Abstract
A new species of Scutiger Theobald, 1868 is described from Medog, southeastern Tibet, China. Scutiger wuguanfui sp.nov. is distinguished from congeners in the following combination of characters: (1) large adult size, female is much largerthan male; (2) maxillary teeth absent; (3) male with a pair of pectoral glands and a pair of axillary glands, axillary glandssimilar to pectoral glands, all of them covered by black spines in breeding season; (4) in breeding male, nuptial spines ondorsal surface of firs and second fingers, and inner side of third finger; (5) male with an internal single subgular vocal sac,a pair of slit-like openings of vocal sac near corners of the mouth. This new species is currently known only from the type locality.References
Das, I., Tron, R.K.L., Rangad, D. & Hooroo, K.R. (2010) A new species of Leptolalax (Anura: Megophryidae) from the sacred groves of Mawphlang, Meghalaya, north-eastern India. Zootaxa 2339, 44–56.
Dubois, A. (1979) Une espèce nouvelle de Scutiger (Amphibiens, Anoures) du nord de la Birmanie. Revue Suisse de Zoologie, 86, 631–640.
Dubois, A. (1973) Diagnoses de trois espèce nouvelles d’amphibiens du Népal. Séance, 23, 495–497.
Delorme, M. & Dubois, A. (2001) Une nouvelle espèce de Scutiger du Bhutan, et quelques remarques sur la classification subgénérique du genre Scutiger (Megophryidae, Leptobrachiinae). Alytes, 2001, 19, 141–153.
Fei, L., Hu, S.Q., Ye, C.Y. & Huang, Y.Z. (2009) Fauna Sinica, Amphibia, Vol. 2. Science Press, 1–957 [in Chinese].
Fei, L. & Ye, C.Y. (1989) Geographical distribution, origin and evolution, and centre of differentiation of high-altitude pelobatid toads in Asia (Amphibia: Pelobatidae). Zoological Research, 10 (4), 295–302 [in Chinese].
Fei, L., Ye, C.Y., Jiang, J.P., Xie, F. & Huang, Y.Z. (2005) An Illustrated Key to Chinese Amphibians. Sichuan Publishing House of Science and Technology, 1–340 [in Chinese].
Fei, L., Ye, C.Y. & Li, S.S. (1989) On the generic classification of Asian high altitude pelobatid toads (Amphibia: Pelobatidae). Acta Zoologica Sinica, 35 (4), 381–389 [in Chinese].
Frost, D.R. (2011) Amphibian Species of the World: an Online Reference. Version 5.5 (31 January, 2011). Electronic Database accessible at http://research.amnh.org/vz/herpetology/amphibia/, American Museum of Natural History, New York, USA.
Fu, J.Z., Lathrop, A. & Murphy R.W. (1997) Phylogeny of genus Scutiger (Amphibia: Megophryidae): a re-evaluation. Asiatic Herpetological Research, 7, 32–37.
Fu, J.Z., Weadic, C.J. & Bi, K. (2007) A phylogeny of the high-elevation Tibetan megophryid frogs and evidence for the multiple origins of reversed sexual size dimorphism. Journal of Zoology, 273, 315–325.
Hu, S.Q. (1987) Amphibia-Reptilia of Xizang. Science Press, 1–153 [in Chinese].
Li, P.P., Lu, Y.Y. & Rao, D.Q. (2008) A new species of cascade frog (Amphibia, Ranidae) from Tibet, China. Acta Zootaxonomica Sinica, 33 (3), 537–541.
Li, P.P., Zhao, E.M. & Dong, B.J. (2010) Amphibians and Reptiles of Tibet. Science Press, 1–251 [in Chinese].
Mahony, S., Sengupta, S., Kamei R.G. & Biju, S.D. (2011) A new low altitude species of Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from Assam, northeast India. Zootaxa, 3059, 36–46.
Mathew, R. & Sen, N. (2007) Description of two new species of Xenophrys from north-east India. Cobra, 1 (2), 18–28.
Myers, G.S. & Leviton, A.E. (1962) Generic classification of the high-altitude pelobatid toads of Asia (Scutiger, Aelurophryne, and Oreolalax). Copeia, 1962 (2), 287–291.
Rao, D.Q. (2005) The study on the phylogenetic relationship of Pelobatidea (Amphibia: Anura). Unpublished Ph.D. thesis, Kunming Institute of Zoology, the Chinese Academy of Sciences, 1–221 [in Chinese].
Sondhi, S. & Ohler, A. (2011) A blue-eyed Leptobrachium (Anura: Megophryidae) from Arunachal Pradesh, India. Zootaxa, 2912, 28–36.
WWF-Living Himalayas Initiative. (2009) The Eastern Himalayas - Where Worlds Collide, New Species Discoveries. http://www.worldwildlife.org/what/wherewework/easternhimalayas/WWFBinaryitem13235.pdf., World Wide Fund for Nature.
Ye, C.Y., Fei, L., Wei, G. & Xu, N. (1992) Study on the phylogenetic relationship between species of Scutiger genus in Qinghai-Xizang Plateau (Amphibia: Pelobatidae). Acta Herpetological Sinica, Vol. 1 and 2, 27–39 [in Chinese].
Zhao, W.G., Rao, D.Q., Lu, S.Q. & Dong, B.J. (2005) Herpetological surves of Xizang Autonomous Region, 2. Mêdog. Sichuan Journal of Zoology, 24 (3), 250–253 [in Chinese].