Abstract
The endemic Australasian species of Ypsiloncyphon are the sister group of the combined Asian species groups 1, 2, and 4. The description of the type species, Y. chlorizans (Klausnitzer), is supplemented by details of male and female genitalia. New species are described and illustrated in several genera: Austrocyphon scissus n. sp., Leptocyphon abnormis n. sp., Petrocyphon bonang n. sp., P. lacteus n. sp., P. televisionarius n. sp., Prionocyphon bidentatus n. sp., P. cacatua n. sp., P. laurae n. sp., P. neboissi n. sp., P. serratus n. sp., P. uncatus n. sp., and P. urbanus n. sp. Genus Prionocyphon is distinguished from Oriental genera with similar antennal modifications. However, a synapomorphy of Prionocyphon as presently understood is not known. Supplementary information on various species in the aforementioned genera and in Pachycyphon and Calvarium is provided. A key to the genera of adult Australian Scirtidae: Scirtinae is presented.
References
Bonzle Digital Atlas of Australia (2012) Bonzle Digital Atlas of Australia. Available from: http://maps.bonzle.com/c/a (accessed 3 August 2015)
Champion, G.C. (1924) Coleoptera from the Seychelles: Lampyridae, Helodidae, Cantharidae, Melyridae and supplement to Cleridae. Transactions of the Entomological Society of London, 1923, 295–304.
Cooper, S.J.B., Watts, C.H.S., Saint, K.M. & Leijs, R. (2014) Phylogenetic relationships of Australian Scirtidae (Coleoptera) based on mitochondrial and nuclear sequences. Invertebrate Systematics, 28, 628–642.
http://dx.doi.org/10.1071/IS13046Google Earth (2012) Available from: http://www.gearth.de (accessed 29 September 2014)
Klausnitzer, B. (1973) Zur Kenntnis der Cyphon-Fauna der Philippinen, Sumbawas, Neuguineas und des Bismarck-Archipels (Col., Helodidae). Folia entomologica Hungarica, 26, 97–110.
Klausnitzer, B. (1980a) Zur Kenntnis der Helodidae des Himalaja-Gebietes (Col.). Entomologica Basiliensia, 5, 195–214.
Klausnitzer, B. (1980b) Zur Kenntnis der Helodidae von Vietnam (Coleoptera). Folia entomologica Hungarica, 33, 87–94.
Klausnitzer, B. (2009a) Beschreibung einer neuen Gattung der Scirtidae aus der Orientalischen Region (Coleoptera) (144. Beitrag zur Kenntnis der Scirtidae). Beiträge zur Entomologie, 59, 247–261.
Klausnitzer, B. (2009b) Insecta: Coleoptera: Scirtidae. Süßwasserfauna von Mitteleuropa, 20/17. Spektrum Akademischer Verlag, Heidelberg, XIV + 326 pp.
Klausnitzer, B. (2010) Neue Arten von Mescirtes Motschulsky, 1863 aus Vietnam (Coleoptera, Scirtidae) (149. Beitrag zur Kenntnis der Scirtidae). Denisia, 29, 165–172.
Klausnitzer, B. (2011a) Redeskription von Prionoscirtes reliquus Champion, 1918 (Coleoptera, Scirtidae) (161. Beitrag zur Kenntnis der Scirtidae). Beiträge zur Entomologie, 61, 271–275.
Klausnitzer, B. (2011b) Beschreibung von Prionocara n. gen. aus der Orientalischen Region (Coleoptera, Scirtidae) (165. Beitrag zur Kenntnis der Scirtidae). Entomologische Blätter, 107, 65–76.
Klausnitzer, B. (2012) Zur Kenntnis der Gattung Prionocyphon L. Redtenbacher 1858 sensu lato (Coleoptera, Scirtidae) aus Südamerika (181. Beitrag zur Kenntnis der Scirtidae). Linzer biologische Beiträge, 44, 1177–1194.
Klausnitzer, B. (2015a) Neue Arten der Gattung Ypsiloncyphon Klausnitzer, 2009 (Coleoptera, Scirtidae) aus China und Nordindien sowie Anmerkungen zu Ypsiloncyphon-Arten aus Nepal. (200. Beitrag zur Kenntnis der Scirtidae). Entomologische Nachrichten und Berichte, 59 (3–4), 157–168 + cover page 4.
Klausnitzer, B. (2015b) Neue Arten der Gattung Ypsiloncyphon Klausnitzer, 2009 (Coleoptera, Scirtidae) aus der Orientalischen Region (201. Beitrag zur Kenntnis der Scirtidae). Entomologische Blätter und Coleoptera, 111, 465–479.
Lawrence, J.F. (2001) A new genus of Valdivian Scirtidae with comments on Scirtoidea and beetle suborders. Sukanahikoma. Special Publication of the Japan Coleopterological Society Osaka, 1, 351–361.
Lawrence, J. F. & Yoshitomi, H. (2007) Nipponocyphon, a new genus of Japanese Scirtidae (Coleoptera) and its phylogenetic significance. Elytra, Tokyo, 35 (2), 507–527.
Libonatti, M.L. (2014) A revision of the genus Ora Clark, 1865 (Coleoptera: Scirtidae) in Argentina (part I) – descriptions of new species. Zootaxa, 3884 (1), 27–44.
http://dx.doi.org/10.11646/zootaxa.3884.1.2Müller, Ph.W. J. (1821) Neue Insekten, beschrieben von Ph.W.J. Müller. Magazin der Entomologie. Herausgegeben von E. F. Germar (Halle), 4, 184–230.
Nyholm, T. (1969) Über Bau und Funktion der Kopulationsorgane bei den Cyphones (Col. Helodidae) Studien über die Familie Helodidae. X. Entomologisk Tidsskrift, 90, 233–271, pls. I–Vii.
Nyholm, T. (1971) Beiträge zur Kenntnis der paläarktischen Helodiden 2. Bemerkungen über die Gattung Prionocyphon Redtb. Mit Beschreibung zweier neuer Arten aus dem Mittelmeergebiet Studien über die Familie Helodidae. XII. Entomologisk Tidsskrift, 92, 28–42.
Nyholm, T. (1972) Zur Morphologie und Funktion des Helodiden-Aedoeagus (Col.). Entomologica Scandinavica, 3, 81–119.
http://dx.doi.org/10.1163/187631272X00193Nyholm, T. (2002) Scirtes japonicus Kiesenwetter and its allies, with description of Scirtes ussuriensis n. sp. (Coleoptera, Scirtidae). Entomologische Blätter, 98, 49–60.
Paykull, G. (1799) Fauna Suecica; Insecta (Col.). Vol. 2. Upsaliae, Edman, 234 pp. [Cyphon: pp. 117–121]
Pic, M. (1914) Fam. Dascillidae, Helodidae, Eucinetidae. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Pars 58. W. Junk, Berlin, pp. 1–65.
Redtenbacher, L. (1858) Fauna Austriaca. Die Käfer. Nach der analytischen Methode bearbeitet. 2. Auflage. Carl Gerold's Sohn, Wien, 136 + 1017 pp.
Ruta, R. (2004). Five new species of Oriental Scirtidae (Coleoptera: Scirtoidea). Genus, 15, 363–379.
Ruta, R. (2007) Scirtidae of India and Sri Lanka. Part 1. The chlorizans-group of Cyphon Paykull, 1799 (Insecta: Coleoptera). Genus, 17, 323–340.
Ruta, R. (2008) Contribution to the knowledge of Seychellois Scirtidae (Coleoptera: Scirtoidea). Zootaxa, 1913, 49–68.
Ruta, R. (2009) Brachycyphon Fairmaire, 1896, a neglected genus of Afrotropical Scirtidae (Coleoptera). Annales Zoologici, 59 (4), 621–628. [Warszawa, 2009]
http://dx.doi.org/10.3161/000345409x484982Ruta, R. (2010) Two new species of Prionocyphon Redtenbacher from Taiwan (Coleoptera: Scirtidae), with notes on Prionocyphon, Mescirtes Motschulsky and Prionoscirtes Champion from East and Southeast Asia. Zootaxa, 2402, 52–60.
Ruta, R., Kiałka, A. & Yoshitomi, H. (2014) A supplement to the revision of the Scirtes flavoguttatus species-group (Coleoptera: Scirtidae: Scirtinae). Zootaxa, 3902 (1), 1–62.
http://dx.doi.org/10.11646/zootaxa.3902.1.1Ruta, R., Thorpe, S. & Yoshitomi, H. (2011) Stenocyphon neozealandicus, a new species from New Zealand of a previously monotypic subfamily from Chile (Coleoptera: Scirtidae: Stenocyphoninae). Zootaxa, 3113, 65–68.
Sharp, D. (1878) On the Dascillidae of New Zealand. Annals and Magazine of Natural History, Series 5, 2, 40–59.
http://dx.doi.org/10.1080/00222937808682380Watts, C.H.S. (2004) Revision of Australian Scirtes Illiger and Ora Clark (Coleoptera: Scirtidae). Transactions of the Royal Society of South Australia, 128, 131–168.
Watts, C.H.S. (2007) Revision of Australian Pseudomicrocara Armstrong (Coleoptera:Scirtidae). Transactions of the Royal Society of South Australia, 131, 1–80.
Watts, C.H.S. (2009) Revision of Heterocyphon Armstrong, Peneveronatus Armstrong and Accolabass gen. nov. (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 133, 108–149.
Watts, C.H.S. (2010a) Revision of the genus Macrohelodes (Scirtidae: Coleoptera; Insecta). Transactions of the Royal Society of South Australia, 134, 19–52.
Watts, C.H.S. (2010b). Revision of Australian Prionocyphon Redtenbacher (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 134, 53–88.
Watts, C.H.S. (2011) Revision of Australian Scirtidae of the genera Chameloscyphon gen. nov., Daploeuros gen. nov., Dasyscyphon gen. nov., Eurycyphon gen. nov., Macrodascillus Carter, Petrocyphon gen. nov. and Spaniosdascillus gen. nov. (Coleoptera). Transactions of the Royal Society of South Australia, 135, 66–10.
Watts, C.H.S. (2014) The larvae of some Australian Scirtidae (Coleoptera) with a key to known genera. Transactions of the Royal Society of South Australia, 138, 1–91.
http://dx.doi.org/10.1080/03721426.2014.10887191Yoshitomi, H. (2005) Systematic revision of the family Scirtidae of Japan, with phylogeny, morphology and bionomics (Insecta: Coleoptera, Scirtoidea). Japanese Journal of Systematic Entomology Monographic Series, 3, 1–212.
Yoshitomi, H. (2015) New species of the genus Ypsiloncyphon (Insecta: Coleoptera: Scirtidae: Scirtinae). Zootaxa, 3904 (4), 451–500.
http://dx.doi.org/10.11646/zootaxa.3904.4.1Zwick, P. (2011) Some Scirtidae (Coleoptera) from Palawan (the Philippines), mainly from phytotelmata. Aquatic Insects, 33, 233–252.
http://dx.doi.org/10.1080/01650424.2011.614257Zwick, P. (2012) Australian Marsh Beetles (Coleoptera: Scirtidae). 1. Additions to genus Petrocyphon. Entomologische Blätter und Coleoptera, 108, 159–180.
Zwick, P. (2013a) Australian Marsh Beetles (Coleoptera: Scirtidae). 2. Pachycyphon, a new genus of presumably terrestrial Australian Scirtidae. Zootaxa, 3626 (3), 326–344.
http://dx.doi.org/10.11646/zootaxa.3626.3.2Zwick, P. (2013b) Australian Marsh Beetles (Coleoptera: Scirtidae). 3. A restricted concept of genus Cyphon, Australian species of Cyphon s. str., and the new Australasian genus Nanocyphon. Genus, 24, 163–189.
Zwick, P. (2013c) Australian Marsh Beetles (Coleoptera: Scirtidae). 4. Two new genera, Austrocyphon and Tasmanocyphon. Zootaxa, 3706 (1), 1–74.
http://dx.doi.org/10.11646/zootaxa.3706.1.1Zwick, P. (2013d) ERRATUM: Peter Zwick (2013) Australian Marsh Beetles (Coleoptera: Scirtidae) 4. Two new genera, Austrocyphon and Tasmanocyphon. Zootaxa, 3706 (1), 001–074. Zootaxa, 3710 (6), 600.
http://dx.doi.org/10.11646/zootaxa.3710.6.6Zwick, P. (2014) Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer. Zootaxa, 3846 (1), 1–41.
http://dx.doi.org/10.11646/zootaxa.3846.1.1Zwick, P. (2015a) Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa, 3981 (3), 301–359.
http://dx.doi.org/10.11646/zootaxa.3981.3.1Zwick, P. (2015b) Australian Marsh Beetles (Coleoptera: Scirtidae). 8. The new genera Cygnocyphon, Eximiocyphon, Paracyphon, Leptocyphon, Tectocyphon, and additions to Contacyphon de Gozis and Eurycyphon Watts. Zootaxa, 3981 (4), 451–490.
http://dx.doi.org/10.11646/zootaxa.3981.4.1Zwick, P. (2015c) To the knowledge of Sarabandus robustus (LeConte) (Col.: Scirtidae: Scirtinae), and on the groundplan of male marsh beetle genitalia. Linzer biologische Beiträge, 47 (2), 1439–1449.
Zwick, P. (2015d) Three new Marsh Beetles (Col.: Scirtidae) from New Guinea and Java. Linzer biologische Beiträge, 47 (2), 1885–1895.
Zwick, P., Klausnitzer, B. & Ruta, R. (2013) Contacyphon Gozis, 1886 removed from synonymy (Coleoptera: Scirtidae) to accommodate species so far combined with the invalid name, Cyphon Paykull, 1799. Entomologische Blätter und Coleoptera, 109, 337–353.