobald. Journal of the Asiatic Society of Bengal, Calcutta, 24, 187.
Blyth, E. (1859) Report of Curator, Zoological Department, for February to May meetings, 1859. Journal and Proceedings of the Asiatic Society of Bengal, Calcutta, 28, 279.
Boettger, O. (1898) Katalog der Reptilien-Sammlung im Museum der Senckenbergischen naturforschenden Gesellschaft in Frankfurt am Main. II. Teil. Schlangen. Gebrüder Knauer, Frankfurt am Main. 160 pp.
Boie, H. (1826) Merkmale einiger japanischer Lurche. Isis von Oken, 19, 203–216.
Bosch, H.A.J. in den (1985) Snakes of Sulawesi: checklist, key, and additional biogeographical remarks. Zoologische Verhandelingen, 217, 1–50.
Boulenger, G.A. (1887) An account of the reptiles and batrachians obtained in Tenasserim by M. L. Fea, of the Genoa Civic Museum. Annali del Museo Civico di Storia Naturale Giacomo Doria, (ser. 2), 5, 474–486.
Boulenger, G.A. (1890) The Fauna of British India, including Ceylon and Burma. Reptilia and Batrachia. Taylor and Francis, London. 541 pp.
Boulenger, G.A. (1896) Catalogue of the Snakes in the British Museum, Vol.3. British Museum (Natural History), London. 727 pp.
Boulenger, G.A. (1905) Report on the batrachians and reptiles. Fasciculi Malayenses Zoology, 1, 1–164
Boulenger, G.A. (1912) A Vertebrate Fauna of the Malay Peninsula from the Isthmus of Kra to Singapore Including the Adjacent Islands. Reptilia and Batrachia. Taylor and Francis, London. 294 pp.
Bourret, R. (1927) La Faune de l'Indochine. Les Vertebratés. Inventaire Général de l’Indochine, 3, 1–265.
Bourret, R. (1934) Notes herpétologiques sur l’Indochine française. IV. Sur une collectio d’Ophidiens de Cochine et du Cambodge. Bulletin Générale de l’Instruction Publique, Hanoi, September (1), 1–20.
Bourret, R. (1936) Les Serpents de L'Indochine. Tome I1. Catalogue systématique descriptif. Henri Basuyau & Co., Toulouse. 505 pp.
Brongersma, L.D. (1929) Appendix III. A list of the reptiles from Java. Pp. 64-68 In: On the zoogeography of Java. K. W. Dammerman (ed.). Treubia, 11, 1–88.
Brongersma, L.D. (1931) Resultats scientifiques du Voyage aux Indes Orientales Néerlandaises. Reptilia. Mémories du Musée Royal d’Histoire Naturelle de Belgique, 5, 1–39.
Brooks, S.E., Reynolds, J.D. & Allison, E.H. (2008) Sustained by snakes? Seasonal livelihood strategies and resource conservation by Tonle Sap fishers in Cambodia. Human Ecology, 36, 835–851.
Brooks, S.E., Allison, E.H. Gill, J.A. & Reynolds, J.D. (2009) Reproductive and Trophic Ecology of an Assemblage of Aquatic and Semi-Aquatic Snakes in Tonle Sap, Cambodia. Copeia, 2009, 7–20.
Brooks, S.E., Allison, E.H. Gill, J.A. & Reynolds, J.D. (2010) Snake prices and crocodile appetites: Aquatic wildlife supply and demand on Tonle Sap Lake, Cambodia. Biological Conservation, 143, 2127–2135.
Brooks, S.E., Allison, E.H. & Reynolds, J.D. (2007) Vulnerability of Cambodian water snakes: Initial assessment of the impact of hunting at Tonle Sap Lake. Biological Conservation, 139, 401–414.
Campbell, J.A. & Lamar, W.W. (2004) The Venomous Reptiles of the Western Hemisphere. 2 Volumes. Comstock Publishing Associates, Ithaca. 870 pp.
Campden-Main, S.M. (1970) A Field Guide to the Snakes of South Vietnam. Smithsonian Institution, Washington, D.C. 114 pp.
Cantor, T. (1836) Sketches of two undescribed venomous Serpentes with fangs behind the maxillary teeth (“a crochets posterieurs”). Transactions of the Medical and Physical Society, 8, 135–142.
Cantor, T. (1839) Spicilegium seperntum indicorum. Proceedings of the Zoological Society of London, 1839, 31–55.
Cantor, T. (1847) Catalogue of reptiles inhabiting the Malayan Peninsula and islands, collected or observed. Journal of the Asiatic Society of Bengal, Calcutta, 16, 1–157.
Chan-ard, T., Grossman, W., Gumprecht, A. & Schulz, K.-D. (1999) Amphibians and Reptiles of Peninsular Malaysia and Thailand, An Illustrated Checklist. Bushmaster Publications, Wuerselen. 240 pp.
Chanhome, L., Pauwels, O.S.G., Jintakune, P. & David, P. (2001) Catalog of the herpetological collection of the Queen Saovabha Memorial Institute, Thai Red Cross Society, Bangkok. Part 1. Snakes (except Elapidae and Viperidae). Bulletin of the Maryland Herpetological Society, 37, 49–72.
Cochran, D. (1929) Natural History collecting by Dr. Hugh M. Smith in Siam. In: Explorations of the Smithsonian Institution in 1927. Pp. 41–44.
Cochran, D. (1930) The herpetological collections made by Dr. Hugh M. Smith in Siam from 1923 to 1929. Proceeding of the United States National Museum, 77, 1–39.
Cox, M.J. (1991) The Snakes of Thailand and Their Husbandry. Krieger Publishing Company, Malabar. 536 pp.
Cox, M.J., van Dijk, P.P., Nabhitabhata, J. & Thirakhaupt, J. (1998) A Photographic Guide to Snakes and other Reptiles of Peninsular Malaysia, Singapore, and Thailand. Ralph Curtis Publishing, Sanibel Island. 142 pp
Dammerman, K.W. (1926) The fauna of Durian and the Rhio-Lingga Archipelago. Treubia, 8, 281–326.
Das, I., Dattagupta, B. & Gayen, N.C. (1998) History and catalogue of reptile types in the collection of The Zoological Survey of India. Journal of South Asian Natural History, 3, 1–52.
Daudin, F.M. (1803a) Historie naturelle, générale et particuliére des reptiles, pour faire suite à l’histoire naturelle de Buffon, rédigée par Sonnini. Volume 6. F. Dufart, Paris.
Daudin, F.M. (1803b) Historie naturelle, générale et particuliére des reptiles, pour faire suite à l’histoire naturelle de Buffon, rédigée par Sonnini. Voume 7. F. Dufart, Paris. 436 pp.
David, P. and G. Vogel. (1996) The Snakes of Sumatra. An Annotated Checklist and Key with Natural History Notes. Edition Chimaira, Frankfurt am Main. 260 pp. 2 maps. 8 figs.
De Lang, R. & Vogel, G. (2000) The Snakes of Sulawesi. Edition Chimaira, Frankfurt am Main. 312 pp.
Despax, R. (1912) Sur trois collection de reptiles et de batraciens provenant de l’Archipel Malais. Bulletin du Muséum d’Historie Naturelle, Paris, 18, 198–205.
Deuve, J. (1970) Serpents du Laos. Mémoire Office de Recherche Scientifique et Technologique Outre Mer, Paris, 39, 1–251.
Dowling, H.G. (1951) A proposed standard system of counting ventrals in snakes. British Journal of Herpetology, 1, 97–99.
Dowling, H.G. (1974) A provisional classification of snakes. In: 1974 HISS Yearbook of Herpetology, H. G. Dowling (ed.). New York: HISS, American Museum of Natural History. Pp.167–170.
Dowling, H. G. & Jenner, J.V. (1988) Snakes of Burma, Checklist of reported species and bibliography. Smithsonian Herpetological Information Service, (76), 1–19.
Dubey, S., Brown, G.P., Madsen, T. & Shine, R. (2009) Sexual selection favours large body size in males of a tropical snake (Stegonotus cucullatus, Colubridae). Animal Behavior, 77, 177–182.
Dunn, E. R. (1927) Results of the Douglas Burden expedition to the Island of Komodo. II. Snakes from the East Indies. American Museum Novitates, 144, 1–7.
Fabien, A., Bonnet, X., Maumelat S., Bradshaw, D. & Schwaner, T. (2004). Diet divergence, jaw size and scale counts in two neighboring populations of tiger snakes (Notechis scutatus). Amphibia-Reptilia, 25, 9–7.
Firth, C.B. (1977) A survey of the snakes of Phuket Island and the adjacent mainland areas of peninsular Thailand. Natural History Bulletin of the Siam Society, 26, 263–316.
Fitzinger, L.I. (1826) Neue Classification der Reptilien. J G Heubner, Wien. 66 pp.
Flower, S. S. (1896) Notes on a collection of reptiles and batrachians made in the Malay Peninsula in 1895–96; with a list of the species recorded from that region. Proceedings of the Zoological Society of London, 1896, 856–914.
Fuchs, K. & Fuchs, M. (2003) The Reptile Skin: A Key-Feature in the Identification of Lizards and Snakes. Edition Chimera, Frankfurt Am Main. 407 pp.
Geissler, P., Nguyen, T. Q., Poyarkov, N.A. & Böhme, W. (2011) New records of snakes from Cat Tien National Park, Dong Nai and Lam Dong provinces, southern Vietnam. Bonn Zoological Bulletin, 60, 9–16
Gibbons, J.W. & Lovich, J.E. (1990) Sexual dimorphism in turtles with emphasis on the slider turtle (Trachemys scripta). Herpetological Monographs, 4, 1–29.
Gmelin, J.F. (1789) Caroli a Linne...Systema naturae per regna tria natural, secundum classes, ordines, genera, species, cum characteribus differentilis, synonymis, locis. Tomus I, Editio decima tertia, aucta, reformata. Pars III. Amphibia et Pisces. Georg. Emanuel Beer, Lipsiae, pp.1033–1516.
Gray, J. (1842) Monographic synopsis of the watersnakes, or the family Hydridae. In: Zoological Miscellany, pp. 59–71.
Gray, J. 1849. Catalogue of the Specimens of Snakes in the Collection of the British Museum. Trustees of the British Museum (Natural History), London. 125 pp.
Grismer, L. L., Neang, T. Chav, T. & Grismer, J. L. (2008) Checklist of the amphibians and reptiles of the Cardamom region of southwestern Cambodia. Cambodian Journal of Natural History, 2008, 12–28.
Günther, A.C.L.G. (1864) Reptiles of British India. Ray Society, London. 443 pp.
Gyi, K.K. (1970) A revision of colubrid snakes of the subfamily Homalopsinae. University of Kansas Publications, Museum of Natural History, 20, 47–223.
Gyldenstolpe, N. (1916) Zoological results of the Swedish Zoological Expeditions to Siam 1911-1912 and 1914-1915. I. Snakes. Kongliga Svenska Vetenskaps-Akademiens. Handlingar. 55, 17-18.
Haas, C.P.J. de (1950) Checklist of the snakes of the Indo-Australian Archipelago (Reptiles, Ophidia). Treubia, 20, 511–625.
Hoesel, J. K.P. van 1959. Ophidia Javanica. Museum Zoologicum Bogoriense, Bogor. 188 pp.
Hundley, H.G. (1964) Checklist of Burmese Reptiles. H.G. Hundley, Kyi, Burma. 32 pp.
Jayne, B.C., Voris, H.K. & Heang, K.B. (1988) Diet, feeding behavior, growth, and numbers of a population of Cerberus rynchops (Serpentes: Homalopsinae) in Malaysia. Fieldiana: Zoology, (50), 1-15.
Jenkins, M. & Broad, S. (1994) International Trade in Reptile Skins. A review and analysis of the main consumer markets 1983-1991. Traffic Net Work Report. 68 pp.
Karns, D.R., Murphy, J.C. & Voris, H.K. (2010) Semi-aquatic Snake Communities of the Central Plain Region of Thailand. The Natural History Journal of Chulalongkorn University, 10, 1–25.
Karns, D.R., Murphy, J.C., Voris, H.K. & Suddeth, J.S. (2005) Comparison of semi-aquatic snake communities associated with the Khorat Basin, Thailand. The Natural History Journal of Chulalongkorn University, 5, 73–90.
Karns, D.R., Voris, H.K. & Goodwin, T.G. (2002) Ecology of the Oriental-Australian rear-fanged water snakes (Colubridae: Homalopsidae) in the Pasir Ris Park Mangrove Forest, Singapore. The Raffles Bulletin of Zoology, 50, 487–498.
Kelly, C.M.R., Barker, N.P. & Villet, M.H. (2003) Phylogenetics of advanced snakes (Caenophidia) based on four mitochondrial genes. Systematic Biology, 52, 439–459.
Kopstein, F. (1930) Herpetologische notizen: II. Oologische Beobachtungen an West Javanischen Reptilien. Treubia, 11, 301–307.
Kopstein, F. (1938) Ein Beitrag zur Eierkunde und zur Fortpflanzung der Malaiischen Reptilien. Bulletin of the Raffles Museum, 14, 168–174.
Kopstein, F. (1941) Über sexualdimorphismus bei Malaiischen schlangen. Temminckia, 6, 109–185.
Kuhl, H. and Hasselt, J.C. van. (1822) Uittreksel uit de brieven van Heeren Kuhl en van Hasselt aan de Heeren C. J. Temminck, Th. Van Swinderen en W. de Haan. Algemeene Konsten Letter Bode voor het jaar 1822, 1, 99-104.
Laidlaw, F.F. 1901. List of a collection of snakes, crocodiles, and chelonians from the Malay Peninsula, made by members of the “Skeat Expedition,” 1899-1900. Proceedings of the Zoological Society of London, 2, 575-583.
Lampe, E. and Lindholm, W.A., 1902. Catalog der Reptilien und Amphibien – Sammlung des Naturhistorischen Museums zu Wiesbaden. Jahrbuch des Nassauischen Vereins für Naturkunde, 55, 1-66.
Lawson, R. Slowinski, J.B., Crother, B.I., and Burbrink, F.T. 2005. Phylogeny of the Colubroidea (Serpentes): new evidence from mitochondrial and nuclear genes. Molecular Phylogenetics and Evolution, 37, 581–601.
Lillywhite, H.B. (2008) Dictionary of Herpetology. Krieger Publishing Company, Malabar. 376 pp.
Lim, B.L. (1955) Snakes collected near Kuala Lumpur. Malayan Nature Journal, 9, 122–125.
Lim, B.L. (1956) The natural food of some Malayan snakes. Malayan Nature Journal, 10, 139–144.
Lim, B.L. (1964) Notes on the elephant trunk snake and the puff-faced water snake in Kuala Lumpur. Malayan Nature Journal, 18, 179–183.
Lim, L.K.L. & Lee, T.M.M. (1988) Fascinating Snakes of Southeast Asia, An Introduction. Tropical Press Sdn. Bhd., Kuala Lumpur 124 pp.
Lim, K.K.P. & Lim, F.L.K. (1992) A Guide to the Amphibians and Reptiles of Singapore. Singapore Science Center, Singapore. 160 pp.
Linnaeus, C. (1754) Museum S:ae R:ae M:tis Adolphi Friderici Regis. Svecorum, Gothorum, Vandalorumque. Hær. Norv. Duc. Slesv. Hols. Storm. Ditm. Com. Oldenb. Delmenhorstiæ &c. &c. in quo Animalia Rariora Imprimis, et Exotica: Quadrupedia, Aves, Amphibia, Pisces, Insecta, Vermes describuntur et determinatur. Typographia Regia, Holmiae, 1-104, pls. 1-33.
Linnaeus, C. (1758) Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii, Holmiæ. 824 pp.
Linnaeus, C. (1789) Systema naturae per regna tria naturae. secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis locis. Tomus I, Editio decima tertia aucta, reformata. Pars III. Amphibia et Pisces. Georg. Emanuel Beer, Lipsiae. Pp. 1033-1516.
Lloyd, D. J., Cotton, F.R., Parker, H.W. & Prain, D. (1933) The collection of reptile skins for commercial purposes with respect to the possibilities in Empire countries. Imperial Institute, London. 36 pp.
Lönnberg, E. (1896) Linnean type-specimens of birds, reptiles, batrachians and fishes in the Zoological Museum of the R. University in Upsala. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar, 22, 1–45.
Manthey, U. & Grossmann, W. (1997) Amphibien & Reptilien Südostasiens. Natur und Tier – Verlag, Münster. 512 pp.
Marx, H. & Rabb, G.B. (1972) Phyletic analysis of fifty characters of advanced snakes. Fieldiana Zoology, 68, 1–321.
McDiarmid, R.W., Campbell, J.A. & Touré, T.A. (1999) Snake Species of the World. A Taxonomic and Geographical Reference. Volume 1. The Herpetologists League, Washington. 519 pp.
McLeod, D.S. (2010) Of Least Concern? Systematics of a cryptic species complex: Limnonectes kuhlii (Amphibia: Anura: Dicroglossidae) Molecular Phylogenetics and Evolution, 56, 991-1000.
Mell, R. (1922) Beiträge zur fauna sinica. Die vertebraten Südchinas: feldlisten und feldnoten der Saüger, Vögel, Reptilien, Batrachier. Archiv für Naturgeschichte, 88A, 1–134.
Mertens, R. (1929) Über eine kleine herpetologische Sammlung aus Java. Senckenbergiana Biologica, 11, 22–33.
Mocquard, F. (1892) Voyage de M. Chaper. A. Borneo. Nouvelle contributiuon a la faune herpétologique de Borneo. Mémoires de la Société Zoologique de France, 5, 190–206.
Mocquard, F. (1907) Les reptiles de l’Indo-Chine (conférence faite au Muséum d’Histoire naturelle, 19 avril 1906). Augustin Challamel, Paris. 59 pp. Morice, A. (1875) Coup d'Oeil sur la Fauna de la Cochinchine Francaise. H. Georg, Lyon, 1–64.
Müller, F. (1878) Katalog der im Museum und Universitätskabinet zu Basel aufgestellten Amphibien und Reptilien nebst Anmerkungen. Verhandlungen der Naturforschenden Gesellschaft in Basel 6, 557–709.
Murphy, J.C., Voris, H.K., Karns, D.R., Chan-ard, T.& Suvurat, K.. (1999) The ecology of the water snakes of Ban Tha Hin, Songkhla Province, Thailand. Natural History Bulletin of the Siam Society, Bangkok, 47, 129–147.
Murphy, J.C. (2007) Homalopsid Snakes, Evolution in the Mud. Krieger Publishing Company, Malabar. 249 pp.
Nguyen, V. S. (2007) “Fauna of Vietnam. 14. Serpentes.” Science and Technics Publishing House, Ha Noi. 247 pp.
Nguyen, V. S. & Ho, T. C. (1996) “Checklist of reptiles and amphibians of Vietnam.” Science and Technics Publishing House, Ha Noi. 264 pp. (in Vietnamese) (Homalopsis buccata from S. Vietnam, p. 127-128).
Nguyen, V. S., Ho, T. C. & Nguyen, Q. T. (2009) Herpetofauna of Vietnam. Edition Chimaira, Frankfurt am Main. 768 pp.
Oldfield Thomas, F. (1911) The mammals of the tenth edition of Linnaeus; an attempt to fix the types of the genera and the exact bases and localities of the species. Proceedings of the Zoological Society of London, 81, 120–158.
Pauwels, O.S.G., Laohawat, O.-A., David, P., Bour, R. Dangsee, P., Puangjit, C. & Chimsunchart, C. (2000) Herpetological investigations in Phang-Nga Province, southern peninsular Thailand, with a list of reptile species and notes on their biology. Dumerilla, 4, 123–154.
Pauwels, O.S.G., Wallach, V. & David, P. (2008) Global diversity of snakes (Serpentes; Reptilia) in freshwater. Hydrobiologia, 595, 599–605.
Peters, J.A. (1964) Dictionary of Herpetology. Hafner Publishing Company, New York. 392 pp.
Presch, W. (1973). A Review of the Tegus, Lizard Genus Tupinambis (Sauria: Teiidae) from South America. Copeia, 1973, 740–746.
Pyron RA & Burbrink FT (2011) Extinction, ecological opportunity, and the origins of global snake diversity. Evolution, online in advance of print. doi:10.1111/j.1558-5646.2011.01437.x
Pyron, R.A., Burbrink, F.T., Colli, G.R. de Oca, A.N.M., Vitt, L.J., Kuczynski, C.A. & Wiens. J.J. (2010) The phylogeny of advanced snakes (Colubroidea), with discovery of a new subfamily and comparison of support methods for likelihood trees. Molecular Phylogenetics and Evolution, 58, 329–342.
Ridley, H.N. (1899) The habits of Malay reptiles. Journal of the Straits Branch Royal Asiatic Society, 32, 185–210.
Rooij, N. de, (1917) The Reptiles of the Indo-Australian Archipelago. II. Ophidia. Leiden: E.J. Brill, Leiden. 334 pp.
Schätti, B. Tillack, F.& Helfenberger, N. (2009) A contribution to Spalerosophis microlepis Jan, 1865, with a short review of the genus and a key to the species (Squamata: Serpentes: Colubridae). Herpetozoa, 22, 115-135.
Schlegel, H. (1837) Essai sur la physionomie des serpens. II. Partie descriptive. Arnz & Companie, Leide. 606 pp.
Schleich, H.H. & Kästle, W. (eds.) (2002) Amphibians and Reptiles of Nepal. A.R.G. Ganter Verlag K.G., Koeltz Scientific Books, Koenigstein, Germany. 1201 pp.
Sclater, W.L. (1891) List of Snakes in the Indian Museum. The Indian Museum, Calcutta. 70 pp.
Seba, A. (1735) Locupletissimi rerum naturalium thesauri accurata descriptio, et iconibus artificiosissimis expressio, per universam physices historiam. Tomus II. H. Janssonio-Waesbergios, J. Wetstenium & Gul. Smith, Amstelaedami. 154 pp. + 114 pls.
Smith, M.A. (1914) The snakes of Bangkok. Part 1. Journal of the Natural History Society of Siam, Bangkok 1, 5–18.
Smith, M.A. (1930) The Reptilia and Amphibia of the Malay Peninsula. A supplement to G. A. Boulenger's Reptilia and Batrachia, 1912. Bulletin of the Raffles Museum, Singapore, 3, 1–149.
Smith, M.A. (1943) The Fauna of British India, Ceylon and Burma. Reptilia and Amphibia. Vol. 3. Serpentes. Taylor and Francis, London. 583 pp.
Spawls, S. & Branch. B. (1995) The Dangerous Snakes of Africa. Ralph Curtus Books, Sanibel Island. 192 pp.
St. Girons, H. & Pfeffer, P. (1971) Le cycle sexuel des serpents du Cambodge. Annales des Sciences Naturelles, 13, 543–572.
St. Girons, H. & Pfeffer, P., (1972) Notes sur l'ecologie des serpents du Cambodge. Zoologische Mededelingen, 47, 65–86.
St. Girons, H. (1972) Les Serpents du Cambodge. Mémoires du Muséum Natlional d’Histoire Naturelle, Paris, (sér. 2), 74A, 1–170.
Stuebing, R. B. (1991) A checklist of the snakes of Borneo. Raffles Bulletin of Zoology, 39, 323–362.
Steubing, R.B. & Inger, R.F. (1999) A Field Guide to the Snakes of Borneo, Natural History Publications, Kota Kinabalu. 254 pp.
Stuart, B.L. (2004) The harvest and trade of reptiles at U Minh Thuong National Park, Southern Viet Nam. TRAFFIC Bulletin, 20, 25–34.
Stuart, B.L., Sok, K. & Neang, T. (2006) A collection of amphibians and reptiles from hilly eastern Cambodia. Raffles Bulletin of Zoology, 5, 129–155.
Stuart, B.L., Smith, J., Davey, K., Prom, D. & Platt, S.G. (2000). Homalopsine watersnakes: the harvest and trade from Tonle Sap, Cambodia. TRAFFIC Bulletin, 18, 115–124.
Sworder, G.H. (1923) A list of the snakes of Singapore Island. The Singapore Naturalist, 1923, 55–73.
Taylor, E. H. & Elbel, R.E. (1958) Contributions to the herpetology of Thailand. University of Kansas Science Bulletin, 38, 1033–1189.
Taylor, E. H. (1965) The serpents of Thailand and adjacent waters. University of Kansas Science Bulletin, 45, 609–1096.
Teo, R. C. H. & Rajathurai, S. (1997) Mammals, reptiles and amphibians in the nature reserves of Singapore – diversity, abundance and distribution. Proceedings of the Nature Reserves Survey Seminar. Garden’s Bulletin, Singapore, 49, 353–425.
Theobald, W., Jr. (1868a) Catalogue of the reptiles of British Burma, embracing the provinces of Pegu, Martaban, and Tenasserim; with descriptions of new or little-known species. The Journal of the Linnean Society (Zoology), London, 10, 4-67.
Theobald, W., Jr. (1868b) Catalogue of Reptiles in the Museum of the Asiatic Society of Bengal, 1866. Journal of the Asiatic Society (extra number), Calcutta. 88 pp.
Theobald, W., Jr. (1876) Descriptive Catalogue of the Reptiles of British India. Thacker, Spink, and Co., Calcutta. 238 pp.
Tirant, G. (1885) Notes sur les reptiles et les batraciens de la Cochinchine et du Cambodge. Impromerie du Gouvernement, Saigon. 104 pp.
Tweedie, M.W.F. (1983) The Snakes of Malaya. Third edition. Singapore National Printers, Singapore. 139 pp.
Volz, W. (1904) Schlangen von Palembang (Sumatra). Zoologischer Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, Jena, 20, 491–508.
Vidal, N., Delmas, A.-S., David, P., Cruaud, C., Couloux, A. & Hedges, S.B. (2007) The phylogeny and classification of caenophidian snakes inferred from seven nuclear protein-coding genes. Comptes Rendus Biologies, 330, 182–187.
Voris, H.K. & Karns, D.R. (1996) Habitat utilization, movements, and activity patterns of Enhydris plumbea (Serpentes: Homalopsinae) in a rice paddy wetland in Borneo. Herpetological Natural History, 4, 111–126.
Voris, H.K. & Murphy, J.C. (2002) Prey and predators of homalopsine snakes. Journal of Natural History, 36, 1621–1632.
Wall, F. (1902) Aids to the differentiation of snakes. Journal of the Bombay Natural History Society, 14, 337–345.
Wall, F. (1903) A prodromus of the snakes hiterto recorded from China, Japan, and the Loo Choo Islands: with some notes. Proceedings of the Zoological Society of London, 1, 84–102.
Wall, F. (1905) Notes on snakes collected in Cannanore from 5th November to 5th August 1904. Journal of the Bombay Natural History Society, 16, 292–317.
Wall, F. (1923) How to Identify the Snakes of India. F. Wall, Karachi. 56 pp.
Wall, F. (1925) Notes on snakes collected in Burma in 1924. Journal of the Bombay Natural History Society, 30, 805–821.
Wall, F. (1924) A hand-list of the snakes of the Indian Empire. Part 3. Journal of the Bombay Natural History Society, 29, 864–878.
Wall, F. (1925). Notes on snakes collected in Burma in 1924. Journal of the Bombay Natural History Society, 30, 805–821.
Wall, F. & Evans G.H. (1901) Burmese snakes. Notes on specimens including 45 species of ophidian fauna collected in Burma from 1st January to 30th June 1900. Journal of the Bombay Natural History Society, 13, 611–620.
Werner, F. (1896) Zweiter Bietrag zur Herpetologie der indo-orientalischen Region. Verhanglungen der Zoologische-Botanischen Gesellscaft in Wien, 46, 6–23.
Werner, F. (1900) Reptilien und Batrachier aus Sumatra, gesammelt von Herrn Gustav Schneider jr. im Jahre 1897-98. Zoologischer Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere, Jena, 12, 461–508.
Westermann, J.H. (1942) Snakes from Bangka and Billiton. Treubia, 18, 611–619.
Whitaker, R. & Captain, A. (2004). Snakes of India, The Field Guide. Draco Books, Chennai, India. 479 pp.
Williams K.L. & Wallach, V. (1989) Snakes of the World. Volume 1. Synopsis of Snake Generic Names. Krieger Publishing Company, Malabar. 234 pp.
Yen, S.-H. (2004) Nomenclatural and Systematic Problems Surrounding Eterusia aedea (Linnaeus, 1763) (Lepidoptera: Zygaenidae: Chalcosiinae). Zoological Studies, 43, 20-34.
Zhou, Z. & Jiang, Z. (2004) International trade status and crisis for snake species in China. Conservation Biology, 18, 1368–1394.
Zug G.R. & Mitchell, J.C. (1995) Amphibians and reptiles of the Royal Chitwan National Park, Nepal. Asiatic Herpetological Research, 6, 172–180.