Abstract
The limits of zalmoxid distribution in Southeast Asia are poorly understood, but a focus of recent research. Here we describesix new species of litter-inhabiting harvestmen in the genus Zalmoxis Sørensen, 1886 (Opiliones: Laniatores: Zalmoxidae)using light microscopy and SEM. Three of these species are from the Philippine Islands (Zalmoxis gebeleizis sp. nov., Zalmoxisderzelas sp. nov., and Zalmoxis sabazios sp. nov.) and the other three from Borneo (Zalmoxis zibelthiurdos sp. nov., Zalmoxisbendis sp. nov., and Zalmoxis kotys sp. nov.). The collecting localities of these species add to the known range of Zalmoxidae,which have not previously been reported from Borneo. The new species add to known morphological variation of Zalmoxis, specifically with respect to sexually dimorphic tarsomeres, body size, and armature of the anal plate.References
Clouse, R.M., General, D.M., Diesmos, A.C. & Giribet, G. (2011) An old lineage of Cyphophthalmi (Opiliones) discovered on Mindanao highlights the need for biogeographical research in the Philippines. The Journal of Arachnology, 39, 147-153.
Forster, R.R. (1949) Opiliones from the Solomon Islands. Records of the Australian Museum, 22, 141–147.
Forster, R.R. (1955) Further Australian harvestmen. Australian Journal of Zoology, 3, 354–411.
Giribet, G., Edgecombe, G.D., Wheeler, W. C. & Babbitt, C. (2002) Phylogeny and systematic position of Opiliones: A combined analysis of chelicerate relationships using morphological and molecular data. Cladistics, 18, 5–70.
Giribet, G., Vogt, L., Pérez-González, A., Sharma, P. & Kury, A.B. (2010) A multilocus approach to harvestman (Arachnida: Opiliones) phylogeny with emphasis on the biogeography and systematics of Laniatores. Cladistics, 26, 408–437.
Herodotus (2003) The Histories, Revised. Translation by Aubery de Sélincourt. Edited by John M. Marincola. Third edition. Penguin Books, London, England, 683 pp.
Kury, A.B. (2003) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, vol. especial monográfico, 1, 1–337.
Kury, A.B. & Pérez-González, A. (2007) Zamoxidae Sørensen, 1886. In: Pinto-da-Rocha, R., Machado, G., & Giribet, G. (Eds.), Harvestmen: the biology of Opiliones. Harvard University Press, Cambridge, pp. 243–246.
Martens, J. & Schwendinger, P.J. (1998) A taxonomic revision of the family Oncopodidae I. New genera and new species of Gnomulus Thorell (Opiliones, Laniatores). Revue Suisse de Zoologie, 105, 499–555.
Murphree, C.S. (1988) Morphology of the dorsal integument of ten opilionid species (Arachnida, Opiliones). The Journal of Arachnology, 16, 237–252.
Roewer, C.F. (1912) Die Familien der Assamiiden und Phalangodiden der Opiliones-Laniatores (Assamiden, Dampetriden, Phalangodiden, Epedaniden, Biantiden, Zalmoxiden, Samoiden, Palpidediden anderer Autoren). Archiv für Naturgeschichte, Berlin, Abteilung A, 78, 1–242.
Roewer, C.F. (1926) Opiliones vom Mount Maquiling auf Luzon Philippinen. Philippine Journal of Science, 29, 545–559.
Roewer, C.F. (1927) Weitere Weberknechte I. Abhandlungen der Naturwissenschaftlichen Verein zu Bremen 26: 261–402.
Roewer, C.F. (1949) Über Phalangodiden I: Subfam. Phalangodinae, Tricommatinae, Samoinae; Weitere Weberknechte XIII. Senckenbergiana, 30, 11–61.
Schwendinger, P.J. (2007) Oncopodidae Thorell, 1876. In: Pinto-da-Rocha, R., Machado, G., & Giribet, G. (Eds.), Harvestmen: the biology of Opiliones. Harvard University Press, Cambridge, pp. 211–214.
Sharma, P. & Giribet, G. (2009) Sandokanid phylogeny based on eight molecular markers—The evolution of a southeast Asian endemic family of Laniatores (Arachnida, Opiliones). Molecular Phylogenetics and Evolution, 52(2), 432–447.
Sharma, P.P. & Giribet, G. (2011) The evolutionary and biogeographic history of the armoured harvestmen–Laniatores phylogeny based on ten molecular markers, with the description of two new families of Opiliones. Invertebrate Systematics, 25, 106–142.
Sharma, P.P. (2012) New Australasian Zalmoxidae (Opiliones: Laniatores) and a new case of male polymorphism in Opiliones. Zootaxa, 3236, 1–35.
Sharma, P.P., Kury, A.B. & Giribet, G. (2011) Zalmoxidae of the Paleotropics: a catalogue of Southeast Asian and Indo-Pacific species. Zootaxa, 2972, 37–58.
Simon, E. (1892) Arachnides des îles Philippines. Annales Société Entomologique de France, séries 6, 61, 35–52.
Sørensen, W. (1886) Opiliones Australasiae. In: Koch L. and Keyserling, E. (Eds.) Die Arachniden Australiens nach der Natur beschrieben und abgebidet. Bauer & Raspe, pp. 53–86.
Staręga, W. (1989) Harvestmen (Opiliones) from the Mascarene Islands and the resurrection of the family Zalmoxidae. Annals of the Natal Museum, 30, 1–8.
Suzuki, S. (1977) Report on a collection of opilionids from the Philippines. Journal of Science of the Hiroshima University B, 1, 27(1), 1–120.