Abstract
The family Phytoseiidae includes more than 2,300 species distributed all over the world. However despite the huge numbers of faunistic surveys carried out for more than 60 years, the fauna of some countries and particular ecosystems remain little explored. This paper reports results of surveys carried out in various regions of Morocco (from Sahara to Atlantic and Mediteranean coasts) in 2002 and 2003. A total of 43 species was found. Among them 19 are new for the Moroccan fauna and five are new to science. This paper provides the descriptions of these five new species, Neoseiulus thymeleae, Transeius audeae, Typhlodromus (Typhlodromus) ballotae, T. (T.) leclanti, T. (T.) mazarii, and re-descriptions of two species (Typhlodromus (T.) setubali and Typhlodromus (Anthoseius) clairathiasae. A key to females of the 52 species now known from Morocco is given.
References
Abad-Moyano, R., Pina, T., Ferragut, F. & Urbaneja, A. (2009) Comparative life-history traits of three phytoseiid mites associated with Tetranychus urticae (Acari: Tetranychidae) colonies in clementine orchards in eastern Spain: implications for biological control. Experimental and Applied Acarology, 47, 121–132.
http://dx.doi.org/10.1007/s10493-008-9197-zAbbassy, M.R., Hendy, H.H., Mowafi, M.H. & Nawar, M.A. (2012) Biology of Euseius scutalis (Acari: Phytoseiidae) on Tetranychus urticae and Panonychus ulmi (Acari: Tetranychidae) at different temperatures. Acarines, 6, 15–19.
Abo-Shnaf, R.I. & Moraes, G.J. (2014) Phytoseiid mites (Acari: Phytoseiidae) from Egypt, with new records, descriptions of new species, and a key to species. Zootaxa, 3865 (1), 1–71.
http://dx.doi.org/10.11646/zootaxa.3865.1.1Al-Shammery, K.A. (2010) Different biological aspects of the predaceous mite Euseius scutalis (Acari: Gamasida: Phytoseiidae) and the effects due to feeding on three tetranychid mite species in Hail, Saudi Arabia. Asian Journal of Biological Sciences, 3, 77–84.
http://dx.doi.org/10.3923/ajbs.2010.77.84Amitai, S. & Grinberg, T. (1971) Description of a new phytoseiid genus and species (Acarina: Mesostigmata) from Israel. Israel Journal of Entomology, 6, 327–335.
André, H.M. (1986) Notes on the ecology of corticolous epiphyte dwellers. 4. Actinedida (especially Tydeidae) and Gamasida (especially Phytoseiidae). Acarologia, 27, 107–115.
Arutunjan, E.S. (1970) Phytoseiid mites (Phytoseiidae) on agricultural crops in the Armenian SSR. Akademii Nauk Armyanskoi SSR, Otdelenie Biologicheskikh Nauk, Dissertatsii na Soiskanie Uchenoi Stepeni Candidata Biologrcheskikh Nauk, Zooliya, 97, 1–31.
Athias-Henriot, C. (1957a) Phytoseiidae et Aceosejidae (Acarina, Gamasina) d'Algérie. I. Genres Blattisocius Keegan, Iphiseius Berlese, Amblyseius Berlese, Phytoseius Ribaga, Phytoseiulus Evans. Bulletin de la Société d’Histoire Naturelle de l’Afrique du Nord, 48, 319–352.
Athias-Henriot, C. (1957b) Typhlodromus ornatus n. sp. et Typhlodromus longilaterus n. sp. (Acarina: Phytoseiidae) mésostigmates prédateurs d'Algérie. Revue de Pathologie Végétale et d’Entomologie Agricole de France, 36, 215–222.
Athias-Henriot, C. (1958a) Contribution à la connaissance du genre Typhlodromus Scheuten (Acariens Parasitiformes, Phytoseiidae). Description de deux espèces nouvelles d'Algérie et clé des espèces du groupe finlandicus. Revue de Pathologie Végétale et d’Entomologie Agricole de France, 37, 179–186.
Athias-Henriot, C. (1958b) Phytoseiidae et Aceosejidae (Acarina: Gamasina) d'Algérie. II. Phytoseiidae. Clé des genres Amblyseius Berlese (Suite) et Seiulus Berlese. Bulletin de la Société d’Histoire Naturelle de l’Afrique du Nord, 49, 23–43.
Athias-Henriot, C. (1959) Acariens planticoles d'Algerie. I. 5e contribution au genre Amblyseius Berlese (Phytoseiidae). II. Première liste d'Actinochitinosi (Cheyletidae, Caligonellidae, Hemisarcoptidae). Bulletin de l’Académie Royale de Belgique, 45, 130–153.
Athias-Henriot, C. (1960a) Nouveaux Amblyseius d'Algérie (Parasitiformes, Phytoseiidae). Acarologia, 2, 288–299.
Athias-Henriot, C. (1960b) Phytoseiidae et Aceosejidae (Acarina: Gamasina) d'Algérie. IV. Genre Typhlodromus Scheuten, 1857. Bulletin de la Société d’Histoire Naturelle de l’Afrique du Nord, 51, 62–107.
Athias-Henriot, C. (1961) Mésostigmates (Urop. excl.) édaphiques Méditerranéens (Acaromorpha, Anactinotrichida). Acarologia, 3, 381–509.
Athias-Henriot, C. (1966) Contribution à l'étude des Amblyseius paléarctiques (Acariens anactinotriches, Phytoseiidae). Bulletin Scientifique de Bourgogne, 24, 181–230.
Athias-Henriot, C. (1975) Nouvelles notes sur les Amblyseiini. II. Le relevé organotaxique de la face dorsale adulte (Gamasides protoadéniques, Phytoseiidae). Acarologia, 17, 20–29.
Athias-Henriot, C. (1978) Typhlodromini du Vaucluse, avec description de trois espèces nouvelles (Arachnides, Gamasides, Phytotseiidae). Annales de Zoologie et d’Ecologie Animale, 10, 695–701.
Beglyarov, G.A. (1958) Species of Phytoseiidae (Parasititormes: Gamasoidea) predatory upon tetranychid mites in orchards of the Krasnodar region. Trudy Vsesoiuznogo Institut Zashchity Rastenii, 10, 98–124.
Beglyarov, G.A. & Malov, N.A. (1977) Predaceous phytoseiid mites (Phytoseiidae: Parasitiformes) of the fauna of Moldavia and north Bukovina (Report II). Entomofagi v Zashchite Rasteniy. Vsesoyuznaya Ordena Lenina Akademiya Selískokhozyaystvennyukh Nauk Imeni V. I. Lenina. Kishinev, Russia, pp. 3–11.
Berlese, A. (1889) Acari, Myriopoda et Scorpiones hucusque in Italia reperta. Tipografia del Seminario, 6, 7–9.
Berlese, A. (1921) Acari, Myriopoda et Pseudoscorpiones hucusque in Italia reperta. I. Indice sinonimico dei generi e delle specie illustrate nei fascicoli 1 a 101. Redia, 14, 77–105.
Boller, E.F. (1984) Eineainfache Ausschwemm-methodezurschellen Erfassung von raumilben, Thrips and andren Kleinarthropodenimb Weinbau. Schweizerische Zeitschrift fur Obst- und Weinbau, 120, 249–255.
Castagnoli, M., Liguori, M. & Nannelli, R. (1984) Contributo alla conoscenza degli Acari del pesco in Toscana e osservazioni sull’andamento delle loro popolazioni. Redia, 67, 493–504.
Castagnoli, M., Liguori, M. & Simoni, S. (1999) Effect of two different host plants on biological features of Neoseiulus californicus (McGregor). International Journal of Acarology, 25, 145–150.
http://dx.doi.org/10.1080/01647959908683626Chant, D.A. (1956) Some mites of the subfamily Phytoseiinae (Acarina: Laelaptidae) from southeastern England, with descriptions of new species. The Canadian Entomologist, 88, 26–37.
http://dx.doi.org/10.4039/Ent8826-1Chant, D.A. (1957) Descriptions of some phytoseiid mites (Acarina, Phytoseiidae). Part I. Nine new species from British Columbia with keys to the species of British Columbia. Part II. Redescriptions of eight species described by Berlese. The Canadian Entomologist, 89, 289–308.
http://dx.doi.org/10.4039/Ent89289-7Chant, D.A. (1959) Phytoseiid mites (Acarina: Phytoseiidae). Part I. Bionomics of seven species in southeastern England. Part II. A taxonomic review of the family Phytoseiidae, with descriptions of thirty-eight new species. Canadian Entomologist Supplement, 12, 1–166.
Chant, D.A. & Hansell, R.I.C. (1971) The genus Amblyseius (Acarina: Phytoseiidae) in Canada and Alaska. Canadian Journal of Zoology, 49, 703–758.
http://dx.doi.org/10.1139/z71-110Chant, D.A. & McMurtry, J.A. (2005) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae) Part VI. The tribe Euseiini N. tribe, subtribes Typhlodromalina, N. subtribe, Euseiina, N. subtribe and Ricoseiina, N. subtribe. International Journal of Acarology, 31, 187–224.
http://dx.doi.org/10.1080/01647950508684424Chant, D.A. & McMurtry, J.A. (2007) Illustrated Keys and Diagnoses for the Genera and Sub-genera of the Phytoseiidae of the World. Indira Publishing House, West Bloomfield, Michigan, 220 pp.
Chant, D.A. & Yoshida-Shaul, E. (1982) A world review of the soleiger species group in the genus Typhlodromus Scheuten (Acarina: Phytoseiidae). Canadian Journal of Zoology, 60, 3021–3032.
http://dx.doi.org/10.1139/z82-385Chant, D.A. & Yoshida-Shaul, E. (1987) A world review of the pyri species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae). Canadian Journal of Zoology, 65, 1770–1804.
http://dx.doi.org/10.1139/z87-272Chant, D.A. & Yoshida-Shaul, E. (1991) Adult ventral setal patterns in the family Phytoseiidae (Acari: Gamasina). International Journal of Acarology, 17, 187–199.
http://dx.doi.org/10.1080/01647959108683906Charlet, L.D. & McMurtry, J.A. (1977) Systematics and bionomics of predaceous and phytophagous mites associated with pine foliage in California. Hilgardia, 45, 173–236.
http://dx.doi.org/10.3733/hilg.v45n07p173Coiutti, C. (1993) Phytoseiid mites on spontaneous and cultivated trees in the Friuli-Venezia Giulia region. Frustula Entomologica, 16, 65–77.
De Leon, D. (1965) A note on Neoseiulus Hughes 1948 and new synonymy (Acarina: Phytoseiidae). Proceedings of the Entomological Society of Washington, 67, 23.
Demite, P.R., Lofego, A.C. & Feres, R.J.F. (2008) A new species of Galendromus (Acari: Phytoseiidae): A case of intraspecific variation and redefinition of carinulatus species group. Annals of the Entomological Society of America, 101, 804–807.
http://dx.doi.org/10.1093/aesa/101.5.804Demite, P.R., McMurtry, J.A. & de Moraes, G.J. (2014) Phytoseiidae Database: a website for taxonomic and distributional information on phytoseiid mites (Acari). Zootaxa, 3795 (5), 571–577.
http://dx.doi.org/10.11646/zootaxa.3795.5.6Daneshvar, H. (1987) Some predatory mites from Iran, with descriptions of one new genus and six new species (Acari: Phytoseiidae, Ascidae). Entomologie et Phytopathologie Appliquées, 54, 13–37.
Denmark, H.A. (1966) Revision of the genus Phytoseius Ribaga, 1904 (Acarina: Phytoseiidae). Florida Department of Agriculture Bulletin, 6, 1–105.
Denmark, H.A. (1992) A revision of the genus Typhlodromus Scheuten (Acari: Phytoseiidae). Occasional Papers of the Florida State Collection of Arthropods, 7, 1–43.
Denmark, H.A. & Muma, M.H. (1989) A revision of the genus Amblyseius Berlese, 1914 (Acari: Phytoseiidae). Occasional Papers of the Florida State Collection of Arthropods, 4, 1–149.
Denmark, H.A. & Rather, A.Q. (1996) Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae). International Journal of Acarology, 22, 43–77.
http://dx.doi.org/10.1080/01647959608684080Denmark, H.A. & Welbourn, W.C. (2002) Revision of the genera Amblydromella Muma and Anthoseius De Leon (Acari: Phytoseiidae). International Journal of Acarology, 28, 291–316.
http://dx.doi.org/10.1080/01647950208684308Denmark, H.A., Evans, G.A., Aguilar, H., Vargas, C. & Ochoa, R. (1999) Phytoseiidae of Central America. Indira Publishing House, West Bloomfield, Michigan, 125 pp.
Doker, I., Stathakis, T.I., Kazak, C., Karut, K. & Papadoulis, G.T. (2014) Four new records and two new species of Phytoseiidae (Acari: Mesostigmata) from Turkey, with a key to the Turkish species. Zootaxa, 3827 (3), 331–342.
http://dx.doi.org/10.11646/zootaxa.3827.3.3Dosse, G. (1958) Uber einige neue Raubmilbenarten (Acar.: Phytoseiidae). Pflanzenschutz Berichte, 21, 44–61.
Dosse, G. (1961) Zur Klarung der Artenfrage von Typhlodromus (Typhlodromus) pyri Scheuten, 1857 (= T. tiliae Oud. 1929) und Typhlodromus (Typhlodromus) sebutali n. sp. (Acari: Phytoseiidae). Zeitschrift fur Angewandte Zoologie, 48, 313–323.
Duso, C. & Pasini, M. (2003) Distribution of the predatory mite Amblyseius andersoni Chant (Acari: Phytoseiidae) on different apple cultivars. Anzeiger für Schadlingskunde, 76, 33–40.
http://dx.doi.org/10.1046/j.1439-0280.2003.03003.xDuso, C. & Sbrissa, F. (1990) Gli Acari fitoseidi (Acari, Phytoseiidae) del melo nell'Italia settentrionale: distribuzione, biologia, ecologia ed importanza economica. Bolletino de Zoologia Agraria e Bachicoltura, Serie II, 22, 53–89.
Duso, C., Pozzebon, A., Capuzzo, C., Malagnini, V., Otto, S. & Borgo, M. (2005) Grape downy mildew spread and mite seasonal abundance in vineyards: effects on Tydeus caudatus and its predators. Biological Control, 32, 143–154.
http://dx.doi.org/10.1016/j.biocontrol.2004.09.004Ehara, S. (1966) A tentative catalogue of predatory mites of Phytoseiidae known from Asia, with descriptions of five new species from Japan. Mushi, 39, 9–30.
Ehara, S. & Amano, H. (1998) A revision of the mite family Phytoseiidae in Japan (Acari: Gamasina), with remarks on its biology. Species Diversity, 3, 25–73.
Ehara, S., Okada, Y. & Kato, H. (1994) Contribution to the knowledge of the mite family Phytoseiidae in Japan (Acari: Gamasina). Journal of the Faculty of Education, Tottori University, Natural Science, 42, 119–160.
El-Sawi, S.A. & Momen, F.M. (2005) Biology of some phytoseiid predators (Acari: Phytoseiidae) on eggs of Phthorimaea operculella and Spodoptera littoralis (Lepidoptera: Gelechiidae and Noctuidae). Acarologia, 45, 23–30.
Espinha, I.G., Costa-Comelles, J., Torres, L.M. & Ferragut, F. (1995) Contribuição para o conhecimen to dos ácaros fitoseídeos associados à macieira no norte interior de Portugal. Boletin de Sanidad Vegetal Plagas, 21, 223–230.
Evans, G.O. (1987) The status of three species of Phytoseiidae (Acari) described by Carl Willmann. Journal of Natural History, 21, 1461–1467.
http://dx.doi.org/10.1080/00222938700770921Evans, G.O. & Edland, T. (1998) The genus Anthoseius De Leon (Acari: Mesostigmata) in Norway. Fauna Norvegica, Series B, 45, 41–62.
Fantinou, A.A., Baxevani, A., Drizou, F., Labropoulos, P., Perdikis, D. & Papadoulis, G. (2012) Consumption rate, functional response and preference of the predaceous mite Iphiseius degenerans to Tetranychus urticae and Eutetranychus orientalis. Experimental and Applied Acarology, 58, 133–144.
http://dx.doi.org/10.1007/s10493-012-9557-6Ferragut, F. & Escudero, A. (1997) Taxonomia y distribucion de los acaros depredadores del genero Euseius Wainstein 1962, en Espana (Acari: Phytoseiidae). Boletin de Sanidad Vegetal Plagas, 23, 227–235.
Ferragut, F. & Pena-Estevez, M.A. (2003) Phytoseiid mites of the Canary Islands (Acari: Phytoseiidae). 1. Gran Canaria Island. International Journal of Acarology, 29, 149–170.
http://dx.doi.org/10.1080/01647950308683654Ferragut, F. & Ueckermann, E. (2012) A new species and new records of the subgenus Typhlodromus Scheuten from Spain, with a key to the world species (Acari: Phytoseiidae). Journal of Natural History, 46, 1731–1745.
http://dx.doi.org/10.1080/00222933.2012.681318Ferragut, F., Garcia-Mari, F. & Marzal, M.C. (1983) Determinacion y abundancia de los fitoseidos (Acari: Phytoseiidae) en los agrios espanoles. I. Congreso Nacional de la Sociedad Espanola de Ciencias Horticolas, Valencia, 28 Noviembre a 1 Diciembre de 1983, Spain, 299–308.
Ferragut, F., Costa-Comelles, J., Gomez Bernardo, J. & Garcia Mari, F. (1985) Contribucion al conocimiento de los acaros fitoseidos (Gamasida: Phytoseiidae) de los cultivos espanoles . Actas II Congreso Iberico de Entomologia, Suplemento No 1, 2, pp. 223–231.
Ferragut, F., Laborda, R., Costa-Comelles, J. & Garcia-Mari, F. (1992) Feeding behaviour of Euseius stipulatus and Typhlodromus phialatus on the citrus red mite Panonychus citri (Acari: Phytoseiidae, Tetranychidae). Entomophaga, 37, 537–543.
http://dx.doi.org/10.1007/BF02372323Fischer, S. & Mourrut-Salesse, J. (2005) L'acariose bronzée de la tomate en Suisse (Aculops lycopersici: Acari, Eriophyidae). Revue Suisse de Viticulture, Arboriculture et Horticulture, 37, 227–232.
Fitzgerald, J., Xu, X.M., Pepper, N., Easterbrook, M. & Solomon, M. (2008) The spatial and temporal distribution of predatory and phytophagous mites in field-grown strawberry in the UK. Experimental and Applied Acarology, 44, 293–306.
http://dx.doi.org/10.1007/s10493-008-9151-0Gambaro, P.I. (1994) The importance of humidity in the development and spread of Amblyseius andersoni Chant (Acarina, Phytoseiidae). Bollettino di Zoologia Agraria e di Bachicoltura, 26, 241–248.
Goven, M.A., Cobanoglu, S. & Guven, B. (2009) Ege Bolgesi bag alanlarndaki avc akar faunas. Bitki Koruma Bulteni, 49, 1–10.
Hadju, Z., Szabo, A. & Penzes, B. (2011) The zoophagous mite fauna on the evergreen ornamental trees and the ornamental shrubberies. Kertgazdasag, 43 (4), 69–76.
Hajidazeh, J. & Nazari, M. (2012) A checklist and key for the phytoseiid mites (Acari: Phytoseiidae) of citrus orchards in Iran, with a new record for Iranian phytoseiid mites. Systematic & Applied Acarology, 17, 388–396.
Hirschmann, W. (1962) Gangystematik der Parasitiformes. Teil 5. Acarologie. Schriftenreihe für Vergleichende Milbenkunde, 5, 1–56.
Houten, Y.M. van, Ostlie, M.L., Hoogerbrugge, H. & Bolckmans, K. (2005) Biological control of western flower thrips on sweet pepper using the predatory mites Amblyseius cucumeris, Iphiseius degenerans, A. andersoni and A. swirskii. Bulletin OILB/SROP, 28, 283–286.
Hughes, A.M. (1948) The Mites Associated with Stored Food Products. Ministry of Agriculture and Fishery, H. M. Stationery Office, London, 168 pp.
Hussein, H. & Momen, F. (2010) Fertilisation and prey deprivation affecting reproduction, life history and life table of the predacious mite Paraseiulus talbii (Athias-Henriot) (Acari: Phytoseiidae). Archives of Phytopathology and Plant Protection, 43, 242–251.
http://dx.doi.org/10.1080/03235400701722210Hwang-Yong, K., Yong-Hwi, L., Jeong-Hwan, K. & Yong-Heon, K. (2008) Comparison on the capability of four predatory mites to prey on the eggs of Bemisia tabaci (Hemiptera: Aleyrodidae). Korean Journal of Applied Entomology, 47, 429–433.
http://dx.doi.org/10.5656/KSAE.2008.47.4.429Kanouh, M., Tixier, M.-S., Okassa, M. & Kreiter, S. (2010a) Phylogenetic and biogeographic analysis of the genus Phytoseiulus (Acari: Phytoseiidae). Zoologica Scripta, 39, 450–461.
http://dx.doi.org/10.1111/j.1463-6409.2010.00439.xKanouh, M., Tixier, M.-S., Guichou, S., Cheval, B. & Kreiter, S. (2010b) Two synonymy cases within the genus Neoseiulella (Acari: Phytoseiidae): is the molecular evidence so evident? Biological Journal of Linnean Society, 101, 323–344.
Karg, W. (1983) Systematische untersuchung der Gattungen und Untergattungen der Raubmilbenfamilie Phytoseiidae Berlese 1916, mit der beschreibung von 8 neuen Arten. Mitteilungen Zoologisches Museum in Berlin, 59, 293–328.
http://dx.doi.org/10.1002/mmnz.4830590203Karg, W. (1989) Neue Raubmilbenarten der Gattuig Proprioseiopsis Muma, 1961 (Acarina, Parasitiformes) mit Bestimmungsschlusseln. Zoologische Jahrbücher Abteilung für Systematik, Ökologie und Geographie der Tiere, 116, 199–216.
Karg, W. (1991) Die Raubmilbenarten der Phytoseiidae Berlese (Acarina) Mitteleuropas sowie angrenzender Gebiete. Zoologische Jahrbücher Abteilung für Systematik, Ökologie und Geographie der Tiere, 118, 1–64.
Karg, W. (1993) Acari (Acarina), Milben, Parasitiformes (Anactinochaeta), Cohors Gamasina Leach, Raubmilben, Die Tierwelt Deutschlands, 59, 1–523.
Kasap, I. & Sekeroglu, E. (2004) Life history of Euseius scutalis feeding on citrus red mite Panonychus citri at various temperatures. BioControl, 49, 645–654.
http://dx.doi.org/10.1023/B:BICO.0000046733.53887.2bKazak, C., Yildiz, S. & Sekeroglu, E. (2002) Biological characteristics and life tables of Neoseiulus umbraticus Chant (Acari, Phytoseiidae) at three constant temperatures. Anzeiger fur Schadlingskunde, 75, 118–121.
http://dx.doi.org/10.1046/j.1472-8206.2002.02034.xKennett, C.E. & Caltagirone, L.E. (1968) Biosystematics of Phytoseiulus persimilis Athias-Henriot (Acarina: Phytoseiidae). Acarologia, 10, 563–577.
Kolodochka, L.A. (1978) Manual for the Identification of Plant-Inhabiting Phytoseiid mites. Akademii Nauk Ukrainian SSR, Instituta Zoologii, Naukova Dumka, Kiev, Ukraine, 79 pp.
Kolodochka, L.A. (1980) New phytoseiid mites (Parasitiformes, Phytoseiidae) from Molodova. Vestnik Zoologii, 4, 39–45.
Kolodochka, L.A. (1995) (1993) A revision of the genus Eharius (Parasitiformes, Phytoseiidae). Zhurnal Ukrainsíkogo Entomologichnogo Tovaristva, 1, 79–96.
Koveos, D.S. & Broufas, G.D. (2000) Functional response of Euseius finlandicus and Amblyseius andersoni to Panonychus ulmi on apple and peach leaves in the laboratory. Experimental and Applied Acarology, 24, 247–256.
http://dx.doi.org/10.1023/A:1006431710313Kreiter, S. & Brian, F. (1987) Les phytoseiidae de la vigne en France. In: Proceedings from International Conference on pests in Agriculture. FRA: ANPP, Paris, pp. 241–249.
Kreiter, S., Tixier, M.-S. & Allam, L. (2007) The diversity of phytoseiid mite communities in the Maghreb comparisons with the fauna of one of the Canary Islands. Journal of the Egyptian Society of Parasitology, 37, 381–394.
Kreiter, S., Tixier, M.-S., Croft, B.A., Auger, P. & Barret, D. (2002) Plants and leaf characteristics influencing the predaceous mite, Kampimodromus aberrans (Oudemans), in habitats surrounding vineyards (Acari: Phytoseiidae). Environmental Entomology, 31, 648–660.
http://dx.doi.org/10.1603/0046-225X-31.4.648Kreiter, S., Tixier, M.-S., Auger, P., Sentenac, G., Salva, G. & Weber, M. (2000) Phytoseiid mites of vineyards in France (Acari: Phytoseiidae). Acarologia, 41, 77–96.
Kreiter, S., Lebdi Grissa, K., Chabaan, S., Chatti, A., Tixier, M.-S., Auger, P., Chermiti, B., Ksantini, M. & Khoualdia, O. (2010) Phytoseiid mites (Acari: Mesostigmata) from Tunisia: catalogue, biogeography and key for identification. Tunisian Journal of Plant Protection, 5, 151–178.
Liguori, M. & Guidi, S. (1990) Influenze del condizionamento alimentare di Typhlodromus exhilaratus Ragusa (Acari: Phytoseiidae) sul suo consumo di preda. Redia, 73, 201–211.
Lindquist, E.E. & Evans, G.O. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata). Memoirs of the Entomological Society of Canada, 47, 1–64.
http://dx.doi.org/10.4039/entm9747fvLombardini, G. (1960) Typhlodromus baccettii nuova specie di acaro predatore. Redia, 45, 19–21.
Lorenzon, M., Pozzebon, A. & Duso, C. (2012) Effects of potential food sources on biological and demographic parameters of the predatory mites Kampimodromus aberrans, Typhlodromus pyri and Amblyseius andersoni. Experimental and Applied Acarology, 58, 259–278.
http://dx.doi.org/10.1007/s10493-012-9580-7Maoz, Y., Gal, S., Argov, Y., Coll, M. & Palevsky, E. (2011) Biocontrol of persea mite, Oligonychus perseae, with an exotic spider mite predator and an indigenous pollen feeder. Biological Control, 59, 147–157.
http://dx.doi.org/10.1016/j.biocontrol.2011.07.014McGregor, E.A. (1954) Two new mites in the genus Typhlodromus (Acarina: Phytoseiidae). Southern California Academy of Science Bulletin, 53, 89–92.
McMurtry, J.A. (1977) Some predaceous mites (Phytoseiidae) on citrus in the Mediterranean region. Entomophaga, 22, 19–30.
http://dx.doi.org/10.1007/BF02372986McMurtry, J.A. & Bounfour, M. (1989) Phytoseiid mites of Morocco, with descriptions of two new species and notes on the genera Kuzinellus, Typhloctonus and Typhlodromus (Acari: Phytoseiidae). Acarologia, 30, 13–24.
Messelink, G.J., Steenpaal, S.E.F. van & Ramakers, P.M.J. (2006) Evaluation of phytoseiid predators for control of western flower thrips on greenhouse cucumber. BioControl, 51, 753–768.
http://dx.doi.org/10.1007/s10526-006-9013-9Meszaros, A., Tixier, M.-S., Cheval, B., Barbar, Z. & Kreiter, S. (2007) Cannibalism and intraguild predation in Typhlodromus exhilaratus Ragusa and T. phialatus Athias-Henriot (Acari: Phytoseiidae) under laboratory conditions. Experimental and Applied Acarology, 41, 37–43.
http://dx.doi.org/10.1007/s10493-006-9046-xMiedema, E. (1987) Survey of phytoseiid mites (Acari: Phytoseiidae) in orchards and surrounding vegetation of northwestern Europe, especially in the Netherlands. Keys, descriptions and figures. Netherlands Journal of Plant Pathology, 93 (Supplement 2), 1–64.
http://dx.doi.org/10.1007/BF01984462Minarro, M. & Kreiter, S. (2012) Fitoseidos en los vinedos de la denominacion Vino de Calidad de Cangas (Asturias). Boletin de Sanidad Vegetal, Plagas, 38, 73–82.
Minarro, M., Dapena, E. & Ferragut, F. (2002) Phytoseid mites (Acari: Phytoseiidae) in plantations of apples in Asturias. Boletin de Sanidad Vegetal, Plagas, 28, 287–297.
Momen, F.M. & Abdel-Khalek, A. (2008) Effect of the tomato rust mite Aculops lycopersici (Acari: Eriophyidae) on the development and reproduction of three predatory phytoseiid mites. International Journal of Tropical Insect Science, 28, 53–57.
http://dx.doi.org/10.1017/S1742758408942594Momen, F.M. & El-Sawi, S. (2008) Life-history traits of the predacious mite Euseius scutalis (Athias-Henriot) (Acari: Phytoseiidae) on eggs of three insects (Lepidoptera: Noctuidae). Acta Phytopathologica and Entomologica Hungarica, 43, 163–170.
http://dx.doi.org/10.1556/APhyt.43.2008.1.16Moraes, G.J., McMurtry, J.A. & Denmark, H.A. (1986) A Catalog of the Mite Family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat. EMBRAPA, DDT, 353 pp.
Moraes, G.J., McMurtry, J.A., Denmark, H.A. & Campos, C.B. (2004) A revised catalog of the mite family Phytoseiidae. Zootaxa, 434, 1–494.
Moraes, G.J., Zannou, I.D., Ueckermann, E.A., Oliveira, A.R., Yaninek, J.S. & Hanna, R. (2007) Phytoseiid mites of the tribes Afroseiulini, Kampimodromini and Phytoseiulini, and complementary notes on mites of the tribes Euseiini and Neoseiulini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa, 1628, 1–22.
Moraza, M.L. & Pena, M.A. (2005) Acaros mesostigmata (Acari: Mesostigmata) en habitats seleccionados de la isla de Tenerife (Islas Canarias). Revista Iberica de Acarachnologia, 11, 61–68.
Muma, M.H. (1961) Subfamiles, genera, and species of Phytoseiidae (Acarina: Mesostigmata). Florida State Museum Bulletin, 5, 267–302.
Nicotina, M. (1996) Phytoseiid mites (Phytoseiidae) associated with vines in vineyards of the Campania Region. In: Mitchell, R., Horn, D.J., Needham, G.R. & Welbourn, W.C. (Eds.), Acarology IX. Proceedings. Ohio Biological Survey, USA, 1, pp. 237–240.
Nesbitt, H.H.J. (1951) A taxonomic study of the Phytoseiidae (Family Laelaptidae) predaceous upon Tetranychidae of economic importance. Zoologische Verhandelingen, 12, 1–64 + 32 pls.
Nomikou, M., Janssen, A., Schraag, R. & Sabelis, M.W. (2001) Phytoseiid predators as potential biological control agents for Bemisia tabaci. Experimental and Applied Acarology, 25, 271–291.
http://dx.doi.org/10.1023/A:1017976725685Ohno, S., Gotoh, T., Miyagi, A., Ganaha-Kikumura, T., Kurima, M., Kijima, K. & Ooishi, T. (2012) Geographic distribution of phytoseiid mite species (Acari: Phytoseiidae) on crops in Okinawa, a subtropical area of Japan. Entomological Science, 15, 115–120.
http://dx.doi.org/10.1111/j.1479-8298.2011.00469.xOudemans, A.C. (1930a) Acarologische Aanteekeningen. CI. Entomologische Berichten, 8, 48–53.
Oudemans, A.C. (1930b) Acarologische Aanteekeningen. CII. Entomologische Berichten, 8, 69–74.
Papadopoulos, G.D. & Papadoulis, G.T. (2008) Effect of seven different pollens on bio-ecological parameters of the predatory mite Typhlodromus foenilis (Acari: Phytoseiidae). Environmental Entomology, 37, 340–347.
http://dx.doi.org/10.1093/ee/37.2.340Papadoulis, G.Th. & Emmanouel, N.G. (1991) The genus Amblyseius (Acari: Phytoseiidae) in Greece, with the description of a new species. Entomologia Hellenica, 9, 35–62.
Papadoulis, G., Emmanouel, N.G. & Kapaxidi, E.V. (2009) Phytoseiidae of Greece and Cyprus. Indira Publishing House, West Bloomfield, Michigan, 200 pp.
Papaioannou-Souliotis, P., Ragusa di Chiara, S. & Tsolakis, C. (1994) Phytophagous mites and their predators observed on cultivated plants in Greece during 1975–1990. Annales Institut Phytopathologique Benaki, N.S., 17, 35–86.
Perez Otero, R., Mansilla Vazquez, P. & Lopez Villarreal, M.J. (2005) Primer inventario de acaros fitoseidos sobre cultivos horticolas en la comarca de O Salnes (Pondevedra). Boletin de Sanidad Vegetal, Plagas, 31, 343–350.
Peverieri, G.S., Simoni, S., Goggioli, D., Liguori, M. & Castagnoli, M. (2009) Effects of variety and management practices on mite species diversity in Italian vineyards. Bulletin of Insectology, 62, 53–60.
Pina, T., Sargolo, P., Urbaneja, A. & Jacas, J.A. (2012) Effect of pollen quality on the efficacy of two different life-style predatory mites against Tetranychus urticae in citrus. Biological Control, 61, 176–183.
http://dx.doi.org/10.1016/j.biocontrol.2012.02.003Pritchard, A.E. & Baker, E.W. (1962) Mites of the family Phytoseiidae from Central Africa, with remarks on genera of the world. Hilgardia, 33, 205–309.
http://dx.doi.org/10.3733/hilg.v33n07p205Ragusa, S. (1977) Notes on phytoseiid mites in Sicily with a description of a new species of Typhlodromus (Acarina: Mesostigmata). Acarologia, 18, 379–392.
Ragusa, S. (2006) Phytoseiid mites (Parasitiformes Phytoseiidae) of some Eptanissan Islands (Greece). Redia, 89, 1–7.
Ragusa, S. & Athias-Henriot, C. (1983) Observations on the genus Neoseiulus Hughes (Parasitiformes, Phytoseiidae). Redefinition. Composition. Geography. Description of two new species. Revue Suisse de Zoologie, 90, 657–678.
http://dx.doi.org/10.5962/bhl.part.82005Ragusa, S. & Ciulla, A.M. (1991) Phytoseiid mites associated with vines in Sicilian vineyards. In: Schuster, R. & Murphy, P.W. (Eds.), The Acari: Reproduction, Development and Life History Strategies. Chapman & Hall, London, pp. 417–423.
Ragusa, S. & Swirski, E. (1978) Description of three new species of Typhlodromus Scheuten from Italy with redescription of Typhlodromus baccettii Lombardini (Acari: Phytoseiidae). International Journal of Acarology, 4, 211–220.
http://dx.doi.org/10.1080/01647957808683118Ragusa, S. & Swirski, E. (1981) A new species of the genus Seiulus (Acarina: Phytoseiidae) from Italy. Redia, 64, 269–276.
Ragusa, S. & Tsolakis, H. (1998) Life-history data of Typhlodromus rhenanoides Athias-Henriot and Typhlodromus cryptus Athias-Henriot (Parasitiformes, Phytoseiidae) fed on pollen under laboratory conditions. Phytophaga, 8, 3–11.
Ragusa, S. & Tsolakis, H. (2001) Phytoseiid faunas of natural and agricultural ecosystems in Sicily. In: Halliday, R.B., Walter, D.E., Proctor, H.C., Norton, R.A. & Coloff, M.J. (Eds.), Acarology: Proceedings of the 10th International Congress. CSIRO Publishing, Melbourne, pp. 522–529.
Ragusa, S. & Vargas, R. (2002) On some phytoseiid mites (Parasitiformes: Phytoseiidae) from Chile. Phytophaga, 12, 129–139.
Ribaga, C. (1904) Gamasidi planticoli. Rivista di Patologia Vegetale, 10, 175–178.
Rahmani, H., Kamali, K. & Faraji, F. (2010) Predatory mite fauna of Phytoseiidae of northwest Iran (Acari: Mesostigmata). Turkish Journal of Zoology, 34, 497–508.
Rioja, T.S. & Vargas, R.M. (2009) Life table parameters and consumption rate of Cydnodromus picanus Ragusa, Amblyseius graminis Chant and Galendromus occidentalis (Nesbitt) on avocado red mite Oligonychus yothersi (McGregor) (Acari: Phytoseiidae, Tetranychidae). Chilean Journal of Agricultural Research, 69, 160–170.
http://dx.doi.org/10.4067/S0718-58392009000200005Rowell, H.J., Chant, D.A. & Hansell, R.I.C. (1978) The determination of setal homologies and setal patterns on the dorsal shield in the family Phytoseiidae. The Canadian Entomologist, 110, 859–876.
http://dx.doi.org/10.4039/Ent110859-8Sahraoui, H., Lebdi Grissa, K., Kreiter, S., Douin, M. & Tixier, M.-S. (2012) Phytoseiid mites (Acari: Mesostigmata) of Tunisian Citrus orchards. Acarologia, 52, 433–452.
http://dx.doi.org/10.1051/acarologia/20122072Santaballa, E., Roca, M. & Laborda, R. (1994) Comportamiento de clofentecin frente al acaro rojo Panonychus citri (McGregor) (Acari: Tetranychidae) y su predador Euseius stipulatus (Athias-Henriot) (Acari: Phytoseiidae) en los agrios espanoles. Boletin de Sanidad Vegetal, Plagas, 20, 419–428.
Schicha, E. & Dosse, G. (1974) A new species of Typhlodromus Scheuten (Acarina: Phytoseiidae) from apple in Australia. Proceedings of the Linnean Society of New South Wales, 99, 79–84.
Schuster, R.O. & Pritchard, A.E. (1963) Phytoseiid mites of California. Hilgardia, 34, 191–285.
http://dx.doi.org/10.3733/hilg.v34n07p191Schweizer, J. (1922) Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandlungen der Naturferschenden Gesellschaft in Basel, 33, 23–112.
Schweizer, J. (1949) Die Landmilben des Schweizerischen Nationalparkes. I. Teil: Parasitiformes Reuter, 1909. Ergebnisse der Wissenschaftlichen Untersuchungen im Schweizerischen Nationalpark, 2, 1–99.
Sengonca, C. & Drescher, K. (2001) Laboratory studies on the suitability of Thrips tabaci Lindeman (Thysanoptera, Thripidae) as prey for the development, longevity, reproduction and predation of four predatory mite species of the genus Amblyseius (Acari, Phytoseiidae). Zeitschrift fur Pflanzenkrankheiten und Pflanzenschutz, 108, 66–76.
Siscaro, G., Longo, S. & Lizzio, S. (1999) Ruolo degli entomofagi di Aonidiella aurantii (Maskell) (Homoptera, Diaspididae) in agrumeti siciliani. Phytophaga, 9, 41–52.
Stathakis, T.I. & Papadoulis, G.T. (2012) New records of phytoseiid mites from Greece with description of Typhlodromus (Anthoseius) creticus sp. nov. (Acari: Phytoseiidae). International Journal of Acarology, 38, 116–119.
http://dx.doi.org/10.1080/01647954.2011.595084Stathakis, T.I., Kapaxidi, E.V. & Papadoulis, G.Th. (2012) A new record and two new species of the genus Typhlodromus Scheuten (Acari: Phytoseiidae) from Greece. International Journal of Acarology, 38, 605–611.
http://dx.doi.org/10.1080/01647954.2012.715671Stathakis, T.I., Kapaxidi, E.V. & Papadoulis, G.Th. (2013) Two new species of the genus Neoseiulus Hughes (Acari: Phytoseiidae) from Greece with re-description of Neoseiulus leucophaeus (Athias-Henriot). Zootaxa, 3681 (5), 563–572.
http://dx.doi.org/10.11646/zootaxa.3681.5.5Swirski, E. & Amitai, S. (1982) Notes on predacious mites (Acarina: Phytoseiidae) from Turkey, with description of the male of Phytoseius echinus Wainstein and Arutunjan. Israel Journal of Entomology, 16, 55–62.
Swirski, E. & Amitai, S. (1984) Notes on phytoseiid mites (Mesostigmata: Phytoseiidae) from the Mediterranean littoral zone of Israel, with a description of a new species of Typhloctonus. Israel Journal of Entomology, 18, 71–82.
Swirski, E. & Amitai, S. (1997a) Notes on phytoseiid mites (Mesostigmata: Phytoseiidae) of Mt. Carmel (Israel), with descriptions of two new species. Israel Journal of Entomology, 31, 1–20.
Swirski, E. & Amitai, S. (1997b) Annotated list of phytoseiid mites (Mesostigmata: Phytoseiidae) in Israel. Israel Journal of Entomology, 31, 21–46.
Tixier, M.-S. (2012) Approaches to assess intraspecific variations of morphological continuous characters: the case study of the family Phytoseiidae (Acari: Mesostigmata). Cladistics, 28, 489–502.
http://dx.doi.org/10.1111/j.1096-0031.2012.00394.xTixier, M.-S. (2013) Statistical approaches to assess interspecific differences for morphological continuous characters: the case study of the family Phytoseiidae (Acari: Mesostigmata). Zoologica Scripta, 42, 327–334.
http://dx.doi.org/10.1111/zsc.12004Tixier, M.-S., Kreiter, S. & Auger, P. (2000) Colonization of vineyards by phytoseiid mites: their dispersal patterns in the plot and their fate. Experimental and Applied Acarology, 24, 191–211.
http://dx.doi.org/10.1023/A:1006332422638Tixier, M.-S., Kreiter, S. & de Moraes, G.J. (2008b) Biogeographic distribution of the mites of the family Phytoseiidae (Acari: Mesostigmata). Biological Journal of the Linnean Society, 93, 845–856.
Tixier, M.-S., Kreiter, S., Auger, P. & Weber, M. (1998) Colonization of Languedoc vineyards by phytoseiid mites (Acari: Phytoseiidae): influence of wind and crop environment. Experimental and Applied Acarology, 22, 523–542.
http://dx.doi.org/10.1023/A:1006085723427Tixier, M.-S., Kreiter, S., Croft, B.A. & Cheval, B. (2008a) Kampimodromus aberrans (Acari: Phytoseiidae) from USA: morphological and molecular assessment of its identity. Bulletin of Entomological Research, 98, 125–134.
Tixier, M.-S., Kreiter, S., Douin, M. & Moraes, G.J. (2012) Rates of description of Phytoseiidae (Acari: Mesostigmata): space, time and body size variations. Biodiversity and Conservation, 21, 993–1013.
http://dx.doi.org/10.1007/s10531-012-0235-0Tixier, M.-S., Kreiter, S., Klaric, V. & Duso, C. (2010) Phytoseiid mite species from Croatia, with description of a new species of the genus Typhlodromus (Typhlodromus) Scheuten. Annals of the Entomological Society of America, 103, 165–180.
http://dx.doi.org/10.1603/AN09092Tixier, M.-S., Kreiter, S., Allam, L., Ouahbi, A. & Hmimina, H. (2003) Phytoseiid and tetranychid mites (Acari: Mesostigmata) of some Moroccan crops. Acarologia, 43, 87–97.
Tixier, M.-S., Kreiter, S., Barbar, Z., Ragusa, S. & Cheval, B.(2006) The status of two cryptic species: Typhlodromus exhilaratus Ragusa and Typhlodromus phialatus Athias-Henriot (Acari: Phytoseiidae): consequences for taxonomy. Zoologica Scripta, 35, 115–122.
http://dx.doi.org/10.1111/j.1463-6409.2006.00222.xTixier, M.S., Tsolakis, H., Ragusa, S., Poinso, A., Ferrero, M., Okassa, M. & Kreiter, S. (2011) An integrative taxonomical approach demonstrates the synonymy between Cydnodromus idaeus and C. picanus (Acari: Phytoseiidae). Invertebrate Systematics, 25, 273–281.
http://dx.doi.org/10.1071/IS11025Tseng, Y.H. (1976) Systematics of the mite family Phytoseiidae from Taiwan, with a revised key to genera of the world (II). Journal of the Agricultural Association of China, 94, 85–128.
Tsolakis, H., Sinacori, M. & Ragusa, S. (2013) Predation of Typhlodromus longilaterus Athias-Henriot (Parasitiformes, Phytoseiidae) females on eggs and juveniles of the tetranychid mites Tetranychus urticae (Koch) and Panonychus citri (McGregor) (Acariformes, Tetranychidae). IOBC/WPRS Bulletin, 93, 129–132.
Tsoukanas, V.I., Papadopoulos, C.D., Fantinou, A.A. & Papadoulis, G.T. (2006) Temperature-dependent development and life table of Iphiseius degenerans (Acari: Phytoseiidae). Environmental Entomology, 35, 212–218.
http://dx.doi.org/10.1603/0046-225X-35.2.212Tuovinen, T. & Lindqvist, I. (2010) Maintenance of predatory phytoseiid mites for preventive control of strawberry tarsonemid mite Phytonemus pallidus in strawberry plant propagation. Biological Control, 54, 119–125.
http://dx.doi.org/10.1016/j.biocontrol.2010.04.006Tuttle, D.M. & Muma, M.H. (1973) Phytoseiidae (Acarina: Mesostigmata) inhabiting agricultural and other plants in Arizona. Agricultural Experiment Station Technical Bulletin, University of Arizona, Tucson, USA, 208, 1–55.
Ueckermann, E.A. & Loots, G.C. (1988) The African species of the subgenera Anthoseius De Leon and Amblyseius Berlese (Acari: Phytoseiidae). Entomology Memoir, Department of Agriculture and Water Supply, Republic of South Africa, South Africa, 73, 1–168.
Vacante, V. & Nucifora, A. (1987) Gli Acari degli agrumi in Italia. II. I Gamasidi (Phytoseiidae Berlese e Ascidae Voigts & Oudemans): chiave per il riconoscimento dei e delle specie e note di biologia. Bolletino de Zoologia Agraria e Bachicoltura, Series II, 19, 145–184.
Van der Merwe, G.G. (1968) A taxonomic study of the family Phytoseiidae (Acari) in South Africa with contributions to the biology of two species. Entomology Memoirs, South Africa Department of Agricultural Technical Services, South Africa, 18, 1–198.
Van Houten, Y.M. & Van Stratum, P. (1995) Control of western flower thrips on sweet pepper in winter with Amblyseius cucumeris (Oudemans) and A. degenerans Berlese. In: Parker, B.L., Skinner, M. & Lewis, T. (Eds.), Thrips Biology and Management. NATO ASI Series A: Life Sciences. Vol. 276. Plenum Press, New York, pp. 245–248.
http://dx.doi.org/10.1007/978-1-4899-1409-5_37Vantornhout, I., Minnaert, H.L., Tirry, L. & De Clercq, P. (2004) Effect of pollen, natural prey and factitious prey on the development of Iphiseius degenerans. BioControl, 49, 627–644.
http://dx.doi.org/10.1007/s10526-004-5280-5Villiers, M. & Pringle, K.L. (2011) The presence of Tetranychus urticae (Acari: Tetranychidae) and its predators on plants in the ground cover in commercially treated vineyards. Experimental and Applied Acarology, 53, 121–137.
http://dx.doi.org/10.1007/s10493-010-9391-7Wainstein, B.A. (1958) New species of mites of the genus Typhlodromus (Parasitiformes: Phytoseiidae) from Georgia. Soobshcheniya Akademii Nauk Gruzinskoy SSR, Georgia, 21, 201–207.
Wainstein, B.A. (1959) New subgenus and species of the genus Phytoseius Ribaga, 1902 (Phytoseiidae: Parasitiformes). Zoologicheskii Zhurnal, 38, 1361–1365.
Wainstein, B.A. (1960) New species and subspecies of the genus Typhlodromus Scheuten (Parasitiformes, Phytoseiidae) of the USSR fauna. Zoologicheskii Zhurnal, 39, 683–690.
Wainstein, B.A. (1962) Révision du genre Typhlodromus Scheuten, 1857 et systématique de la famille des Phytoseiidae (Berlese 1916) (Acarina: Parasitiformes). Acarologia, 4, 5–30.
Wainstein, B.A. (1973) Predatory mites of the family Phytoseiidae (Parasitiformes) of the fauna of the Moldavian SSR. Fauna i Biologiya Nasekomykh Moldavii, Akademiya Nauk Moldavskoy SSR, Institut Zoologii, Moldova, 12, 176–180.
Wainstein, B.A. (1975) Predatory mites of the family Phytoseiidae (Parasitiformes) of Yaroslavl Province. Entomologicheskoe Obozrenie, 54, 914–922.
Wainstein, B.A. (1977) A contribution to the fauna of the family Phytoseiidae (Parasitiformes) in Australia. Zoologicheskii Zhurnal, 56, 1413–1416.
Wainstein, B.A. & Arutunjan, E.S. (1967) New species of predaceous mites of the genera Typhlodromus Scheuten and Paraseiulus Muma (Parasitiformes, Phytoseiidae). Zoologicheskii Zhurnal, 46, 1764–1770.
Wainstein, B.A. & Vartapetov, S.G. (1973) Predatory mites of the family Phytoseiidae (Parasitiformes) of Adzharskaya ASSR. Akademiya Nauk Armyanskoy SSR, Biologicheskiy Zhurnal Armenii, 26, 102–105.
Westerboer, I. & Bernhard, F. (1963) Die Familie Phytoseiidae Berlese 1916. In: Stammer, H. (Ed.), Beitrage zur Systematik und Okologie mitteleuropaischer Acarina, Band II, Mesostigmata I, pp. 451–791.