Abstract
The Oxytelus Gravenhorst, 1802 is one of the most speciose genus in the subfamily Oxytelinae (Coleoptera:Staphylinidae) and its members are mainly ground dwellers and are distributed in almost all geographical regions.This paper reports four new species from China: O. abiturus sp. nov. from Sichuan and Yunnan, O. ailaoshanicussp. nov. from Shaanxi, Sichuan, and Yunnan, O. bajiei sp. nov. from Guangxi, Hubei, Hunan, Sichuan, Yunnan, andO. solus sp. nov. from Sichuan. The following four names are synonymized: O. ginyuenensis Bernhauer, 1935 andO. takahashii Ito, 1994 (= O. lucidulus Cameron, 1929); O. discalis Cameron, 1930 (= O. punctipennis Fauvel,1905); O. nilgiriensis Cameron, 1930 (= O. subferrugineus Cameron, 1941). The Chinese fauna of the genus is thusincreased to twenty-one species. All other species known to occur in China are also revised and redescribed to updatethe knowledge on their identifications. Some important characters are discussed, detailed line drawing is offered foreach species, and a key to all the Chinese species is provided. All the types of the new species are deposited in IZCAS.key words: Staphylinidae, Oxytelinae, Oxytelus, Taxonomy, ChinaReferences
Abdullah, M. & Qadri, N.-N. (1970) The Staphylindiae [sic], Coleoptera of Pakistan. Part III. A key to the genera and species of the Piestinae, Osoriinae, Pseudopsinae and Oxytelinae, with descriptions of new genera, subgenera and species from Karachi. Pakistan Journal of Scientific and Industrial Research, 13, 114–131.
Ades, G.W.J. & Kendrick, R.C. (2004) Hong Kong Fauna: a Checklist of Selected Taxa. Kadoorie Farm & Botanic Garden Corporation, Hong Kong, 84 pp.
Allen, A.J. (2003) Oxytelus migrator Fauvel (Staphylinidae) in Wiltshire. Coleopterist, 12, 40.
Bernhauer, M. (1903) Beitrag zur Staphylinidenfauna von Ostindien und den Sundainseln. Stettiner Entomologische Zeitung, 64, 21–36.
Bernhauer, M. (1907) Zur Staphyliniden-fauna von Japan. Verhandlungen der kaiserlich-königlichen Zoologisch-Botanischen Gesellschaft in Wien, 57, 371–414.
Bernhauer, M. (1910) Beitrag zur Kenntnis der Staphyliniden-Fauna von Zentralamerika. Verhandlungen der kaiserlich-königlichen Zoologisch-Botanischen Gesellschaft in Wien, 60, 350–393.
Bernhauer, M. (1912) Neue Staphyliniden der Philippinen. The Philippine Journal of Science, 7, 245–254.
Bernhauer, M. (1922) Sauter’s Formosa-Ausbeute: Stapliylinidae I. Archiv für Naturgeschichte. Abteilung A, 88, 220–237.
Bernhauer, M. (1933) Neuheiten der chinesischen Staphylinidenfauna. Wiener Entomologische Zeitung, 50, 25–48.
Bernhauer, M. (1935) Neue Kurzflügler aus China. Entomologisches Nachrichtenblatt, Troppau, 9, 4–14.
Bernhauer, M. & Schubert, K. (1911) Staphylinidae II (Pars 29). In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. vol. V. W. Junk, Berlin, pp. 87–190.
Biswas, D.N. (2003) Insecta: Coleoptera: Staphylinidae. In: Zoological Survey of India (Ed.), Fauna of Sikkim [State Fauna Series 9]. Zoological Survey of India, Kolkata, pp. 237–281.
Blackburn, T. (1885) [New species]. In: Blackburn, T. & Sharp, D. (Eds.), Memoirs on the Coleoptera of the Hawaiian Islands. vol. 3(2). The Scientific Transactions of the Royal Dublin Society, pp. 119–300.
Blackwelder, R.E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. United States National Museum Bulletin, 182, i–viii, 1–658.
Blackwelder, R.E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. United States National Museum Bulletin, 200, i–iv, 1–483.
Cameron, M. (1919) New species of Staphylinidae from Ceylon. Part I. The Entomologist’s Monthly Magazine, 55, 224–228, 251–255.
Cameron, M. (1928) Fauna sumatrensis. Staphylinidae (Col.). Entomologische Mitteilungen, Berlin. 17, 90–110.
Cameron, M. (1929) New Staphylinidae from the Malay Peninsula. Journal of the Federated Malay States Museums, Kuala Lumpur, 14, 436–452.
Cameron, M. (1930) The fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. I. Taylor and Francis, London, xvii+471 pp.
Cameron, M. (1933) New species of Staphylinidae (Col.) from the Belgian Congo. Bulletin et Annales de la Société Entomologique de Belgique, 73, 35–53.
Cameron, M. (1934) Further remarks on some of Motschulsky’s types of oriental Staphylinidae (Col.). The Entomologist’s Monthly Magazine, 70, 76–80.
Cameron, M. (1936) Fauna Javanica The Staphylinidae (Col.) collected by Mr. F. C. Drescher. Tijdschrift voor Entomologie Amsterdam, 79, 25–54.
Cameron, M. (1938) New species of Staphylinidae from the Belgian Congo. Bulletin et Annales de la Société Entomologique de Belgique, 14, 1–16.
Cameron, M. (1941) New species of Staphylinidae (Col.) from the Philippines. Annals and Magazine of Natural History, (11)7, 430–447.
Cameron, M. (1949) New species and records of staphylinid beetles from Formosa, Japan, and South China. Proceedings of the United States National Museum, 99, 455–477.
Coiffait, H. (1983) Nouveaux staphylinides du Cachemire (Coleoptera). Annales de la Société entomologique de France (n. ser.), 19, 273–289.
Erichson, W.F. (1840) Genera et Species Staphylinorum Insectorum Coleopterorum Familiae. F. H. Morin, Berlin, pp. 401–954.
Fabricius, J.C. (1775) Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, adiectis Synonymis, Locis, Descriptionibus, Observationibus. Libraria Kortii, Flensburgi et Lipsiae, xxxi+832 pp.
Fagel, G. (1956) Contribution à la connaissance des Staphylinidae. XXXVIII. Démembrement du genre Oxytelus Erichson. Bulletin et Annales de la Société Royale d’Entomologie de Belgique, 92, 267–275.
Fagel, G. (1957) Contributions à l’étude de la faune entomologique du Ruanda-Urundi (Mission P. Balilewsky 1953). CXIX. Coleoptera Staphylinidae: Piestinae, Omaliinae, Proteininae, Osoriinae, Oxytelinae. Annales du Musée Royal du Congo belge, Tervuren (ser. in 8). Sciences Zoologiques, 58, 26–72.
Fagel, G. (1960) Contribution à la connaissance des Staphylinidae. LXVII. Le complexe des Anisopsis. Bulletin Institut royal des Sciences naturelles de Belgique, 36, 1–51.
Fairmaire, L. (1893) Coléoptères des Iles Comores. Annales de la Société Entomologique de Belgique, 37, 521–555.
Fauvel, A. (1875a) Faune Gallo-Rhénane ou species des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, la Prusse Rhénane, le Nassau et le Valais avec tableaux synoptiques et planches gravées. Coléoptères. Catalogue systématique des staphylinides de la Faune Gallo-Rhénane avec l’addition synonymique des espèces européennes, siberiénnes, caucasiques et Mediterranéennes et descriptions nouvelles. Bulletin de la Société Linnéenne de Normandie, (2)9, 202–236.
Fauvel, A. (1875b) Faune Gallo-Rhénane ou species des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, la Prusse Rhénane, le Nassau et le Valais avec tableaux synoptiques et planches gravées. Coléoptères. Catalogue systématique des staphylinides de la Faune Gallo-Rhénane avec l’addition synonymique des espèces européennes, siberiénnes, caucasiques et Mediterranéennes et descriptions nouvelles. vol. 3, livraison 6. F. le Blanc-Hardel, Caen, xxxviii+4+228–231 pp.
Fauvel, A. (1876a) Faune Gallo-Rhénane ou species des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, la prusse Rhénane, la Nassau et la Valais avec tableaux synoptiques et planches gravées. 3e supplément aux staphylinides. vol. 3, livraison 6. F. le Blanc-Hardel, Caen, pp. 47–82.
Fauvel, A. (1876b) Faune Gallo-Rhénane ou species des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, la prusse Rhénane, la Nassau et la Valais avec tableaux synoptiques et planches gravées. Bulletin de la Société Linnéenne de Normandie, (2)10, 26–267.
Fauvel, A. (1895) Staphylinides nouveaux de l’Inde et de la Malaisie. Revue d’Entomologie publiée par la Société Française d’Entomolodie. Caen, 14, 180–286.
Fauvel, A. (1901) Voyage de M. le Dr. Ed. Bugnion au Venezuela, en Colombie et aux Antilles. Revue d’Entomologie publiée par la Société Française d’Entomolodie. Caen, 20, 69–91.
Fauvel, A. (1902) Catalogue des Staphylinides de la Barbarie de la Basse-Égypte et des Iles Açores, Madères, Salvages et Canaries. Revue d’Entomologie publiée par la Société Française d’Entomolodie. Caen, 21, 45–189.
Fauvel, A. (1904a) Staphylinides de l’Hindoustan et de la Birmanie. Revue d’Entomologie publiée par la Société Française d’Entomolodie. Caen., 23, 43–70.
Fauvel, A. (1904b) Staphylinides exotiques nouveaux. 2e Partie. Revue d’Entomologie publiée par la Société Française d’Entomolodie. Caen, 23, 76–112.
Fauvel, A. (1905) Staphylinides exotiques nouveaux. 3e Partie. Revue d’Entomologie publiée par la Société Française d’Entomolodie. Caen, 24, 113–147.
Fauvel, A. (1907) Voyage de M. Ch. Alluaud dans l’Afrique Orientale. Staphylinidae. Revue d’Entomologie publiée par la Société Française d’Entomolodie. Caen, 26, 10–70.
Gaedike, H. (1981) Katalog der in den Sammlungen der Abteilung Taxonomie der Insekten des Institutes für Pflanzenschutzforschung, Bereich Eberswalde (ehemals Deutsches Entomologisches Institut), aufbewahrten Typen — XIX. Beiträge zur Entomologie, Berlin, 31, 175–232.
Ganglbauer, L. (1895) Die Käfer von Mitteleuropa. Die Käfer der österreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des französischen und italienischen Alpengebietes. 2. Familienreihe Staphylinoidea. Theil I. Staphylinidae, Pselaphidae. Carl Gerold’s Sohn, Wien, 881 pp.
Gebler, F.A. von (1848) Verzeichniss der im Kolywano-Woskresenskischen Hüttenbezirke Süd-West Sibiriens beobachteten Kaefer mit Bemerkungen und Beschreibungen. Bulletin de la Société Impériale des Naturalistes de Moscou. 21, 3–85.
Gemminger, M. & Harold, E. von (1868) Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. vol. 2. E. H. Gummi, Monachii, pp. 425–752+vi.
Gollkowski, V. (2007) Oxytelus migrator Fauvel, 1904 (Coleoptera, Staphylinidae) in Portugal. Entomologische Nachrichten und Berichte (Faunistische Notiz Nr. 867), 51, 134.
Gravenhorst, J.L.C. (1802) Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera familias et species distribuit. vol. 1. Carolus Reichard, Brunsuigae, lxvi+206 pp.
Gravenhorst, J.L.C. (1806) Monographia Coleopterorum Micropterorum. Henricus Dieterich, Gottingae, 248 pp.
Gusarov, V.I. (1991) New and little-known Palearctic Staphylinidae (Coleoptera). Communication 2 [Новые и Малоизвестные Палеарктические стафилиниды (Coleoptera, Staphylinidae). Сообщение 2]. Bulletin of the Leningrad University. Biology [Вестник Ленинградского университета: Биология], 1991, 3–12, 133.
Hammond, P.M. (1975) Report from the Lund University Ceylon Expedition in 1962. Report No. 34. Coleoptera: Staphylinidae Oxytelini from Ceylon. Entomologica scandinavica, Supplementum. 4, 141–178.
Hammond, P.M. (1976a) A review of the genus Anotylus C.G. Thomson (Coleoptera: Staphylinidae). Bulletin of the British Museum (Natural History) (Entomology), 33, 139–187.
Hammond, P.M. (1976b) The crassicornis species-group of the genus Anotylus (Staphylinidae). Systematic Entomology, 1, 19–51.
Hammond, P.M. (1998) Oxytelus migrator Fauvel (Col., Staphylinidae), an Asian rove-beetle established in Britain. Entomologist’s Monthly Magazine, 134, 273–276.
Heer, O. (1839) Fauna Coleopterorum Helvetica. vol. 1(2). Orelii, Fuesslini et Sociorum, Turici, pp. 145–360.
Herman, L.H. (1970) Phylogeny and reclassification of the genera of the rove-beetle subfamily Oxytelinae of the world (Coleoptera, Staphylinidae). Bulletin of the American Museum of Natural History, 142, 345–454.
Herman, L.H. (2001) Catalog of the Staphylinidae (Insecta, Coleoptera) : 1758 to the end of the second millennium. Vol. 3, Oxyteline group. Bulletin of the American Museum of Natural History, No. 265, 1067–1806.
Hochhuth, J.H. (1851) Beitraege zur naeheren Kenntniss der Staphylinen Russlands. Enthaltend Beschreibung neuer Genera und Arten, nebst Erläuterungen noch nicht hinlänglich bekannter Staphylinen des russischen Reichs. Bulletin de la Société Impériale des Naturalistes de Moscou, 24,pt.2, 3–58.
Horion, A. (1963) Faunistik der mitteleuropäischen Käfer. Staphylinidae. 1. Micropeplinae bis Euaesthetinae. vol. 9. A. Feyel, Überlingen-Bodensee, xii+412 pp.
Ito, T. (1994) Notes on the species of Staphylinidae (Coleoptera) from Japan 3. Descriptions of two new species of the genus Oxytelus Gravenhorst. Natural History Research, 3, 41–46.
Koch, C. (1932) Eine neue mediterrane Oxytelus-Art. In: Livre du Centenaire. Société Entomologique de France, Paris, pp. 649–656.
Koch, C. (1934) Wissenschaftliche ergebnisse der Entomologische Expeditionen seiner Durchlaucht des Fuersten Alexandro C. della Torre e Tasso nach Aegypten und auf die Halbinsel Sinai. IV. Staphylinidae (Coleoptera). Bulletin de la Société Royale Entomologique d’Égypte, 1934, 33–91.
Kraatz, G. (1859) Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan. Archiv für Naturgeschichte. 25, 1–196.
Latreille, P.A. (1804) Histoire Naturelle, Générale et Particulière, des Crustacés et des Insectes. vol. IX. F. Dufart, Paris, 416 pp.
Latreille, P.A. (1810) Considerations Générales sur l’Ordre Naturel des Animaux Composant les Classes des Crustacés, des Arachnides et des Insectes; Avec un Tableau Méthodique de leurs Genres, Disposés en Familles. F. Schoell, Paris, 444 pp.
Lee, S.-I. & Ahn, K.-J. (2007) The Korean species of the genus Oxytelus Gravenhorst (Coleoptera: Staphylinidae: Oxytelinae). Korean Journal of Systematic Zoology, 23, 21–33.
Linné, C. von (1767) Systema Naturae per Regna Tria Naturae, secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Duodecima Reformata. vol. 1, pt. 2. Laurentii Salvii, Holmiae, pp. 533–1327+xxxvii.
Lott, D.A. (2009) The Staphylinidae (rove beetles) of Britain and Ireland part 5: Scaphidiinae, Piestinae, Oxytelinae. vol. 12. Royal Entomological Society, St Albans, iv+99 pp.
Lucas, R. (1920) Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Archiv für Naturgeschichte (A), 84, 1–696.
Makranczy, G. (2006) Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae). Annales Historico-Naturales Musei Nationalis Hungarici, 98, 29–119.
Makranczy, G. (2012) Nomenclatural changes removing secondary homonymy in the genus Oxytelus (Coleoptera: Staphylinidae: Oxytelinae). Acta Entomologica Musei Nationalis Pragae, 52, 193–205.
Motschulsky, V. de (1857) Énumération des nouvelles espèces de Coléoptères rapportés de ses voyages. Bulletin de la Société Impériale des Naturalistes de Moscou, 30, 490–517.
Motschulsky, V. de (1862) Entomologie spéciale, Insectes du Japan. I. Coleopteres. Etudes Entomologiques. Helsingfors, 9(1860), 4–41.
Mukhopadhyay, P. & Sar, A. (2007) Coleoptera: Staphylinidae. In: Zoological Survey of India (Ed.), Fauna of Andhra Pradesh (Part 3): Insects [State Fauna Series 5]. Zoological Survey of India, Kolkatta, pp. 417–438.
Müller, O.F. (1776) Zoologiae Danicae prodromus, seu animalium Daniae et Norvegiae indigenarum characteres, nomina, et synonyma imprimis popularium. Hallager, Havniae, xxxii+282 pp.
Naomi, S.-I. (1989) Comparative morphology of the Staphylinidae and the allied groups (Coleoptera, Staphylinoidea) X. Eighth to 10th segments of abdomen. Japanese Journal of Entomology, 57, 720–733.
Naomi, S.-I. (1990) Comparative morphology of the Staphylinidae and the allied groups (Coleoptera, Staphylinoidea) XI. Abdominal glands, male genitalia and female spermatheca. Japanese Journal of Entomology, 58, 16–23.
Newton, A.F., Thayer, M.K., Ashe, J.S. & Chandler, D.S. (2000) 22. Staphylinidae. In: Arnett, R.H. & Thomas, M.C. (Eds.), American Beetles. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. vol. 1. CRC Press, Boca Raton, London, New York, and Washington D.C., pp. 272–418.
Normand, H. (1947) Contribution au catalogue des Coléoptères de la Tunisie. Bulletin Société de Géographie et d’Archélogie de la Province d’Oran. Troisième supplément, fascicule 2. 66/67(1945–1946), 1–16.
Olivier, A.G. (1811) Encyclopédie Méthodique. Histoire naturelle. Insectes. vol. 8(2). H. Agasse, Paris, pp. 361–722.
R Development Core Team (2012) R: A language and environment for statistical computing. R Foundation for Statistical Computing, Vienna, Austria, Available from: http://www.R-project.org/.
Samin, N., Zhou, H.-Z. & Imani, S. (2011) A contribution to the Oxyteline group of rove beetles (Coleoptera: Staphylinoidea: Staphylinidae) from Iran. Entomofauna, 32, 277–282.
Scheerpeltz, O. (1933) Staphylinidae VII (Pars 129). In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. vol. VI. W. Junk, Berlin, pp. 989–1500.
Schubert, K. (1906) Einige neue paläalrktische Staphyliniden aus Kashmir. Deutsche Entomologische Zeitschrift, 1906, 378–383.
Schülke, M. (2009) Zur Taxonomie und Faunistik westpalaearktischer Staphylinidae (Coleoptera: Staphylinidae: Omaliinae, Oxytelinae et Tachyporinae). Linzer Biologische Beitraege, 41, 803–844.
Schülke, M. & Uhlig, M. (1988) Faunistisch neue und bemerkenswerte Kurzfluglerarten aus der DDR (Coleoptera, Staphylinidae, Micropeplinae—Tachyporinae). Entomologische Nachrichten und Berichte, 32, 1–16.
Sharp, D.S. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1874, 1–103.
Shibata, Y. (1973) Preliminary check list of the family Staphylinidae of Taiwan (Insecta: Coleoptera). Annual Bulletin of the Nichidai Sanko, 16, 21–88.
Shibata, Y. (1986) A list of genera and species new to Taiwan and new data on distribution of the Staphylinidae discovered from Taiwan since 1973 (Insecta: Coleoptera). Annual Bulletin of the Nichidai Sanko, 24, 109–128.
Shibata, Y. (1993) New records of staphylinid beetles (Coleoptera) from Taiwan. Elytra, 21, 317–318.
Smetana, A. (2004) Subfamily Oxytelinae Fleming, 1821. In: Löbl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera: Hydrophiloidea—Histeroidea—Staphylinoidea. vol. 2. Apollo Books, Stenstrup, pp. 511–535.
Smetana, A. & Davies, A. (2000) Reclassification of the North Temperate Taxa Associated with Staphylinus Sensu Lato, Including Comments on Relevant Subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, No.3287, 1–88.
Stan, M. (2008) New data on the rove beetle fauna (Coleoptera: Staphylinidae) from Bucuresti and its surroundings. Travaux du Museum National d’Histoire Naturelle “Grigore Antipa”, 51, 369–386.
Staniec, B. (1992) The morphology of the larva of Oxytelus piceus (Linnaeus, 1767) (Coleoptera, Staphylinidae). Annales Universitatis Mariae Curie-Sklodowska Sectio C Biologia, 47, 31–39.
Staniec, B. (1993) The morphology of the pupa of coprophagous beetles Oxytelus piceus (Linnaeus, 1767) and Platystethus arenarius (Fourcroy, 1785) Coleoptera, Staphylinidae. Annales Universitatis Mariae Curie-Sklodowska Sectio C Biologia, 48, 241–250.
Szujecki, A. (2008) Beetles—Coleoptera. Issue 24a Rove beetles—Staphylinidae: introduction and subfamilies Micropeplinae, Piestinae, Osoriinae, Pseudopsiinae, Phleocharinae, Olisthaerinae, Proteininae, Omaliinae, Oxytelinae, Oxyporinae. Polskie Towarzystwo Entomologiczne Klucze do Oznaczania Owadow Polski, 175, 1–229.
Thomson, C.G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. vol. I. Berlingska Boktryckeriet, Lund, 290 pp.
Thomson, C.G. (1861) Skandinaviens Coleoptera, synoptiskt bearbetade. vol. III. Berlingska Boktryckeriet, Lund, 278 pp.
Walker, F. (1859) Characters of some apparently undescribed Ceylon insects. The Annals and Magazine of Natural History, 3, 50–56.
Winkler, A. (1925) Catalogus Coleopterorum Regionis Palaearcticae. Winkler & Wagner, Wien, pp. 241–624.
Yuh, J.-H., Paik, W.-H., Kwon, Y.-J. & Lee, S.-M. (1985) Check list of rove beetles from Korea (Coleoptera: Staphylinidae). Insecta Koreana, 5, 223–255.
Zhu, L., Wang, Y.-Y. & Li, L.-Z. (2007) Two new records of the Genus Oxytelus from China (Coleoptera: Staphylinidae, Oxytelinae). Journal of Shanghai Normal University (Natural Sciences), 36, 74–77.