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Issue: Vol. 4069 No. 1: 27 Jan. 2016
Type: Monograph
Published: 2016-01-27
DOI: 10.11646/zootaxa.4069.1.1
Page range: 1–183
Abstract views: 241
PDF downloaded: 3

Revision and phylogenetic analysis of the orb-weaving spider genus Glenognatha Simon, 1887 (Araneae, Tetragnathidae)

Departamento de Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, SP, 05508-090, Brazil.
Laboratório Especial de Coleções Zoológicas, Instituto Butantan, São Paulo, SP, 05503-900, Brazil.
Araneae parsimony taxonomy morphology Dyschiriognatha Tetragnathinae

Abstract

A taxonomic revision and phylogenetic analysis of the spider genus Glenognatha Simon, 1887 is presented. This analysis is based on a data set including 24 Glenognatha species plus eight outgroups representing three related tetragnathine genera and one metaine as the root. These taxa were scored for 78 morphological characters. Parsimony was used as the optimality criterion and a sensitivity analysis was performed using different character weighting concavities. Seven unambiguous synapomorphies support the monophyly of Glenognatha. Some internal clades within the genus are well-supported and its relationships are discussed. Glenognatha as recovered includes 27 species, four of them only known from males. A species identification key and distribution maps are provided for all. New morphological data are also presented for thirteen previously described species. Glenognatha has a broad distribution occupying the Neartic, Afrotropic, Indo-Malaya, Oceania and Paleartic regions, but is more diverse in the Neotropics. The following eleven new species are described: G. vivianae n. sp., G. caaguara n. sp., G. boraceia n. sp. and G. timbira n. sp. from southeast Brazil, G. caparu n. sp., G. januari n. sp. and G. camisea n. sp. from the Amazonian region, G. mendezi n. sp., G. florezi n. sp. and G. patriceae n. sp. from northern Andes and G. gouldi n. sp. from Southern United States and central Mexico. Females of G. minuta Banks, 1898, G. gaujoni Simon, 1895 and G. gloriae (Petrunkevitch, 1930) and males of G. globosa (Petrunkevitch, 1925) and G. hirsutissima (Berland, 1935) are described for the first time. Three new combinations are proposed in congruence with the phylogenetic results: G. argyrostilba (O. P.-Cambridge, 1876) n. comb., G. dentata (Zhu & Wen, 1978) n. comb. and G. tangi (Zhu, Song & Zhang, 2003) n. comb., all previously included in Dyschiriognatha Simon, 1893. The following taxa are newly synonymized: Dyschiriognatha montana Simon, 1897, Glenognatha mira Bryant, 1945 and Glenognatha maelfaiti Baert, 1987 with Glenognatha argyrostilba (Pickard-Cambridge, 1876) and Glenognatha centralis Chamberlin, 1925 with Glenognatha minuta Banks, 1898.

 

References

  1. Agnarsson, I. (2004) Morphological phylogeny of cobweb spiders and their relatives (Araneae, Araneoidea, Theridiidae). Zoological Journal of the Linnean Society, 141 (4), 447–626.

    http://dx.doi.org/10.1111/j.1096-3642.2004.00120.x

    Agnew, C.W., Dean, D.A. & Smith, J.W. (1985) Spiders collected from peanuts and non-agricultural habitats in Texas west cross-timbers. The Southwestern Naturalist, 30 (1), 1–12.

    Álvarez-Padilla, F. (2007) Systematics of the spider genus Metabus O. P.-Cambridge, 1899 (Araneoidea: Tetragnathidae) with additions to the tetragnathid fauna of Chile and comments on the phylogeny of Tetragnathidae. Zoological Journal of the Linnean Society, 151 (2), 285–335.

    http://dx.doi.org/10.1111/j.1096-3642.2007.00304.x

    Álvarez-Padilla, F. & Hormiga, G. (2008) A protocol for digesting internal soft tissues and mounting spiders for scanning electron microscopy. The Journal of Arachnology, 35 (3), 538–542.

    http://dx.doi.org/10.1636/Sh06-55.1

    Álvarez-Padilla, F., Dimitrov, D., Giribet, G. & Hormiga, G. (2009) Phylogenetic relationships of the spider family Tetragnathidae (Araneae, Araneoidea) based on morphological and DNA sequence data. Cladistics, 25 (2), 109-–146.

    http://dx.doi.org/10.1111/j.1096-0031.2008.00242.x

    Álvarez-Padilla, F. & Hormiga, G. (2011) Morphological and phylogenetic atlas of the orb-weaving spider family Tetragnathidae (Araneae: Araneoidea). Zoological Journal of the Linnean Society, 162 (4), 713–879.

    http://dx.doi.org/10.1111/j.1096-3642.2011.00692.x

    Armendano, A. & González, A. (2010) Comunidad de arañas (Arachnida, Araneae) del cultivo de alfalfa (Medicago sativa) en Buenos Aires, Argentina. Revista de Biología Tropical, 58 (2), 757–767.

    Armendano, A. & González, A. (2011) Spider fauna associated with wheat crops and adjacent habitats in Buenos Aires, Argentina. Revista Mexicana de Biodiversidad, 82 (4), 1176–1182.

    Arnedo, M.A., Hormiga, G. & Scharff, N. (2009) Higher-level phylogenetics of linyphiid spiders (Araneae, Linyphiidae) based onmorphological and molecular evidence. Cladistics, 25 (3), 231--–262.

    http://dx.doi.org/10.1111/j.1096-0031.2009.00249.x

    Baert, L. (1987) Spiders of the Galápagos Islands. Part IV. Miscellaneous families II. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 57, 141–155.

    Baert, L., Desender, K. & Maelfait, J.-P. (1991) Spider communities of Isla Santa Cruz (Galapagos, Ecuador). Journal of Biogeography, 18 (3), 333–340.

    Balfour, R.A. & Rypstra, A.L. (1998) The influence of habitat structure on spider density in a no-till soybean agroecosystem. The Journal of Arachnology, 26, 221–226.

    Banks, N. (1896) New North American spiders and mites. Transactions of the American Entomological Society, 23, 57–77.

    Banks, N. (1898) Arachnida from Baja California and other parts of Mexico. Proceedings of the California Academy of Sciences, 1, 205–308.

    Banks, N. (1904) The Arachnida of Florida. Proceedings of the Academy of natural Sciences of Philadelphia, 56, 120–147.

    Banks, N. (1913) Notes on the types of some American spiders in European collections. Proceedings of the Academy of natural Sciences of Philadelphia, 65, 177-–188.

    Banks, N. (1929) Spiders from Panama. Bulletin of the Museum of Comparative Zoology, 69, 53–96.

    Barrion, A.T. & Litsinger, J.A. (1995) Riceland Spiders of South and Southeast Asia. CAB International, Wallingford, UK, xix + 700 pp.

    Barrion, A.T., Villarreal, S.S., Catindig, J.L.A., Cai, D., Yuan, Q.H. & Heong, K.L. (2012) The spider fauna in the rice agricultural landscape of Hainan Island, China: Composition, abundance and feeding structure. Asia Life Sciences, 21 (2), 625-–651.

    Barrows, W.M. (1919) The taxonomic position of Mysmena bulbifera (Glenognatha bulbifera) Banks, with some observations of its habits. Ohio Journal of Science, 19 (3), 210–212.

    Benoit, P.L.G. (1977) Fam. Tetragnathidae. In: La faune terrestre de l'île de Saite-Hélène IV. Annales Musée Royal de l'Afrique Centrale, 220, 156-–162.

    Berland, L. (1929) Araignées (Araneida). In: Insects of Samoa and other Samoan terrestrial Arthropoda. London, 8, 35–78.

    Berland, L. (1933) Araignées des Iles Marquises. Bernice Pauahi Bishop Museum Bulletin, 114, 39–70.

    Berland, L. (1935) Nouvelles araignées marquisiennes. Bernice Pauahi Bishop Museum Bulletin, 142, 31–63.

    Berland, L. (1942) Polynesian spiders. Occasional Papers of the Bernice Pauahi Bishop Museum, 17 (1), 1–24.

    Birkhofer, K., Scheu, S. & Wise, D.H. (2007) Small-scale spatial pattern of web-building spiders (Araneae) in alfalfa: relationship to disturbance from cutting, prey availability, and intraguild interactions. Environmental Entomology, 36 (4), 801–810.

    http://dx.doi.org/10.1603/0046-225X(2007)36[801%3ASSPOWS]2.0.CO%3B2

    de Bivort, B.L., Clouse, R.M. & Giribet, G. (2010) A morphometrics-based phylogeny of the temperate Gondwanan mite harvestmen (Opiliones, Cyphophthalmi, Petallidae). Journal of Zoological Systematics and Evolutionary Research, 48 (4), 294–309.
    http://dx.doi.org/10.1111/j.1439-0469.2009.00562.x

    Bodkhe, A.K., Manthen, S.V. & Tanikawa, A. (2014) A new species of Dyschiriognatha (Araneae: Tetragnathidae) from India. Indian Journal of Arachnology, 3 (1), 28-–34

    Bosmans, R. & Bosselaers, J. (1994) Spiders of the genera Pachygnatha, Dyschiriognatha and Glenognatha (Araneae, Tetragnathidae), with a revision of the Afrotropical species. Zoologica Scripta, 23 (4), 325–352.
    http://dx.doi.org/10.1111/j.1463-6409.1994.tb00392.x

    Breene, R.G., Dean, D.A., Nyffeler, M. & Edwards, G.B. (1993) Biology, Predation Ecology, and Significance of Spiders in Texas Cotton Ecosystems with a Key to Species. Texas Agriculture Experiment Station, College Station, 115 pp.

    Bremer, K. (1988) The limits of aminoacid sequence data in angiosperm phylogenetic reconstruction. Evolution, 42 (4), 795–803.

    Bryant, E.B. (1945) The Argiopidae of Hispaniola. Bulletin of the Museum of Comparative Zoology, 95, 357-–422.

    Cabra-García, J., Brescovit, A.D. & Hormiga, G. (2014) Female genital morphology in the secondarily haplogyne spider genus Glenognatha Simon, 1887 (Araneae, Tetragnathidae), with comments on its phylogenetic significance. Journal of Morphology, 275 (9), 1027–1040.

    http://dx.doi.org/10.1002/jmor.20280

    Caporiacco, L. di (1947) Diagnosi preliminari de specie nuove di aracnidi della Guiana Brittanica raccolte dai professori Beccari e Romiti. Monitore zoologico italiano, 56, 20–34.

    Caporiacco, L. di (1948) Arachnida of British Guiana collected in 1931 and 1936 by Professors Beccari and Romiti. Proceedings of the Zoological Society of London, 118 (3), 607-–747.

    http://dx.doi.org/10.1111/j.1096-3642.1948.tb00402.x

    Chamberlin, R.V. (1925) Diagnoses of new American Arachnida. Bulletin of the Museum of Comparative Zoology, 67, 209–248.

    Chamberlin, R.V. & Ivie, W. (1936) New spiders from Mexico and Panama. Bulletin of the University of Utah, 27 (5), 1–103.

    Chapman, E.G., Schmidt, J.M., Welch, K.D. & Harwood, J.D. (2013) Molecular evidence for dietary selectivity and pest suppression potential in an epigeal spider community in winter wheat. Biological Control, 65 (1), 72–86.

    http://dx.doi.org/10.1016/j.biocontrol.2012.08.005

    Chen, X.E. & Gao, J.C. (1990) The Sichuan farmland spiders in China. Sichuan Science and Technology Publishing House, Chengdu, 226 pp.

    Coddington, J.A. (1983) A Temporary Slide Mount Allowing Precise Manipulation of Small Structures. Verhandlungen Naturwissenchaften Vereins Hamburg, 26, 291–292.

    Coddington, J.A. (1990) Ontogeny and homology in the male palpus of orb-weaving spiders and their relatives, with comments on phylogeny (Araneoclada: Araneoidea, Deinopoidea). Smithsonian Contributions to Zoology, 496, 1–52.

    Crosby, C.R. & Bishop, S.C. (1928) Araneae. In: A list of the insects of New York. Memoirs of the Cornell University Agricultural Experiment Station, 101, 1034–1074.

    Crosby, C.R. & Bishop, S.C. (1933) American spiders: Erigonae, males with cephalic pits. Annals of the Entomological Society of America, 26, 105–172.

    Danielson-François, A.M. (2006) Natural History of Glenognatha emertoni (Araneae, Tetragnathidae): mating behavior and sperm release in an haplogyne. The Journal of Arachnology, 34 (2), 387--–398.

    http://dx.doi.org/10.1636/S04-46.1

    Daza, J.D., Abdala, V., Arias, J.S., García-López, D. & Ortiz, P. (2012) Cladistic Analysis of Iguania and a Fossil Lizard from the Late Pliocene of Northwestern Argentina. Journal of Herpetology, 46 (1), 104–119.

    http://dx.doi.org/10.1670/10-112

    Dean, D.A. & Sterling, W.L. (1985) Size and phenology of ballooning spiders at two locations in eastern Texas. The Journal of Arachnology, 13 (1), 111–120.

    Dierkens, M. & Charlat, S. (2011) Contribution à la connaissance des araignées des îles de la Société (Polynésie française). Reveu Arachnologique, 17 (5), 63–81.

    Dimitrov, D., Alvarez-Padilla, F. & Hormiga, G. (2007) The female genitalic morphology of the orb weaving spider genus Agriognatha (Araneae, Tetragnathidae). Journal of Morphology, 268 (9), 758–770.

    http://dx.doi.org/10.1002/jmor.10543

    Dimitrov, D. & Hormiga, G. (2009) Revision and cladistic analysis of the orbweaving spider genus Cyrtognatha Keyserling, 1881 (Araneae, Tetragnathidae). Bulletin of the American Museum of Natural History, 317, 1–139.

    http://dx.doi.org/10.1206/317.1

    Dimitrov, D. & Hormiga, G. (2011) An extraordinary new genus of spiders from Western Australia with an expanded hypothesis on the phylogeny of Tetragnathidae (Araneae). Zoological Journal of the Linnean Society, 161 (4), 735–768.

    http://dx.doi.org/10.1111/j.1096-3642.2010.00662.x

    Döbel, H.T., Denno, R.F. & Coddington, J.A. (1990) Spider (Araneae) community structure in an intertidial salt marsh: effects of vegetation structure and tidal flooding. Environmental Entomology, 19 (5), 1356–1370.

    http://dx.doi.org/10.1093/ee/19.5.1356

    Draney, M.L. (1997) Ground-layer spiders (Araneae) of a Georgia piedmont floodplain agroecosystem: species list, phenology and habitat selection. The Journal of Arachnology, 25 (3), 333–351.

    Dupo, A.L.B. & Barrion, A.T. (2009) Taxonomy and general biology of delphacid planthoppers in rice agroecosystems. In: Heong, K.L. & Hardy, B. (Eds.), Planthoppers: new threats to the sustainability of intensive rice production systems in Asia. International Rice Research Institute, Los Baños, pp. 3–156. [Philippines]

    Eberhard, W.G. (1982) Behavioral characters for the higher classification of orb-weaving spiders. Evolution, 36 (5), 1067–1095.

    Eberhard, W.G. (2010) Possible functional significance of spigot placement on the spinnerets of spiders. The Journal of Arachnology, 38 (3), 407–414.

    http://dx.doi.org/10.1636/B09-97.1

    Eberhard, W.G & Huber, B.A. (2010) Spider genitalia: precise maneuvers with a numb structure in a complex lock. In: Leonard, J.L. & Córdoba-Aguilar, A. (Eds.), Evolution of primary sexual characters in animals. Oxford University Press, Oxford, pp. 249–284.

    Edwards, R.L. & Senske, C.W. (2001) Observations on the breeding behavior of the spider Glenognatha heleios (Araneae: Tetragnathidae). Entomological News, 112, 71–72.

    Fitch, W.M. (1971) Toward defining the course of evolution: minimum change for a specific tree topology. Systematic Zoology, 20 (4), 404–416.

    http://dx.doi.org/10.1093/sysbio/20.4.406

    Frick, H. & Scharff, N. (2014) Phantoms of Gondwana? - phylogeny of the spider subfamily Mynogleninae (Araneae: Linyphiidae). Cladistics, 30 (1), 67–106.

    http://dx.doi.org/10.1111/cla.12025

    Gavish-Regev, E., Hormiga, G. & Scharff, N. (2013) Pedipalp sclerite homologies and phylogenetic placement of the spider genus Stemonyphantes (Linyphiidae, Araneae) and its implications for linyphiid phylogeny. Invertebrate Systematics, 27 (1), 38–52.

    http://dx.doi.org/10.1071/IS12014

    Gillespie, R.G. (2002) Hawaiian spiders of the genus Tetragnatha: IV new, small species in the spiny leg clade. The Journal of Arachnology, 30 (1), 159–172.

    http://dx.doi.org/10.1636/0161-8202(2002)030[0159%3AHSOTGT]2.0.CO%3B2

    Gillespie, R.G., Croom, H.B. & Hasty, G.L. (1997) Phylogenetic Relationships and Adaptive Shifts among Major Clades of Tetragnatha Spiders (Araneae: Tetragnathidae) in Hawaii. Pacific Science, 51 (4), 380–394.

    Gillespie, R.G., Claridge, E.M. & Goodcare, S.L. (2008) Biogeography of the fauna of French Polynesia: diversification within and between a series of hot spot archipelagos. Philosophical Transactions of the Royal Society B, 363 (1508), 3335–3346.

    http://dx.doi.org/10.1098/rstb.2008.0124

    Giribet, G. (2003) Stability in phylogenetic formulations and its relationship to nodal support. Systematic Biology, 52 (4), 554–564.

    http://dx.doi.org/10.1080/10635150390223730

    Goloboff, P.A. (1993) Estimating character weights during tree search. Cladistics, 9 (1), 93–91.

    http://dx.doi.org/10.1111/j.1096-0031.1993.tb00209.x

    Goloboff, P.A., Farris, J.S., Källesjö, M., Oxelman, B., Ramírez, M.J. & Szumik, C.A. (2003) Improvements to resampling measures of group support. Cladistics, 19 (4), 324–332.

    http://dx.doi.org/10.1111/j.1096-0031.2003.tb00376.x

    Goloboff, P.A, Farris, J.S. & Nixon, K.C. (2008a) TNT, a free program for phylogenetic analysis. Cladistics, 24 (5), 774–786.

    http://dx.doi.org/10.1111/j.1096-0031.2008.00217.x

    Goloboff, P.A., Carpenter, J.M., Arias, J.S. & Miranda Esquivel, D.R. (2008b) Weighting against homoplasy improves phylogenetic analysis of morphological data sets. Cladistics, 24 (5), 758–773.

    http://dx.doi.org/10.1111/j.1096-0031.2008.00209.x

    Goodman, M., Olson, C.B., Beeber, J.E. & Czelusniak, J. (1982) New perspectives in the molecular biological analysis of mammalian phylogeny. Acta Zoologica Fennica, 169, 19–35.

    Grandjean, F. (1949) Observation et conservation des très petits arthropodes. Bulletin du Muséum National d’Histoire naturelle, 21, 363–370.

    Grant, T. & Kluge, A. (2008a) Credit where credit is due: The Goodman-Bremer support metric. Molecular Phylogenetics and Evolution, 49 (1), 405–406.

    http://dx.doi.org/10.1016/j.ympev.2008.04.023

    Grant, T. & Kluge, A. (2008b) Clade support measures and their adequacy. Cladistics, 24, 1051–1064.

    http://dx.doi.org/10.1111/j.1096-0031.2008.00231.x

    Griswold, C.E., Coddington, J.A., Hormiga, G. & Scharff, N. (1998) Phylogeny of the orb-web building spiders (Araneae, Orbiculariae: Deinopoidea, Araneoidea). Zoological Journal of the Linnaean Society, 123 (1), 1–99.

    http://dx.doi.org/10.1111/j.1096-3642.1998.tb01290.x

    Griswold, C.E., Ramírez, M.J., Coddington, J.A. & Platnick, N.I. (2005) Atlas of phylogenetic data for Entelegyne spiders (Araneae: Araneomorphae: Entelegynae) with comments on their phylogeny. Proceeding of the California Academy of Sciences, 56, 1–324.

    Heiss, J.S. & Meisch, M.V. (1985) Spiders (Araneae) associated with rice in Arkansas with notes on species compositions of populations. The Southwestern Naturalist, 30 (1), 119–127.

    Holm, Å. (1969) A new tetragnathid species from the island St-Helena (Araneae). Revue de Zoologie et de Botanique Africaines, 80, 62–66.

    Hormiga, G. (1994) A revision and cladistics analysis of the spider family Pimoidae (Araneae: Araneoidea). Smithsonian Contributions to Zoology, 549, 1–105.

    Hormiga, G. (2000) Higher level phylogenetics of erigonine spiders (Araneae, Linyphiidae, Erigoninae). Smithsonian Contributions to Zoology, 609, 1–160.

    Hormiga, G. (2002) Orsonwelles, a new genus of giant linyphiid spiders (Araneae) from the Hawaiian islands. Invertebrate Systematics, 16, 369–448.

    http://dx.doi.org/10.1071/IT01026

    Hormiga, G. & Döbel, H.G. (1990) A new Glenognatha (Araneae, Tetragnathidae) from New Jersey, with redescriptions of G. centralis and G. minuta. The Journal of Arachnology, 18 (2), 195–204.

    Hormiga, G., Eberhard, W.G. & Coddington, J.A. (1995) Web-construction behaviour in Australian Phonognatha and the phylogeny of nephiline and tetragnathid spiders (Araneae: Tetragnathidae). Australian Journal of Zoology, 43 (4), 313–364.

    http://dx.doi.org/10.1071/ZO9950313

    Hormiga, G., Alvarez-Padilla, F. & Benjamin, S.P. (2007) First records of extant Hispaniolan spiders of the families Mysmenidae, Symphytognathidae, and Ochyroceratidae (Araneae), including a new species of Ochyrocera. American Museum Novitates, 3577, 1–21.

    http://dx.doi.org/10.1206/0003-0082(2007)3577[1:FROEHS]2.0.CO;2

    Hu, J.L. (1984) The Chinese spiders collected from the fields and the forests. Press of Science and Techniques, Tianjin, 482 pp.

    Huber, B.A. & Senglet, A. (1997) Copulation with contralateral insertion in entelegyne spiders (Araneae: Entelegynae: Tetragnathidae). Netherlands Journal of Zoology, 47 (1), 99–102.

    http://dx.doi.org/10.1163/156854297X00274

    Izquierdo, M.A. & Ramírez, M.J. (2008) Two new spider species of the genera Aysenia and Aysenoides from southern Chile and Argentina: description and phylogenetic relationships (Araneae: Anyphaenidae, Amaurobioidinae). Zootaxa, 1861, 29–43.

    Kaston, B.J. (1945) New Micryphantidae and Dictynidae with notes on other spiders. American Museum Novitates, 1292, 1–14.

    Kaston, B.J. (1948) Spiders of Connecticut. Bulletin of the Connecticut State Geological and Natural History Survey, 70, 1–874.

    Keyserling, E. (1883) Neue Spinnen aus Amerika. IV. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 32, 195–226.

    Keyserling, E. (1884) Neue Spinnen aus America. V. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 33, 649–684.

    Kuntner, M., Coddington, J.A. & Hormiga, G. (2008) Phylogeny of extant nephilid orb-weaving spiders (Araneae, Nephilidae): testing morphological and ethological homologies. Cladistics, 24 (2), 147–217.

    http://dx.doi.org/10.1111/j.1096-0031.2007.00176.x

    Kuntner, M., Arnedo, M.A., Trontelj, P., Lokovšek, T. & Agnarsson, I. (2013) A molecular phylogeny of nephilid spiders: Evolutionary history of a model lineage. Molecular Phylogenetics and Evolution, 69 (3), 961–979.

    http://dx.doi.org/10.1016/j.ympev.2013.06.008

    Latreille, P.A. (1804) Histoire naturelle générale et particulière des Crustacés et des Insectes. Tome 7. F. Dufart, Paris, 162 pp. [pp. 144–305]

    Levi, H.W. (1980) The orb-weaver genus Mecynogea, the subfamily Metinae and the genera Pachygnatha, Glenognatha and Azilia of the subfamily Tetragnathinae north of Mexico (Araneae: Araneidae). Bulletin of the Museum of Comparative Zoology, 149, 1–75.

    Levi, H.W. (1986) The Neotropical orb-weaver genera Chrysometa and Homalometa (Araneae: Tetragnathidae). Bulletin of the Museum of Comparative Zoology, 151, 91–215.

    Litsinger, J.A., Alviola, A.L., Dela Cruz, C.G., Canapi, B.L., Batay-An III, E.H. & Barrion, A.T. (2006) Rice white stemborer Scirpophaga innotata (Walker) in southern Mindanao, Philippines. II. Synchrony of planting and natural enemies. International Journal of Pest Management, 52 (1), 23–37.

    http://dx.doi.org/10.1080/09670870600552463

    Lopardo, L. (2005) Phylogenetic revision of the genus Negayan (Araneae, Anyphaenidae, Amaurobioidinae). Zoologica Scripta, 34 (3), 245–277.

    http://dx.doi.org/10.1111/j.1463-6409.2005.00194.x

    Lopardo, L., Giribet, G. & Hormiga, G. (2011) Morphology to the rescue: molecular data and the signal of morphological characters in combined phylogenetic analyses—a case study from mysmenid spiders (Araneae, Mysmenidae), with comments on the evolution of web architecture. Cladistics, 27 (3), 278–330.

    http://dx.doi.org/10.1111/j.1096-0031.2010.00332.x

    Lopardo, L. & Hormiga, G. (2015) Out of the twilight zone: phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea). Zoological Journal of the Linnean Society, 173 (3), 527–786.

    http://dx.doi.org/10.1111/zoj.12199

    Maddison, W.P. & Maddison, D.R. (2015) Mesquite: a modular system for evolutionary analysis. Version 3.02. Available from: http://mesquiteproject.org (accessed 15 January 2015)

    Magalhães, I.L.F. & Santos, A.J. (2012) Phylogenetic analysis of Micrathena and Chaetacis spiders (Araneae: Araneidae) reveals multiple origins of extreme sexual size dimorphism and long abdominal spines. Zoological Journal of the Linnean Society, 166 (1), 14–53.

    http://dx.doi.org/10.1111/j.1096-3642.2012.00831.x

    Mansour, F., Ross, J.W, Edwards, G.B., Whitcomb, W.H. & Richman, D.B. (1982) Spiders of Florida citrus groves. The Florida Entomologist, 65 (4), 514--–522.

    Marples, B.J. (1955) Spiders from western Samoa. Journal of the Linnean Society of London, Zoology, 42 (287), 453–504

    http://dx.doi.org/10.1111/j.1096-3642.1955.tb02217.x

    McCook, H.C. (1894) American spiders and their spinningwork. Vol III. Published by the Author, Academy of Natural Sciences of Philadelphia, Philadelphia, 285 pp.

    Mello-Leitão, C.F. de (1944) Arañas de la provincia de Buenos Aires. Revista del Museo de La Plata, New Series, Zoology, 3, 311–393.

    Mendes, A.C. (2011) Phylogeny and taxonomic revision of Heteropachylinae (Opiliones: Laniatores: Gonyleptidae). Zoological Journal of the Linnean Society, 163 (2), 437–483.

    http://dx.doi.org/10.1111/j.1096-3642.2011.00706.x

    Miller, J.A. & Hormiga, G. (2004) Clade stability and the addition of data: a case study from erigonine spiders (Araneae: Linyphiidae, Erigoninae). Cladistics, 20 (5), 385–442.

    http://dx.doi.org/10.1111/j.1096-0031.2004.00033.x

    Mirande, J.M. (2009) Weighted parsimony phylogeny of the family Characidae (Teleostei: Characiformes). Cladistics, 25 (6), 574–613.

    http://dx.doi.org/10.1111/j.1096-0031.2009.00262.x

    Nixon, K.C. (2002) Winclada. Version 10.00.08. Published by the author, Ithaca, NY, USA. Available from: http://www.cladistics.com (accessed 15 January 2015)

    Ojanguren-Affilastro, A.A. & Ramírez, M.J. (2009) Phylogenetic analysis of the scorpion genus Brachistosternus (Arachnida, Scorpiones, Bothriuridae). Zoologica Scripta, 38 (2), 183–198.

    http://dx.doi.org/10.1111/j.1463-6409.2008.00367.x

    Okuma, C. (1987) A revision of the australasian species of the genus Tetragnatha (Araneae, Tetragnathidae). Esakia, 25, 37–96.

    Okuma, C. (1991) A new record of Japanese spider Dyschiriognatha dentata Chu et Wen. Heptathela, 5 (1), 15–18.

    Okuma, C., Kamal, N.Q., Hirashima, Y., Alam, M.Z. & Ogata, K. (1993) Illustrated Monograph of the Rice Field Spiders of Bangladesh. Institute of Postgraduate Studies in Agriculture (Salna, Gazipur, Bangladesh). Japan International Cooperation Agency Project Publication, 1, 1–93.

    Opell, B.D. (1979) Revision of the genera and tropical American species of the spider family Uloboridae. Bulletin of the Museum of Comparative Zoology, 148, 443–549.

    Opell, B.D. (1990) The relationship of book lung and tracheal systems in the spider family Uloboridae. Journal of Morphology, 206 (2), 211–216.

    http://dx.doi.org/10.1002/jmor.1052060207

    Pavesi, P. (1895) Esplorazione del Guiba e dei suoi affluenti compiuta dal Cap. Bottego. XVIII. Aracnidi. Annali del Museo Civico di Storia Naturale di Genova, 35, 491–537.

    Petrunkevitch, A. (1925a) Arachnida from Panama. Transactions of the Connecticut Academy of Arts and Sciences, 27, 51-–248.

    Petrunkevitch, A. (1925b) New Erigoninae from Tennessee. Journal of the New York Entomological Society, 33, 170–176.

    Petrunkevitch, A. (1930) The spiders of Porto Rico. Part two. Transactions of the Connecticut Academy of Arts and Sciences, 30, 159–356.

    Pickard-Cambridge, O. (1876) Catalogue of a collection of spiders made in Egypt, with descriptions of new species and characters of a new genus. Proceedings of the Zoological Society of London, 44 (1), 541–630.

    http://dx.doi.org/10.1111/j.1096-3642.1876.tb02595.x

    Platnick, N.I. (1993) Advances in spider taxonomy 1988-1991, with synonymies and transfers 1940-1980. The New York Entomological Society, New York, 846 pp.

    Ramírez, M.J. (1996) Descripción de la hembra de Glenognatha lacteovittata (Mello-Leitão), nueva combinación (Araneae, Tetargnathidae). Physis (Buenos Aires), 52, 35–37.

    Ramírez, M.J. (2003) The spider subfamily amaurobioidinae (Araneae: Anyphaenidae): A phylogenetic revision at generic level. Bulletin of the Museum of Natural History, 277, 1–262.
    http://dx.doi.org/10.1206/0003-0090(2003)277<0001:TSSAAA>2.0.CO;2

    Ramírez, M.J. (2014) The Morphology And Phylogeny Of Dionychan Spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1–374.

    http://dx.doi.org/10.1206/821.1

    Saaristo, M.I. (2003) Tetragnathid spiders of Seychelles (Araneae, Tetragnathidae). Phelsuma, 11, 13-–28.

    Saaristo, M.I. (2010) Araneae. In: Gerlach, J. & Marusik, Y. (Eds.), Arachnida and Myriapoda of the Seychelles islands. Siri Scientific Press, Mancheste, pp. 8–306.

    Sereno, P.C. (2007) Logical basis for morphological characters in phylogenetics. Cladistics, 23 (6), 565-–587.

    http://dx.doi.org/10.1111/j.1096-0031.2007.00161.x

    Scharaschkin, T. & Doyle, J.A. (2006) Character evolution in Anaxagorea (Annonaceae). The American Journal of Botany, 93 (1), 36–54.

    http://dx.doi.org/10.3732/ajb.93.1.36

    Sebastian, P.A., Mathew, M.J., Pathummal Beevi, S., Joseph, J. & Biju, C.R. (2005) The spider fauna of the irrigated rice ecosystems in central Kerala, India across different elevational ranges. The Journal of Arachnology, 33 (2), 247–255.

    http://dx.doi.org/10.1636/05-08.1

    Simon, E. (1887) Observation sur divers arachnides: synonymies et descriptions. Annales de la Société entomologique de France, Serie 6, Tome 7, 158–159, 167, 175–176, 186–187, 193–195.

    Simon, E. (1893) Arachnides de l'archipel Malais. Revue suisse de Zoologie, 1, 319–328.

    Simon, E. (1894) Histoire naturelle des araignées. Librairie Enciclopédique de Roret, Paris, 1084 pp.

    Simon, E. (1895) Etudes arachnologiques. 26e. XLI. Descriptions d'espèces et de genres nouveaux de l'ordre des Araneae. Annales de la Société Entomologique de France, 64, 131–160.

    Simon, E. (1897) On the spiders of the island of St Vincent. III. Proceedings of the zoological Society of London, 65 (4), 860-–890.

    http://dx.doi.org/10.1111/j.1096-3642.1898.tb01390.x

    Song, D.X., Zhu, M.S. & Chen, J. (1999) The Spiders of China. Hebei University of Science and Technology Publishing House, Shijiazhuang, 640 pp.

    Soto, M.E. & Ramírez, M.J. (2012) Revision and phylogenetic analysis of the spider genus Philisca Simon (Araneae: Anyphaenidae, Amaurobioidinae). Zootaxa, 3443, 1–65.

    Sudhikumar, A.V., Mathew, M.J., Sunish, E. & Sebastian, P.A. (2005) Seasonal variation in spider abundance in Kuttanad rice agroecosystem, Kerala, India (Araneae). Acta Zoologica Bulgarica, 1 (Supplement), 181–190.

    Sundevall, C.J. (1823) Specimen academicum genera araneidum Sueciae exhibens. Lundae, 1–22.

    Sundevall, C.J. (1833) Svenska spindlarnes beskrifning. Fortsättning och slut. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar, 1832, 172–272.

    Tanikawa, A. (2007) An identification guide to the Japanese spiders of the families Araneidae, Nephilidae and Tetragnathidae. Arachnological Society of Japan, Osaka, 121 pp.

    Tanikawa, A. (2009) Hersiliidae. Nephilidae, Tetragnathidae, Araneidae. In: Ono, H. (Ed.), The Spiders of Japan with keys to the families and genera and illustrations of the species. Tokai University Press, Kanagawa, pp. 149, 403–463.

    Uhl, G. (1994) Genital morphology and sperm storage in Pholcus phalangioides (Fuesslin, 1775) (Pholcidae; Araneae). Acta Zoologica, 75 (1), 1–12.

    http://dx.doi.org/10.1111/j.1463-6395.1994.tb00957.x

    Uhl, G. (2000) Two distinctly different sperm storage organs in female Dysdera erythrina (Araneae: Dysderidae). Arthropod Structure & Development, 29 (2), 163–169.

    http://dx.doi.org/10.1016/S1467-8039(00)00019-0

    Vega, A.S., Rua, G.H., Fabbri, L.T. & Rúgolo de Agrasar, Z.E. (2009) A Morphology-Based Cladistic Analysis of Digitaria (Poaceae, Panicoideae, Paniceae). Systematic Botany, 34 (2), 312–323.

    http://dx.doi.org/10.1600/036364409788606325

    Vellard, J. (1926) Contribution à l'étude des araignées. Nouvelles espèces d'Argiopidae. Memórias do Instituto Butantan, 3, 327–331.

    Walckenaer, C.A. (1841) Histoire naturelle des Insects. Aptères. Librairie Enciclopédique de Roret, Paris, 549 pp.

    Welch, K., Crain, P.R. & Harwood, J.D. (2011) Phenological dynamics of web-building spider populations in alfalfa: implications for biological control. The Journal of Arachnology, 39 (2), 244–249.

    http://dx.doi.org/10.1636/CP10-83.1

    Werenkraut, V. & Ramírez, M.J. (2009) A revision and phylogenetic analysis of the spider genus Coptoprepes Simon (Araneae: Anyphaenidae, Amaurobioidinae). Zootaxa, 2212, 1–40.

    World Spider Catalog (2015) World Spider Catalog. Natural History Museum Bern. Available from: http://wsc.nmbe.ch (accessed 15 January 2015)

    Young, O.P. & Edwards, G.B. (1990) Spiders in United States field crops and their potential effect on crop pests. The Journal of Arachnology, 18 (1), 1–27.

    Zhu, C.D. & Wen, Z.G. (1978) A new species of spider of the genus Dyschiriognatha (Araneae: Tetragnathidae) from China. Journal of Jilin University-Medicine Edition, 3, 16–18.

    Zhu, M.S., Song, D.X. & Zhang, J.X. (2003) Fauna Sinica: Invertebrata. Vol. 35. Arachnida: Araneae: Tetragnathidae. Science Press, Beijing, vii + 418 pp.