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Published: 2012-08-24
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Observations on the biology of Afrotropical Hesperiidae (Lepidoptera) principally from Kenya. Part 4. Hesperiinae: Aeromachini and Baorini

C/o CABI, Bakeham Lane, Egham, TW20 9TY, UK
African Butterfly Research Institute (ABRI), P.O. Box 14308, Nairobi, Kenya
Lepidoptera Ampittia Zenonia Pelopidas Borbo Parnara Gegenes food plant life history leaf shelter parasitoid

Abstract

Partial life histories for two Afrotropical Aeromachini and seven Afrotropical Baorini (Hesperiidae: Hesperiinae) are described and illustrated: Ampittia capenas (Hewitson), A. kilombero Larsen & Congdon, Zenonia zeno (Trimen), Pelopidas mathias mathias (Fabricius), P. thrax (Hübner), Borbo borbonica borbonica (Boisduval), B. fatuellus fatuellus (Hopffer), B. lugens (Hopffer) and Gegenes niso brevicornis (Plötz).  Distinctive features of the pupa of Ampittia spp. (Aeromachini) are noted.  Amongst Baorini, the pupae of Pelopidas spp., Borbo spp., Gegenes spp. and Zenonia spp. are all similar, being elongate, green, with a frontal spike, while the pupae of Parnara spp. are brown, rounded, and have no frontal spike.  We hypothesise that the pupae of the first four genera reflect a common need for camouflage in an open or partially open shelter made from a single leaf of grass, while the pupa of Parnara spp. are formed in a closed shelter using several leaves as needed.  This may also explain the similarity of the camouflaged pupae to those of many other genera in other tribes, which also feed on grasses, palms etc., and pupate in partially or completely open situations.

References

  1. Ackery, P.R., Smith, C.R. & Vane-Wright, R.I. (1995) Carcasson’s African Butterflies. An annotated catalogue of the Papilionoidea and Hesperioidea of the Afrotropical Region. CSIRO Publications, Melbourne, Australia, 803 pp.

    Aurivillius, C. (1925) Hesperiidae. In: Seitz, A. (Ed.) The Macrolepidoptera of the World. Vol. 13. The African Rhopalocera. Alfred Kernen Verlag, Stuttgart, Germany, pp 505–588, plates 75–80.

    Bascombe, M.J., Johnston, G. & Bascombe, F.S. (1999) The butterflies of Hong Kong. Academic Press, London, xiii + 422 pp, 222 plates.

    Bell, T.R. (1920) The common butterflies of the plains of India (including those met with in the hill stations of the Bombay presidency). Part XXVI. Journal of the Bombay Natural History Society, 27, 211–227, plate M.

    Bell, T.R. (1925) The common butterflies of the plains of India (including those met with in the hill stations of the Bombay Presidency.). Part XXXV. Journal of the Bombay Natural History Society, 30, 822–837.

    Boisduval, [J.A.] (1833) Faune entomologique de Madagascar, Bourbon et Maurice. Lépidoptères. Libraire Encyclopédique du Roret, Paris, 122 pp, 16 plates.

    Bouriquet, G. (1949) Les maladies cryptogamiques et les principaux ennemis végétaux et animaux du riz à Madagascar. Agronomie tropicale, 4(1–2), 81–89.

    Box, H.E. (1953) List of Sugar-Cane Insects. Commonwealth Institute of Entomology, London, 101 pp.

    Brenière, J. (1976) Reconnaissance des principaux lepidopteres du riz de l'Afrique de l'ouest. Agronomie Tropicale, 31(3), 212–231.

    Brévignon, L. & Brévignon, C. (2003) A la Découverte des Papillons du Jour des Antilles Françaises. PLB Editions, Le Gosier, Guadeloupe, 64 pp.

    Brown, F.M. & Heineman, B. (1972) Jamaica and its Butterflies. E.W. Classey Ltd., London, 478 pp.

    Burns, J.M. (1964) Evolution in skipper butterflies of the genus Erynnis. University of California Publications in Entomology, 37, 1–217.

    Burns, J.M. (1989) Phylogeny and zoogeography of the bigger and better genus Atalopedes (Hesperiidae). Journal of the Lepidopterists' Society, 43, 11–32.

    Burns, J.M., Janzen, D.H., Hajibabaei, M., Hallwachs, W. & Hebert, P.D.N. (2008) DNA barcodes and cryptic species of skipper butterflies in the genus Perichares in Area de Conservación Guanacaste, Costa Rica. Proceedings of the National Academy of Sciences of the United States of America, 105, 6350–6355.

    Burns, J.M., Janzen, D.H., Hallwachs, W., Hajibabaei, M. & Hebert, P.D.N. (2009) Genitalia, DNA barcodes, and life histories synonymize Telles with Thracides—a genus in which Telles arcalaus looks out of place (Hesperiidae: Hesperiinae). Journal of the Lepidopterists’ Society, 63, 141–153.

    Butler, A.G. [1870] Catalogue of Diurnal Lepidoptera Described by Fabricius in the Collection of the British Museum. Taylor and Francis, London, v + 303 pp., 3 pls. ("1869").

    CABI (2011) Crop Pest Compendium. http://www.cabi.org/cpc/ (Last accessed 11 Nov 2011).

    Carnegie, A.J.M., Dick, J. & Harris, R.H.G. (1974) Insects and nematodes of South African sugarcane. Entomology Memoir, Department of Agriculture and Technical Services, Republic of South Africa, 39, [2+] 19 pp.

    Carpenter, G.D.H. (1935) The Rhopalocera of Abyssinia: a faunistic study. Transactions of the Entomological Society of London, 83, 313–440. [This includes two new genera descriptions by W.H. Evans].

    Carvalho, M.U.P. & Cardoso, H.L. (1971) Lista de pragas de Angola repeitante ao ano de 1970. Instituto de Investigação Agronómica de Angola, Série Técnica, 25, 15 pp. [The date on the cover is 1971, but in the footers on all pages is 1972].

    Chiba, H. & Eliot, J.N. (1991) A revision of the genus Parnara (Lepidoptera: Hesperiidae), with special reference to the Asian species. Tyô to Ga, 42, 179–194.

    Clark, G.C. (1940) On the life-histories of some South African Lepidoptera. Journal of the Entomological Society of Southern Africa, 3, 42–56, plates I–VIII.

    Cock, M.J.W. (2001) Notes on butterflies seen or collected on a short visit to Nevis, W.I., including the life history of Epargyreus zestos Geyer (Hesperiidae). Living World, Journal of the Trinidad and Tobago Field Naturalists' Club, 2001, 27–36.

    Cock, M.J.W. (2003) The skipper butterflies (Hesperiidae) of Trinidad Part 11, Hesperiinae, Genera group O. Living World, Journal of the Trinidad and Tobago Field Naturalists' Club, 2003, 14–48.

    Cock, M.J.W. (2005) The Skipper Butterflies (Hesperiidae) of Trinidad. Part 13, Hesperiinae, Genera group K. Living World, Journal of the Trinidad and Tobago Field Naturalists' Club, 2005, 23–47.

    Cock, M.J.W. [2010a] Observations on the biology of Pelopidas thrax (Hübner) (Lepidoptera: Hesperiidae: Hesperiinae) in the Hajar Mountains, Oman. Tribulus, 18 (2009), 42–49.

    Cock, M.J.W. (2010b) Observations on the biology of Afro–tropical Hesperiidae (Lepidoptera) principally from Kenya. Part 1. Introduction and Coeliadinae. Zootaxa, 2547, 1–63.

    Cock, M.J.W. (2012, in press) The skipper butterflies (Hesperiidae) of Trinidad. Part 19. Hesperiinae, Moncini: the remaining genera with pale spots: Cymaenes, Cobalopsis, Arita, Lerema, Morys, Tigasis. Living World, Journal of the Trinidad and Tobago Field Naturalists' Club, 2012, 20–40.

    Cock, M.J.W. & Congdon, T.C.E. (2011a) Observations on the biology of Afro–tropical Hesperiidae (Lepidoptera) principally from Kenya. Part 2. Pyrginae: Tagiadini. Zootaxa, 2893, 1–66.

    Cock, M.J.W. & Congdon, T.C.E. (2011b) Observations on the biology of Afrotropical Hesperiidae (Lepidoptera) principally from Kenya. Part 3. Pyrginae: Celaenorrhinini. Zootaxa, 3033, 1–67.

    Common, I.F.B. & Waterhouse, D.F. (1981) Butterflies of Australia. Revised edition. Angus & Robertson Publishers, London, xiv + 682 pp.

    Congdon, C., Bampton, I. & Collins, S. (2008) Early stages and larval shelters of some Platylesches Holland (Lepidoptera: Hesperiidae). Metamorphosis, 19, 160–164.

    Crawford, J.C. (1911) Descriptions of new Hymenoptera. 2. Proceedings of the United States National Museum, 40, 439–449.

    Crawford, J.C. (1913) Descriptions of new Hymenoptera, no. 6. Proceedings of the United States National Museum, 45, 241–260.

    Davidson, J., Bo, C.S. & Aitken, E.H. (1890) Notes on the larvae and pupae of some butterflies of the Bombay Presidency. Journal of the Bombay Natural History Society, 5, 260–278, plates A–C, 349–375, plates D–F.

    Davidson, J., Bell, T.R. & Aitken, E.H. (1897) The butterflies of the North Canara District of the Bombay Presidency. Part IV. Journal of the Bombay Natural History Society, 11, 22–63, plates 7–8.

    Davis, P.M.H. & Barnes, M.J.C. (1993) The butterflies of Mauritius. Journal of Research on the Lepidoptera, 30 (1991), 145–161.

    De Jong, R. (1982) Secondary sexual characters in Celaenorrhinus and the delimitation of the genus (Lepidoptera, Hesperiidae). Journal of Natural History, 16, 695–705.

    Descamps, M. (1956) Insectes nuisibles au riz dans le Nord Cameroun. Agronomie tropicale, 11(6), 732–755.

    Dethier, V.G. (1939) Metamorphoses of Cuban Hesperiinae. Psyche, 46, 147–155.

    Dickson, C.G.C. & Kroon, D.M. (Eds.) (1978) Pennington’s Butterflies of Southern Africa. A.D. Donker, Johannesburg, South Africa, 670 pp.

    Doherty, W. (1886) A list of butterflies taken in Kumaon. Journal of the Asiatic Society of Bengal, 55 (Part II) (2), 103–140.

    Eliot, J.N. (1978) The Butterflies of the Malay Peninsula by A. Steven Corbet and H. M. Pendlebury. Third edition. Malayan Nature Society, Kuala Lumpur, Malaysia, xiv + 578 pp, 35 plates.

    Evans, W.H. (1935) see Carpenter (1935).

    Evans, W.H. (1937a) A catalogue of the African Hesperiidae indicating the classification and nomenclature adopted in the British Museum. British Museum (Natural History), London, UK, 212 pp, 30 plates.

    Evans, W.H. (1937b) Indo-Australian Hesperiidae: descriptions of new genera, species and subspecies. Entomologist, 70, 16–19, 38–40, 64–66, 81–83.

    Evans, W.H. (1947) Revisional notes on African Hesperiidae. Annals and Magazine of Natural History, Series 11, 13, 641–648.

    Evans, W.H. (1949) A catalogue of the Hesperiidae from Europe, Asia and Australia in the British Museum (Natural History). British Museum (Natural History), London, 502 pp, 53 plates.

    Evans, W.H. (1951) Revisional notes on African Hesperiidae. Annals and Magazine of Natural History, Series 12, 4, 1268–1272.

    Fabricius, J.C. (1798) Supplementum. Entomologiae systematicae. Proft et Storch, Hafniae [=Denmark], [iv] + 572 pp.

    Feulner, G.R. & Karki, N. (2010) Hidden in plain view: First UAE record of the wadi grass Saccharum kajkaiense and notes on its distribution in the UAE and neighbouring Oman. Tribulus, 18 (2009), 50–55.

    Fleming, W.A. (1983) Butterflies of West Malaysia and Singapore, 2nd edition. Longman Malaysia, Kuala Lumpur, Malaysia, 148 pp, 92 plates.

    Fletcher, T.B. (1914) Some South Indian Insects and Other Animals of Importance considered especially from an economic point of view. Superintendent, Government Press, Madras, India, 565 pp.

    Fletcher, T.B. (1920) Annotated list of Indian crop pests. In: Fletcher, T.B. (Ed.) Report of the proceedings of the Third Entomological meeting held at Pusa on the 3rd to 15th February 1919. Volume 1. Superintendent of Government Printing, Calcutta, India, pp 33–313.

    Forsyth, J. (1966) Agricultural Insects of Ghana. Ghana Universities Press, Accra, Ghana, 163 pp.

    Frappa, C. (1937) Note sur deux espéces de Pamphila nuisibles au riz á Madagascar. Riz et Riziculture, 11(1), 19–24.

    Gaede, M. (1916) Neu afrikanische Lepidoptera des Berliner Zoologischen Museums. Internationale Entomologische Zeitschrift, 9, 105–106, 109–112, 125–126.

    Gaskin, J.F., Bon, M.-C., Cock, M.J.W., Cristofaro, M., De Biase, A., De Clerck-Floate, R., Ellison, C.A., Hinz, H.L., Hufbauer, R.A., Julien, M. & Sforza, R. (2011) Applying molecular-based approaches to classical biological control of weeds. Biological Control, 58, 1–21.

    Ghosh, C.C. (1914) Life-histories of Indian insects. V. Lepidoptera. (Butterflies). Memoirs of the Department of Agriculture in India, Entomology Series, 5(1), 1–72, plates I–IX.

    Ghosh, C.C. (1940) Insect pests of Burma. Supdt., Govt. Printing and Stationery, Burma; Rangoon, Burma, ii + 216 pp.

    Gifford, D. (1965) A list of the Butterflies of Malawi. The Society of Malawi (Historical and Scientific), Blantyre, Malawi, 151 pp.

    Grist, D.H. & Lever, R.J.A.W. (1969) Pests of Rice. Longmans, London, 520 pp.

    Guillermet, C. (2011) Les papillons diurnes de La Réunion. http://christian.guillermet.perso.neuf.fr/rhopalo.htm (Last accessed 6 Nov 2011).

    Hargreaves, H. (1932) Annual Report of the Government Entomologist. Annual Report of the Department of Agriculture of Uganda, 1931(ii), 43–47. [As abstracted in Review of Applied Entomology]

    Hargreaves, H. (1939) Notes on some pests of maize and millets in Uganda. East African Agricultural Journal, 5, 104–109.

    Hargreaves, H. (1940) Insect pests of sugar-cane. In: Tothill, J.D. (Ed.) Agriculture in Uganda. Oxford University Press, London, pp 395–396.

    Heath, A., Newport, M.A. & Hancock, D. (2002) The Butterflies of Zambia. African Butterfly Research Institute and The Lep-idopterists’ Society of Africa, Nairobi, Kenya, 137 pp. + CD.

    Heinrichs, E.A. & Barrion, A.T. (2004) Rice-feeding insects and selected natural enemies in West Africa: Biology, ecology, identification. International Rice Research Institute, Los Baños, Philippines; and WARDA–The Africa Rice Center, Abidjan, Côte d’Ivoire, 243 pp.

    Henning, G.A., Henning, S.F., Joannou, J.G. & Woodhall, S.E. (1997) Living Butterflies of Southern Africa. Biology, Ecology and Conservation. Volume I Hesperiidae, Papilionidae and Pieridae of South Africa. Umdaus Press, Hatfield, South Africa, 395 pp.

    Hewitson, W.C. (1868) Descriptions of one hundred new species of Hesperidae. John Van Voorst, London, vol. 2, 25–56.

    Higgins, L.G. (1975) The Classification of European Butterflies. Collins, London, 320 pp.

    Hoffmannsegg, J.C.G. (1804) Alphabetisches Verseichniss zu J. Hübner’s Abbildungen der Papilionen mit den beigefügten vorzüglichsten Synonymen. Magazin für Insektenkunde (Illiger), 4, 181–206.

    Holland, W.J. (1896) A preliminary revision and synonymic catalogue of the Hesperiidae of Africa and the adjacent islands, with descriptions of some apparently new species. Proceedings of the Zoological Society of London, 1896, 2–107, plates I–V.

    Hopffer, [C.] (1855) Lepidoptera diurna. In: Peters, [W.C.H.] Reisewerke über Mossambique. Bericht über die zur Bekanntmachung geeigneten Verhandlungen der Königlich-Preussischen Akademie der Wissenschaften zu Berlin, 1855, 636–660.

    Hübner, J. [1819–1826] Zuträge zur Sammlung exotischer Schmettlinge [sic]: bestehend in Bekundigung einzelner Fliegmuster neuer oder rarer nichteuropäischer Gattungen. Vol 2. The author, Augsburg, Germany, unpaginated.

    IRRI (International Rice Research Institute) (1983) Field problems of tropical rice. Revised Edition. International Rice Research Institute, Los Baños, Philippines, 172 pp.

    Jandu, A.S. (1943) Biological notes on the butterflies of Delhi. Part II. Danaidae, Satyridae, Nymphalidae, Lycaenidae and Hesperiidae. Indian Journal of Entomology, 5, 223–241.

    Janzen, D.H. & Hallwachs, W. (2011) Dynamic database for an inventory of the macrocaterpillar fauna, and its food plants and parasitoids, of Area de Conservacion Guanacaste (ACG), northwestern Costa Rica. http://janzen.sas.upenn.edu (Last accessed 11 Nov 2011).

    Kalshoven, L.G.E. & van den Laan, P.A. (1981) The Pests of Crops in Indonesia. P.T. Ichtiar Baru-Van Hoeve, Jakarta, Indonesia, 701 pp.

    Kerrich, G.J. (1973) A revision of the tropical and subtropical species of the eulophid genus Pediobius Walker (Hymenoptera: Chalcidoidea). Bulletin of the British Museum (Natural History) (Entomology), 29, 113–199.

    Kielland, J. (1990) Butterflies of Tanzania. Hill House, Melbourne, Australia, 363 pp.

    Kroon, D.M. (1973) Notes on southern African Rhopalocera (Lepidoptera). Entomologists' Record and Journal of Variation, 85, 57–61, plates 1–5.

    Larsen, T.B. (1982) Gegenes pumilio Hoffmannsegg, 1804; a review with cytological evidence that two species are involved (Hesperiidae). Nota Lepidopterologica, 5 (2–3), 103–110.

    Larsen, T.B. (1984) Butterflies of Saudi Arabia and its neighbours. Transworld Arabian Library, Riyadh & Stacey International, London, 160 pp.

    Larsen, T.B. (1991) The Butterflies of Kenya and their Natural History. Oxford University Press, Oxford, UK, 490 pp.

    Larsen, T.B. (2002) The butterflies of Delhi, India – an annotated check-list. Esperiana, 9, 459–479.

    Larsen, T.B. (2005) Butterflies of West Africa. 2 vols. Apollo Books, Stenstrup, Denmark, 595 + 270 pp.

    Larsen, T.C.E. & Congdon, T.C.E. (2012) The genus Ampittia in Africa with the description of a new species (Hesperiinae; Aeromachini) and three new species in the genera Andronymus and Chondrolepis (Hesperiinae, incertae sedis) (Lepidoptera; Hesperiidae). Zootaxa, 3322, 49–62.

    Latreille, P.A. (1809) Genera crustaceorum et insectorum secundum ordinem naturalem in familias disposita, iconibus, exemplisque plurimis _tal._ate. Tomus quartus et ultimus. Amand König, Parisiis et Argentorati, 399 pp.

    Latreille, P.A. & Godart, J.B. [1819, 1824] Encyclopédie Méthodique. Histoire Naturelle [Zoologie] 9 Entomologie. Veuve Agasse, Paris, i–iv, 1–328 [1819], 329–828 [1824].

    Le Pelley, R.H. (1959) Agricultural insects of East Africa. East Africa High Commission, Nairobi, Kenya, 307 pp.

    Linnaeus, C. (1764) Museum S:ae R:ae M:tis Ludovicae Ulricae Reginae Svecorum, Gothorum, Vandalorumque &c. &c. &c. In quo animalia rariora, exotica, imprimis Insecta & Conchilia describuntur & determinantur. Prodromi instar editum. Laurentius Salvius, Holmiae, [=Stockholm, Sweden], [viii] + 720 + [2] pp.

    Litsinger, J.A., Barrion, A.T. & Soekarna, D. (1987) Upland rice insect pests: their ecology, importance and control. IRRI Research Paper Series, 123, 41 pp.

    Litsinger, J.A., Bumroongsri, V., Morrill, W.L. & Sarnthoy, O. (1994) Rearing, developmental biology and host plant range of the rice skipper Pelopidas mathias (F.) (Lepidoptera: Hesperiidae). Insect Science and its Application, 15, 9–17.

    Mabille, P. (1884) Descriptions de Lépidoptères exotiques. Bulletin ou Comptes Rendus des Séances de la Societe Entomologique de Belgique, 28, clxxxiv–cxci [184–191].

    Makris, C. (2003) Butterflies of Cyprus. Bank of Cyprus Cultural Foundation, Nicosia, Cyprus, 329 pp.

    Mansfield-Aders, W. (1920) Insects injurious to economic crops in the Zanzibar Protectorate. Bulletin of Entomological Research , 10, 145–155, plates XIII–XV.

    Maxwell-Lefroy, H. (1907) The more important insects injurious to Indian Agriculture. Memoirs of the Department of Agriculture in India, Entomology Series, 1(2), 113–253.

    Maxwell-Lefroy, H. & Howlett, F.M. (1909) Indian Insect Life. A Manual of the Insects of the Plains (Tropical India). Thacker Spink and Co., Calcutta and Simla, India, 786 pp.

    Migdoll, I. (1988) Ivor Migdoll's Field Guide to the Butterflies of Southern Africa. New Holland Press, London, 256 pp.

    Miller, L.D. & Miller, J.Y. (1997) Gondwanan butterflies: the Africa-South American connection. The Inaugural Conference on African Lepidoptera. Nairobi – Kenya – 1–8 May 1977. Metamorphosis, occasional supplement no. 3, 42–51.

    Minno, M.C., Butler, J.F. & Hall, D.W. (2005) Florida butterfly caterpillars and their host plants. University Press of Florida, Gainesville, Florida, xi + 341 pp.

    Moore, F. (1880–1881) Lepidoptera of Ceylon. Vol. 1. L. Reeve and Co., London, 190 pp.

    Murray, D. (1959) The genitalia and food-plants of the South African Hesperiidae (Lepidoptera: Rhopalocera). Durban Museum Novitates, 5, 219–227, plates i–iii.

    Nakasuji, F. (1982) Seasonal changes in native host plants of a migrant skipper, Parnara guttata Bremer & Grey (Lepidoptera: Hesperiidae). Journal of Applied Entomology and Zoology, 17(1), 146–148.

    Odak, S.C. & Dhamdhere, S.V. (1970) Record of a new alternate host plant of Parnara mathias Fabricius. Indian Journal of Entomology, 32(4), 397.

    Pinhey, E.C.G. (1949) Butterflies of Rhodesia with a short introduction to the insect world. Rhodesia Scientific Association, Salisbury, 208 pp.

    Pinhey, E. (1965) Butterflies of Southern Africa. Thomas Nelson & Sons (Africa) Ltd., Johannesburg, South Africa, 240 pp.

    Plötz, C. (1883) Die Hesperiinen-Gattung Hesperia Aut. und ihre Arten. [part] Entomologische Zeitung, 44, 26–64.

    Plötz, C. (1884) Die Hesperiinen-Gattung Thymelicus Hüb. und ihre Arten. Entomologische Zeitung, 45, 284–290.

    Pringle, E.L.L., Henning, G.A. & Ball, J.B. (Eds.) (1994) Pennington’s Butterflies of Southern Africa. Second Edition. Struik Winchester, Cape Town, South Africa, 800 pp.

    Pro Natura – Ligue Suisse pour la Protection de la Nature (Ed.) (1999) Les papillons et leurs biotopes. Espèces ·Dangers qui les menacent · Protection. Suisse et regions limitrophes. Tome 2. Fotorotar SA, Egg, Switzerland, xi + 667 pp.

    Purohit, M.L., Patel, R.K. & Kharti, A.K. (1972) Note on the record of a new host of the rice skipper, Parnara mathias Fabricius (Lepidoptera: Hesperiidae). Indian Journal of Agricultural Science, 42, 429–430.

    Raich, K.V. & Choudhary, B.S. (1974) A new alternate host plant of the rice skipper, Parnara mathias Fabricius (Lepidoptera: Hesperiidae). Madras Agricultural Journal, 61, 234.

    Robinson, G.S., Ackery, P.R., Kitching, I.J., Beccaloni, G.W. & Hernández, L.M. (2001) Hostplants of the moth and butterfly caterpillars of the Oriental Region. The Natural History Museum, London, 744 pp.

    Sen, P. & Chakravorty, S. (1970) Mode of formation of larval shelters in certain lepidopterous pests of rice. International Rice Commission Newsletter, 19(2), 13–19.

    Sevastopulo, D.G. (1973) The food plants of Indian Rhopalocera. Journal of the Bombay Natural History Society, 72(3), 156–183.

    Sevastopulo, D.G. (1974). Butterflies of the Shimba Hills. Entomologist's Record and Journal of Variation, 86, 131–136.

    Sevastopulo, D.G. (1975) Hesperiidae. Bulletin of the Amateur Entomologists' Society, 34, 131–132. In: Sevastopulo, D.G. (1975–1978) A list of the food-plants of East African Macrolepidoptera. Bulletin of the Amateur Entomologists' Society, 34, 84–92, 124–132, 175–184; 35, 94–100, 177–195; 36, 45–50; 37, 147–172, 202–216.

    Sevastopulo, D.G. (1981) A second list of the food-plants of East African Macrolepidoptera. Bulletin of the Amateur Entomologists' Society, 40, 48–55.

    Sevastopulo, D.G. (unpublished) Manuscript and Photograph Collection of Demetrius George Sevastopulo (1903–1987). Entomology Library, Natural History Museum, London, UK. [See http://www.nhm.ac.uk/research-curation/collections/collections-management/collections-navigator/index.jsp for details].

    Shroff, K.D. (1920) A list of the pests of cereals in Burma. In: Fletcher, T.B. (Ed.) Report of the proceedings of the Third Entomological meeting held at Pusa on the 3rd to 15th February 1919. Volume 1. Superintendent of government Printing, Calcutta, India, pp. 341–343.

    Teotia, T.P.S. & Nand, S. (1966) Bionomics of the rice skipper, Parnara mathias Fabricius (Lepidoptera: Hesperiidae). Indian Journal of Entomology, 28(2), 181–186.

    Trimen, R. (1864) On some new species of South African butterflies. Transactions of the Entomological Society of London, Series 3, 2, 175–180.

    Trimen, R. (1889) South–African Butterflies: a Monograph of the Extra–Tropical Species. Vol. III. Papilionidae and Hesperidae. Trübner & Co., Ludgate Hill, London, 438 pp, plates 10–12.

    Trimen, R. (1893) On some new or imperfectly known species of butterflies from South Africa. Transactions of the Entomological Society of London, 1893, 123–143, pl. 8.

    Tutt, J.W. (1906) Catalogue of the Palaearctic Urbicolides. Entomologist's Record and Journal of Variation, 18, 195–198.

    Van Someren, V.G.L. (1939) Butterflies of the Chyulu Range. A systematic list of the species taken by the Museum Expedition to the hills. April–July 1938. Journal of the East Africa and Uganda Natural History Society, 14, 130–151.

    Van Someren, V.G.L. (1974) List of foodplants of some East African Rhopalocera, with notes on the early stages of some Lycaenidae. Journal of the Lepidopterists' Society, 28, 315–331.

    van Vreden, G. & Ahmadzabidi, A.L. (1986) Pests of rice and their natural enemies in Peninsular Malaysia. PUDOC, Wageningen, Netherlands, 230 pp.

    Vuattoux, R. (1999) Les lepidopteres hesperiides de la Station de Lamto (Côte d’Ivoire). Lambillionea, 94, 562–568.

    Walker, F. (1870) A list of the butterflies collected by J. K. Lord in Egypt, along the African shore of the Red Sea; and in Arabia; with descriptions of the species new to science. The Entomologist, 5(4), 48–57.

    Warren, A.D., Ogawa, J.R. & Brower, A.V.Z. (2008) Phylogenetic relationships of subfamilies and circumscription of tribes in the family Hesperiidae (Lepidoptera: Hesperioidea). Cladistics, 24, 1–35.

    Warren, A.D., Ogawa, J.R. & Brower, A.V.Z. (2009) Revised classification of the family Hesperiidae (Lepidoptera: Hesperioidea) based on combined molecular and morphological data. Systematic Entomology, 34, 467–523.

    Williams, J.R. (1989) Butterflies of Mauritius. The author, Mauritius, 41 pp.

    Veenakumari, K., Mohanraj, P. & Sreekumar, P.V. (1997) Host plant utilization by butterfly larvae in the Andaman and Nicobar Islands (Indian Ocean). Journal of Insect Conservation, 1, 235–246.

    Vishakantaiah, M. & Visweswara Gowda, B.L. (1975) A list of insect pests infesting barley and oats in Karnataka. Madras Agricultural Journal, 62, 91–93.