Abstract
A provisional list of lower taxa (genera, subgenera) for the butterfly family Pieridae is provided based on review of the literature. A total of 98 taxa (83 genera, 15 subgenera) is recognised. Most of the genera appear in relatively good order but there is considerable doubt concerning the monophyly of Eurema sensu lato, Eurema sensu stricto, Catasticta sensu lato and Appias sensu lato. Systematic revision of Eurema sensu lato, Phoebis sensu lato, Hesperocharis sensu lato and Appias sensu lato is also required to determine the status or taxonomic rank (i.e. genera or subgenera) of taxa currently or previously included in these genera in the broad sense. The status of subgenera currently assigned to Colias sensu stricto appears doubtful. It is recommended that future systematic studies on lower classification focus on recognition of species-groups rather than subgenera within monophyletic genera.References
Ackery, P.R. (1975) A new pierine genus and species with notes on the genus Tatochila (Lepidoptera: Pieridae). Bulletin of the Allyn Museum, No. 30, 1–9.
Ackery, P.R. (1984) Systematics and faunistic studies on butterflies. In: Vane-Wright, R.I. & Ackery, P.R. (Eds.), The Biology of Butterflies. Symposium of the Royal Entomological Society of London, Number 11. Academic Press, London, pp. 11–21.
Ackery, P.R., de Jong, R. & Vane-Wright, R.I. (1999) The Butterflies: Hedyloidea, Hesperioidea and Papilionoidea. In: Kristensen, N.P. (Ed), Lepidoptera, Moths and Butterflies. Volume 1. Evolution, Systematics and Biogeography. de Gruyter, Berlin, pp. 263–300.
Ackery, P.R., Smith, C.R. & Vane-Wright, R.I. (1995) Carcasson’s African Butterflies. An Annotated Catalogue of the Papilionoidea and Hesperioidea of the Afrotropical Region. CSIRO Publishing, Melbourne, xi+803 pp.
Berger, L.A. (1986) Systématique du genre Colias F. Lepidoptera-Pieridae. Lambillionea, 86, 1–68 (suppl.).
Bernardi, G. (1947) Revision de la classification des espèces holarctiques des genres Pieris Schr. et Pontia Fabr. Miscellanea Entomologica, 44, 65–80.
Bernardi, G. (1953) Note sur la taxonomie des genres Belenois HB. et Anaphaeis HB., (Lep. Pieridae). Revue Française D'Entomologie, Paris, 20, 47–53.
Bernardi, G. (1958) Taxinomie et zoogéographie de Talbotia naganum Moore. Revue Française D'Entomologie, Paris, 25, 125–128.
Bridges, C.A. (1988) Catalogue of Papilionidae and Pieridae (Lepidoptera: Rhopalocera). Charles A. Bridges, Urbana, Illinois.
Brunton, C.F.A. (1998) The evolution of ultraviolet patterns in European Colias butterflies (Lepidoptera, Pieridae): a phylogeny using mitochondrial DNA. Heredity, 80, 611–616.
Brunton, C.F.A. & Hurst, G.D.D. (1998) Mitochondrial DNA phylogeny of brimstone butterflies (genus Gonepteryx) from the Canary Islands and Madeira. Biological Journal of the Linnean Society, 63, 69–79.
Cheong, S.W. (1990) Comparative morphology and phylogeny on the female internal genitalia of the Pieridae (Lepidoptera). Ph.D Thesis, Department of Biology, Kyungpook National University, Taegu, Korea, 80 pp.
Corbet, A.S. (1946) The conspecificity of Phrissura aegis (C. & R. Feld) and Udaiana cynis (Hew.) (Lepidoptera: Pieridae). The Entomologist, 79, 145–146.
Courtney, S.P. (1986) The ecology of pierid butterflies: dynamics and interactions. Advances in Ecological Research, 15, 51–131.
De Baar, M. (2004) Notes on the status of some Elodina C. & R. Felder species (Lepidoptera: Pieridae). Australian Entomologist, 31, 37–42.
De Baar, M. & Hancock, D.L. (1993) The Australian species of Elodina C. & R. Felder (Lepidoptera: Pieridae). Australian Entomologist, 20, 25–43.
de Jong, R., Vane-Wright, R.I. & Ackery, P. R. (1996) The higher classification of butterflies (Lepidoptera): problems and prospects. Entomologica Scandinavica, 27, 65–101.
Edwards, E.D., Newland, J. & Regan, L. (2001) Lepidoptera: Hesperioidea, Papilionoidea. Zoological Catalogue of Australia. Vol. 31.6. CSIRO Publishing, Melbourne, x+615 pp.
Ehrlich, P.R. (1958) The comparative morphology, phylogeny and higher classification of the butterflies (Lepidoptera: Papilionoidea). The University of Kansas Science Bulletin, 39, 305–364.
Eitschberger, U. & Racheli, T. (1998) Catasticta studies (Lepidoptera: Pieridae). Neue Entomologische Nachrichten, 41, 5–93.
Ferris, C.D. (1993) Reassessment of the Colias alexandra group, the legume-feeding species, and preliminary cladistic analysis of the North American Colias (Pieridae: Coliadinae). Bulletin of the Allyn Museum, No. 138, 1–85.
Field, W.D. (1950) A revision of Eurema Hübner subgenus Teriocolias Röber. Acta Zoologica Lilloana, 9, 359–374.
Field, W.D. (1958) A redefinition of the butterfly genera Tatochila, Phulia, Piercolias, and Baltia, with descriptions of related genera and subgenera. Proceedings of the United States National Museum, 108, 103–131.
Field, W.D. & Herrera, J.V. (1977) The pierid butterflies of the genera Hypsochila Ureta, Phulia Herrich-Schaffer, Infraphulia Field, Pierphulia Field, and Piercolias Staudinger. Smithsonian Contributions to Zoology, 232, 1–64.
Geiger, H.J. (1981) Enzyme electrophoretic studies on the genetic relationships of pierid butterflies (Lepidoptera: Pieridae) I. European taxa. The Journal of Research on the Lepidoptera, 19, 181–195.
Geiger, H.J. & Scholl, A. (1985) Systematics and evolution of holarctic Pierinae (Lepidoptera). An enzyme electrophoretic approach. Experientia, 41, 24–29.
Hemming, F. (1964) Paper XLIII. Establishment of a new genus in the family Pieridae. Annotationes Lepidopterologicae, 4, 148–149.
Henning, G.A., Henning, S.F., Joannou, J.G. & Woodhall, S.E. (1997) Living Butterflies of Southern Africa. Biology, Ecology and Conservation. Volume I. Hesperiidae, Papilionidae and Pieridae of South Africa. Umdaus Press, Hatfield, South Africa, 397 pp.
Herrera, G.J. & Field, W.D. (1959) A revision of the butterfly genera Theochila and Tatochila (Lepidoptera: Pieridae). Proceedings of the United States National Museum, 108, 467–514.
Klots, A.B. (1933) A generic classification of the Pieridae (Lepidoptera) together with a study of the male genitalia. Entomologica america, 12, 139–242.
Kudrna, O. (1974) Artogeia Verity, 1947, Gen. Rev. for Papilio napi Linnaeus (Lep., Pieridae). Entomologist's Gazette, 25, 9–12.
Lamas, G. (1979) Los Dismorphiinae (Pieridae) de México, América Central y las Antillas. Revista de la Sociedad Mexicana de Lepidopterologia, 5, 3–37.
Lamas, G. (2005) Pieridae. In: Lamas, G. (Ed), Checklist: Part 4A. Hesperioidea-Papilionoidea. In: Heppner, J. B. (Ed), Atlas of Neotropical Lepidoptera. Volume 5A. Association for Tropical Lepidoptera/Scientific Publishers, Gainesville (in press).
Lamas, G. & Bollino, M. (2004) Revisional notes on the "amastris" group of Catasticta Butler, 1870 (Lepidoptera: Pieridae), with descriptions of new species and subspecies. Zootaxa, 605, 1–19.
Larsen, T. (1991) Butterflies of Kenya and Their Natural History. Oxford University Press, Oxford, xxii+490 pp.
Mariani, M. (1937) Anatomia e fisiologia degli organi genitali femminili delle Pieris Schrk. (Lepidoptera, Pieridae). Festschrift 60 Geburtstag E. Strand. Vol. III, Riga Universitate, 3, 434–450.
Müller, C.J. (1999) A new species of Leuciacria Rothschild & Jordan (Lepidoptera: Pieridae) from montane New Ireland, Papua New Guinea. Australian Entomologist, 26, 65–70.
Müller, C.J. (2001) The previously unknown females of Leuciacria olivei Müller and Pseudodipsas una D'Abrera (Lepidoptera: Pieridae and Lycaenidae). Australian Entomologist, 28, 23–24.
Petersen, B. (1963) The male genitalia of some Colias species. The Journal of Research on the Lepidoptera, 1, 135–156.
Pitkin, B. & Jenkins, P. (2004) Butterflies and moths of the world: generic names and their type species. Department of Entomology, The Natural History Museum, London. Available from: http://www.nhm.ac.uk/entomology/butmoth/index.html (accessed April 2004)
Pollock, D.D., Watt, W.B., Rashbrook, V.K. & Iyengar, E.V. (1998) Molecular phylogeny for Colias butterflies and their relatives (Lepidoptera: Pieridae). Annals of the Entomological Society of America, 91, 524–531.
Reissinger, E. (1972) Zur Taxonomie und Systematik der Gattung Catasticta Butler (Lepidoptera, Pieridae). Entomologische Zeitschrift, 82, 97–124.
Robbins, R.K. (1982) How many butterfly species? News of the Lepidopterists' Society, 3, 40–41.
Robbins, R.K. & Henson, P. M. (1986) Why Pieris rapae is a better name than Artogeia rapae (Pieridae). Journal of the Lepidopterists' Society, 40, 79–92.
Shapiro, A.M. (1991) The zoogeography and systematics of the Argentine Andean and Patagonian Pierid Fauna. The Journal of Research on the Lepidoptera, 28, 137–238.
Shapiro, A.M. (1993) The proposed Magellanic type-locality of Colias imperialis Butler (Lepidoptera: Pieridae). Acta Entomológica Chilena, 18, 77–82.
Shapiro, A.M. (1994) Why are there so few butteflies in the high Andes? The Journal of Research on the Lepidoptera, 31, 25–56.
Stoneham, H.F. (1940) Notes on the African Genus Colotis, Hübnr (Teracolus, Swin.) and some new forms. Bulletin of the Stoneham Museum, Nairobi, 40, 1–4.
Stoneham, H.F. (1957) Two new generic names, and new subspecies and forms of Pieridae. Bulletin of the Stoneham Museum, Nairobi, 70, 1–3.
Talbot, G. (1928–37) A Monograph of the Pierine Genus Delias. Parts I–VI. British Museum (Natural History), London, 656 pp.
Talbot, G. (1944) A preliminary revision of the genus Mylothris Hübn. (1819) (Lep. Rhop. Pieridae). Transactions of the Royal Entomological Society of London, 94, 155–185.
Ureta, E. (1955) Nuevas especies [sic] de Pieridae (Lep. Rhopalocera) de Chile y Argentina. Boletín del Museo Nacional de Historia Natural (Santiago), 26, 57–71.
van Son, G. (1949) The Butterflies of Southern Africa. Part 1. Papilionidae and Pieridae. The Transvaal Museum, Pretoria, vi+237 pp., 41 pls.
Vane-Wright, R.I. (2003) Evidence and identity in butterfly systematics. In: Boggs, C.L., Watt, W. B. & Ehrlich, P.R. (Eds.), Butterflies: Ecology and Evolution Taking Flight. University of Chicago Press, Chicago, pp. 477–513.
Vane-Wright, R.I. & de Jong, R. (2003) The butterflies of Sulawesi: annotated checklist for a critical island fauna. Zoologische Verhandelingen, Leiden (343), 3–267.
Venables, B.A.B. (1993) Phylogeny of the White and Sulphur Butterflies (Pieridae). Unpublished dissertation project proposal, made available by Dr R.K. Robbins, National Museum of Natural History, Smithsonian Institution, Washington, 75 pp.
Verhulst, J.T. (2000) Les Colias du Globe. Goecke & Evers, Keltern, 308 pp.
Verity, R. (1947) Le farfalle diurne d'Italia. Vol. III. Casa Editrice Marzocco, Firenze, 318 pp.
Yamauchi, T. & Yata, O. (2000) Systematics and biogeography of the genus Gandaca Moore (Lepidoptera: Pieridae). Entomological Science, 3, 331–343.
Yata, O. (1985) Part 1: Pieridae. In: Tsukada, E. (Ed), Butterflies of the South East Asian islands. II: Pieridae, Danaidae. Plapac, Tokyo, pp. 33–120, 205–438.
Yata, O. (1989) A revision of the Old World species of the genus Eurema Hübner (Lepidoptera, Pieridae). Part I. Phylogeny and zoogeography of the subgenus Terias Swainson and description of the subgenus Eurema Hübner. Bulletin of the Kitakyushu Museum of Natural History, 9, 1–103.