Abstract
A review of the West-Palaearctic Hebridae and their distribution is given, including two genera and 21 species-group taxa (18 species and 3 subspecies). A new species, Hebrus (Hebrus) atlas sp. nov. (Hemiptera: Heteroptera: Hebroidea: Hebridae: Hebrinae), is described from Morocco and is the first micropterous species of Hebrus s. str. in the Palaearctic Region. The species Hebrus fulvinervis Horváth, 1929, not recorded since its original description, is rediscovered, redescribed, and its species status is confirmed. Based on the morphological characters, H. fulvinervis is considered closely related to H. ruficeps Thomson, 1871, and transferred to the subgenus Hebrusella Poisson, 1944. Remarks on the morphology of Hebrus syriacus Horváth, 1896 are provided and its placement in the subgenus Hebrus s. str. is confirmed. The following new or confirmed distribution records are provided: Hebrus acutiscutatus Brown, 1951 from Israel, H. fulvi-nervis from Bulgaria, H. jeanneli djaneti Poisson, 1953 from Morocco, H. montanus Kolenati, 1857 from Israel and Romania, H. pusillus arabicus Linnavuori, 1971 from Iran (Hormozgan) and Oman, H. pusillus pusillus (Fallén, 1807) from Greece and Lithuania, H. syriacus from Cyprus, and H. vaillanti Poisson, 1953 from Morocco. Several corrections concerning type localities, type material and depositories are also provided.
References
Andersen, N.M. (1982) The semiaquatic bugs (Hemiptera, Gerromorpha). Phylogeny, adaptations, biogeography and classification. Entomograph, Vol. 3. Scandiavian Science Press Ltd., Klampenborg, 455 pp.
Andersen, N.M. (1995) Infraorder Gerromorpha Popov, 1971—semiaquatic bugs, pp. 17–114. In: Aukema, B. & Rieger, C. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 1. Enicocephalomorpha, Dipsocoromorpha, Nepomorpha, Gerromorpha and Leptopodomorpha. The Netherlands Entomological Society, Amsterdam, xxvi + 222 pp.
Andersen, N.M. & Gaun, S. (1974) Fortegnelse over Danmarks tæger (Hemiptera-Heteroptera). (Check-list of land and water bugs of Denmark). Entomologiske Meddelelser, 42, 113–134. [in Danish, English summary].
Andersen, N.M. & Weir, T.A. (2004) Australian water bugs. Their biology and identification (Hemiptera-Heteroptera, Gerromorpha & Nepomorpha). Entomograph, Vol. 14. Apollo Books, Stenstrup & CSIRO Publishing, Collingwood, 344 pp.
Anonymus (2016) Zoologische Staatssammlung München. Sammlungsbestände der Sektion Hemiptera. Heteroptera. Available from: http://www.zsm.mwn.de/rhy/heteroptera_families.htm (Accessed 8 June 2016)
Assmann, A. (1858) Synonymische Bemerkungen. Ueber Hemiptera. Berliner Entomologische Zeitschrift, 2, 81.
Aukema, B., Cuppen, J.G.M., Nieser, N. & Tempelman, D. (2002) Verspreidingsatlas Nederlandse wantsen (Hemiptera: Heteroptera). Deel I: Dipsocoromorpha, Nepomorpha, Gerromorpha & Leptopodomorpha. [Distribution atlas of true bugs of the Netherlands (Hemiptera: Heteroptera). Volume I: Dipsocoromorpha, Nepomorpha, Gerromorpha & Leptopodomorpha]. European Invertebrate Survey—Nederland, Leiden, 169 pp.
Aukema, B., Duffels, H., Günther, H., Rieger, Ch. & Strauß, G. (2013b) New data on the Heteroptera fauna of La Palma, Canary Islands (Insecta: Hemiptera). In: Kment, P., Malenovský, I. & Kolibáč, J. (Eds.), Studies in Hemiptera in Honour of Pavel Lauterer and Jaroslav L. Stehlík. Acta Musei Moraviae, Scientiae Biologicae 98 (2), pp. 459–493.
Aukema, B., Rieger, Ch. & Rabitsch, W. (2013a) Catalogue of the Heteroptera of the Palaearctic Region. VI. Supplement. The Netherlands Entomological Society, Amsterdam, xxiii + 629 pp.
Baena, M. (1996) Notas sobre Hebrus Curtis, 1833 Españoles (Heteroptera: Hebridae). Zapateri, 6, 123–125.
Baena, M. & Báez, M. (1990) Los Heterópteros acuáticos de las Islas Canarias (Heteróptera; Nepomorpha, Gerromorpha). Vieraea, 19, 233–244.
Baena, M. & Vázquez, M.A. (1986) Catálogo preliminar de los Heterópteros acuáticos ibéricos (Heteroptera: Nepomorpha, Gerromorpha). Graellsia, 42, 61–89.
Báez, M., Heiss, E., García, A. & Cabrera, A. (2005) Hemiptera: Heteroptera. In: Arechavaleta, M., Zurita, N., Marrero, M.C. & Martín, J.L. (Eds.), Lista Preliminar de Especies Silvestres de Cabo Verde (Hongos, Plantas y Animales Terrestres). Consejería de Medio Ambiente y Ordenación Territorial, Gobierno de Canarias, pp. 71–78.
Benedek, P. (1969) Poloskák VII. (Heteroptera. VII). Magyarország Állatvilága. (Fauna Hungariae). Akadémiai Kiadó, Budapest, 86 pp. [in Hungarian]
Benedek, P. (1970) The semiaquatic Heteroptera in the Carpathian Basin with notes on the distribution and phenology of the species. Faunistische Abhandlungen Staatliches Museum für Tierkunde Dresden, 3, 27–49.
Berchi, G.M. (2013) Checklist and distribution of the family Notonectidae in Romania, with the first record of Notonecta maculata Fabricius, 1794 (Hemiptera: Heteroptera: Nepomorpha). Zootaxa, 3682 (1), 121–123.
http://dx.doi.org/10.11646/zootaxa.3682.1.5Berchi, G.M. & Kment, P. (2015) Review of the family Veliidae in Romania (Hemiptera: Heteroptera: Gerromorpha). Zootaxa, 3963 (1), 74–88.
http://dx.doi.org/10.11646/zootaxa.3963.1.5Berchi, G.M., Kment, P., Copilaș-Ciocianu, D., Rákosy, L. & Damgaard, J. (2016) Water treaders of Romania and adjacent countries and their phylogenetic relationships (Hemiptera: Heteroptera: Mesoveliidae). Annales Zoologici (Warszawa), 66 (2), 193–212.
http://dx.doi.org/10.3161/00034541anz2016.66.2.004Bertrand, H. (1958) Ricerche Zoologiche sul Massiccio del Pollino (Lucania-Calabria). XXVI. Hemiptères [sic] aquatiques recueillis en Calabre. Annuario dell’Istituto e Museo di Zoologia della Università di Napoli, 10 (3), 1–4.
Blatchley, W.S. (1926) Heteroptera or true bugs of eastern North America, with special reference to the faunas of Indiana and Florida. Nature Publishing Company, Indianapolis, 1116 pp.
http://dx.doi.org/10.5962/bhl.title.6871Boda, P., Bozóki, T., Vásárhelyi, T., Bakonyi, G. & Várbíró, G. (2015) Revised and annotated checklist of aquatic and semiaquatic Heteroptera of Hungary with comments on biodiversity patterns. ZooKeys, 501, 89–108.
http://dx.doi.org/10.3897/zookeys.501.8964Brown, E.S. (1951) Aquatic and semi-aquatic Hemiptera. In: Anonymus (Ed.), Expedition to S.W. Arabia, 1937–1938. Vol. 1. British Museum (Natural History), London, pp. 221–273.
Burmeister, H.C.C. (1835) Handbuch der Entomologie. Band 2, Abt. 1. Ordnung Rhynchota, Abt. 1. Hemiptera. Enslin, Berlin, xii + 400 pp. [Abt. I was published in 1835]
Chen, P.-P., Nieser, N. & Lekprayoon, C. (2006) Notes on the Asian water bugs, with descriptions of two new species of Timasius Distant (Hemiptera: Gerromorpha). Tijdschrift voor Entomologie, 149, 55–66.
http://dx.doi.org/10.1163/22119434-900000187Chen, P.-P., Nieser, N. & Zettel, H. (2005) The aquatic and semi-aquatic bugs (Heteroptera: Nepomorpha & Gerromorpha) of Malesia. Fauna Malesiana Handbook, Vol. 5. Brill, Leiden—Boston, i–x + 1–546 pp.
Cianferoni, F. (2015) The forgotten Hebrid (Gerromorpha, Hebridae). In: Anonymus (Ed.), 7th European Hemiptera Congress and 9th International Workshop on Leafhoppers and Planthoppers of Economic Importance. Programme. Abstracts of Talks and Posters. Seggau Castle / Graz (Austria), July 19th–24th, 2015. Ökoteam – Institute for Animal Ecology and Landscape Planning, Graz, pp. 26.
Cianferoni, F., Buzzetti, F.M. & Zettel, H. (2016) The “Italian hebrid” Hebrus franzi (Wagner, 1957): disentangling a half-century dilemma (Hemiptera: Heteroptera: Gerromorpha). Zootaxa, 4132 (1), 127–134.
http://dx.doi.org/10.11646/zootaxa.4132.1.11Cianferoni, F. & Mazza, G. (2012) The aquatic Heteroptera (Insecta: Hemiptera) of the “Foreste Casentinesi, Monte Falterona e Campigna” National Park (Central Italy). Zootaxa, 3568, 36–52.
Cobben, R.H. (1982a) A new hebrid species from a cave in Zaire (Hebridae, Heteroptera). Revue Zoologique Africaine, 96 (4), 797–802.
Cobben, R.H. (1982b) The Hebrid fauna of the Ethiopian Kaffa province, with considerations on species grouping (Hebridae, Heteroptera). Tijdschrift voor Entomologie, 125, 1–24.
Costa, A. (1864) Peregrinazione zoologica. Annuario del Museo Zoologico della R. Università di Napoli, 2, 119–125.
Coulianos, C.-C. (1998) Annotated catalogue of the Hemiptera-Heteroptera of Norway. Fauna Norvegica, Series B, 45, 11–40.
Coulianos, C.-C. (2005) Annotated checklist and distribution of the true bugs (Hemiptera-Heteroptera) of Estonia. Proceedings of the Estonian Academy of Sciences, Biology, Ecology, 54, 136–165.
Coulianos, C.-C., Økland, J. & Økland, K.A. (2008) Norwegian water bugs. Distribution and ecology (Hemiptera-Heteroptera: Gerromorpha and Nepomorpha). Norwegian Journal of Entomology, 55, 179–222.
Coulianos, C.-C. & Ossiannilsson, F. (1976) Catalogus Insectorum Sueciae. VII. Hemiptera-Heteroptera. 2nd Ed. Entomologisk Tidskrift, 97, 135–173.
Curtis, J. (1833) Characters of some undescribed genera and species, indicated in the “Guide to an arrangement of British Insects”. Entomological Magazine, 1, 186–199.
Derzhansky, V.V. (1984) Fauna vodnykh klopov (Heteroptera) Moldavii. [Fauna of water bugs (Heteroptera) of Moldavia)]. Izvestiya Akademii Nauk Moldavskoy SSR, Seriya Biologicheskikh i Khimicheskikh Nauk, 1984 (2), 74–75. [in Russian]
Derzhansky, V.V. (1997) List of the Heteroptera of the Republic of Moldova. Zoological Institut, Russian Academy of Sciences, St. Petersburg, 22 pp.
Dethier, M. & Matthey, W. (1977) Contribution à la connaissance des Hétéroptères aquatiques de Suisse. Revue Suisse de Zoologie, 84, 583–591.
Drake, C.J. (1917) A survey of the North American species of Merragata. Ohio Journal of Science, 17, 101–105.
Drake, C.J. (1952) A new tropical hebrid (Hemiptera). Pan-Pacific Entomologist, 28, 194.
Drake, C.J. & Chapman, H.C. (1958) New Neotropical Hebridae, including a catalogue of the American species (Hemiptera). Journal of the Washington Academy of Sciences, 48, 317–326.
Drake, C.J. & Cobben, R.H. (1960) The Heteroptera of the Netherlands Antilles – II. Hebridae. Studies on the Fauna of Curaçao and Other Caribbean Islands, 51 (11), 35–43.
Esenbekova, P.A., Nurushev, M.Zh. & Homziak, J. (2015) Aquatic Hemiptera (Heteroptera) of Kazakhstan, with notes on life history, ecology and distribution. Zootaxa, 4013 (2), 195–206.
http://dx.doi.org/10.11646/zootaxa.4013.2.2Fallén, C.F. (1807) Monographia Cimicum Sveciæ. C.G. Proft, Hafniæ, 123 pp.
Fent, M., Kment, P., Çamur-Elipek, B. & Kırgız, T. (2011) Annotated catalogue of Enicocephalomorpha, Dipsocoromorpha, Nepomorpha, Gerromorpha and Leptopodomorpha (Hemiptera: Heteroptera) of Turkey with new records. Zootaxa, 2856, 1–84.
Fieber, F.X. (1860) Die europäischen Hemiptera. Halbflügler (Rhynchota Heteroptera). Gerold’s Sohn, Wien, i–vi + 1–112 pp.
Gadalla, S.M. & Saleh Ahmed, R. (2005) Contribution to the study of Heteroptera, Gerromorpha (semiaquatic bugs). VI – Hebridae and Hydrometridae of Egypt. Efflatounia, 5, 16–25.
Ghahari, H., Moulet, P., Ostovan, H. & Linnavuori, R.E. (2013) An annotated catalog of the Iranian Dipsocoromorpha, Enicocephalomorpha, Gerromorpha, Leptopodomorpha and Nepomorpha (Hemiptera: Heteroptera). Zootaxa, 3641 (4), 301–342.
http://dx.doi.org/10.11646/zootaxa.3641.4.1Gheit, A. (1995) Catalogue des Hémiptères Hydrocorises et Amphibiocorises des Provinces nord Marocaines (1re note). Entomologiste (Paris), 51, 241–249.
Gogala, A. (2003) Heteroptera of Slovenia, I.: Dipsocoromorpha, Nepomorpha, Gerromorpha and Leptopodomorpha. Annals for Istrian and Mediterranean Studies, Seria Historia Naturalis, 13 (2), 229–240.
Golub, V.B., Vinokurov, N.N. & Soboleva, V.A. (2014) Novye ukazaniya poluzhestkokrylykh iz Karachaebo-Cherkesskoy Respubliki i sopredel’nykh territoriy Severnogo Kavkaza. Dipsocoridae, Hebridae, Saldidae, Tingidae, Lygaeidae (Heteroptera). (New records of bugs for Karachay-Cherkess Republic (Russia) and adjacent territories of the North Caucasus. Dipsocoridae, Hebridae, Saldidae, Tingidae, Lygaeidae (Heteroptera)). Euroasian Entomological Journal, 13, 572–576. [in Russian, English abstract].
Grandova, M.A. (2013) Aquatic Heteroptera (Nepomorpha, Gerromorpha) in small intermittent rivers of Ukraine steppe zone. ZooKeys, 319, 107–118.
Grandova, M.A. (2014) Aquatic Heteroptera of great rivers of the Ukrainian steppe zone and seasonal changes of abundance and biomass. Vestnik Zoologii, 48 (5), 441–450.
http://dx.doi.org/10.3897/zookeys.319.4328Hoberlandt, L. (1977) Heteroptera, pp. 61–82. In: Dlabola, J. (Ed.), Enumeratio Insektorum Bohemoslovakiae. Acta Faunistica Entomologica Musei Nationalis Pragae, Supplementum, 4, 1–158.
Hoffmann, H.-J. & Melber, A. (2003) Verzeichnis der Wanzen (Heteroptera) Deutschlands, pp. 209–272. In: Klausnitzer, B. (Ed.), Entomofauna Germanica. Vol. 6. Entomologische Nachrichten und Berichte, Beiheft, 8, 1–344.
Horváth, G. (1896) Hemiptera nova palæarctica. Természetrajzi Füzetek, 19, 322–329.
Horváth, G. (1897) Ordo Hemiptera. In: Paszlavszky, J. (Ed.), A Magyar birodalom állatvilága. A Magyar Birodalomból Eddig Ismert Állatok Rendszeres Lajstroma. Fauna Regni Hungariae. Animalium Hungariae Hucusque Cognitorum Enumeratio Systematica. A. K. M. Természettudományi Társulat, Budapest, pp. 5–64. [in Hungarian and Latin].
Horváth, G. (1929) Species novae Hebridarum (Hem. Het.) in Museo Nationali Hungarico asservata. Annales Historico-Naturales Musei Nationalis Hungarici, 26, 313–317.
ICZN (International Comission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. 4th Edition. International Trust for Zoological Nomenclature, London, 306 pp.
Jacquemin, G. (2006) Hétéropteres aquatiques des sites salés de Lorraine et d’Alsace (Heteroptera, Gerromorpha, Nepomorpha). Bulletin de la Société Entomologique de France, 111, 185–190.
Jaczewski, T. & Wróblewski, A. (1976) Hebridae, Mesoveliidae, Hydrometridae, Veliidae i nartniki – Gerridae. Klucze do oznaczania owadów polski. Tom. 18, Vol. 4. [Hebridae, Mesoveliidae, Hydrometridae, Veliidae and pond skaters – Gerridae. Keys for identification of Polish insects. Section 18, Vol. 4.] Państwowe Wydawnictwo Naukowe, Warszawa, Wrocław, 31 pp. [in Polish].
Jordan, K.H.C. (1954) Über einen neuen Naeogeus aus Südfrankreich (Heteroptera: Hebridae). Beiträge zur Entomologie, 4, 601−604.
Josifov, M. (1958) Khemipterologichno s’obshschenie III. (Hemipterologische Mitteilung III). Izvestiya na Zoologicheskiya Institut (Sofiya), 7, 343–349. [in Bulgarian, Russian and German summaries]
Josifov, M. (1959) Hemiptera-Heteroptera ot ostrov Tasos. (Hemiptera-Heteroptera von der Insel Thasos). Izvestiya na Zoologicheskiya Institut (Sofiya), 8, 265–269. [in Bulgarian, Russian and German summaries]
Josifov, M. (1960) Vidov s’stav i razprostranenie na nasekomite ot razreda Heteroptera v B’lgariya. (Artenzusammensetzung und Verbreitung der Insekten von der Ordnung Heteroptera in Bulgarien). Izvestiya na Zoologicheskiya Institut (Sofiya), 9, 107–177. [in Bulgarian, Russian and German summaries]
Josifov, M. (1963) Polukrili nasekomi (Heteroptera) ot okolnostite na Petrich (yugozapadna B’lgariya). (Heteropteren aus der Umgebung von Petrič (SW Bulgarien)). Izvestiya na Zoologicheskiya Institut (Sofiya), 13, 93–132. [in Bulgarian, Russian and German summaries]
Josifov, M. (1964) Heteroptera ot Trakiyskata nizina. (Heteroptera aus der Thrakischen Tiefebene). In: Anonymus (Ed.), Fauna na Trakiya, Chast I. (Die Fauna Thrakiens, Band I.), B’lgarska Akademiya na Naukite, Zoologicheski Institut s Muzey, Sofiya, pp. 207–246. [in Bulgarian, Russian and German summaries]
Josifov, M. (1970) Ergebnisse der Albanien-Expedition 1961 des Deutschen Entomologischen Institutes. 82. Beitrag. Heteroptera. Beiträge zur Entomologie, 20, 825–956.
Kanyukova, E.V. (1997) Hebridae of Russia and adjacent countries (Heteroptera). Zoosystematica Rossica, 6, 223–236.
Kanyukova, E.V. (2006) Vodnye poluzhestkokrylye nasekomye (Heteroptera: Nepomorpha, Gerromorpha) fauny Rossii i sopredelnykh stran. [Aquatic and semiaquatic (Heteroptera: Nepomorpha, Gerromorpha) of the fauna of Russia and neighbouring countries]. Dal’nauka, Vladivostok, 296 pp. [in Russian, English abstract]
Kment, P. (2006) A contribution to the faunistics of aquatic and semiaquatic bugs (Heteroptera: Nepomorpha, Gerromorpha) in Portugal, with the review of biology of the Nearctic corixid Trichocorixa verticalis (Fieber, 1851). Boletín de la Sociedad Entomológica Aragonesa, 138, 359–361.
Kment, P. & Beran, L. (2011) Check-list of water bugs (Hemiptera: Heteroptera: Nepomorpha, Gerromorpha) in Croatia with two new records and three rediscoveries. Natura Croatica, 20, 159–178.
Kment, P., Bryja, J., Jindra, Z., Hradil, K. & Baňař, P. (2003) New and interesting records of true bugs (Heteroptera) from the Czech Republic and Slovakia II. Klapalekiana, 39, 257–306.
Kment, P. & Jindra, Z. (2006) New and interesting records of true bugs (Heteroptera) from Turkey, southeastern Europe, Near and Middle East. Acta Entomologica Musei Nationalis Pragae, 45 [2005], 3–16.
Kment, P. & Kanyukova, E.V. (2010) New faunistic records of Hebridae (Hemiptera: Heteroptera) from the Mediterranean, Near and Middle East. Acta Musei Moraviae, Scientiae Biologicae, 95 (2), 11–18.
Kment, P. & Kolínová, Z. (2013) Catalogue of type specimens of true bugs (Hemiptera: Heteroptera) deposited in the National Museum, Prague, Czech Republic. Enicocephalomorpha, Dipsocoromorpha, Nepomorpha, Gerromorpha, and Leptopodomorpha. Acta Entomologica Musei Nationalis Pragae, 53, 821–890.
Kolenati, F.A. (1857) Meletemata Entomologica. Fasc. VI. Hemipterorum Heteropterorum Caucasi. Harpagocorisiae, monographice dispositae. Bulletin de la Société Impériale des Naturalistes de Moscou, 29 [1856], 419–491 + pl. III.
Laporte, F.L. de (1833) Essai d’une classification systematique de l’ordre des Hémipteres. Magasin de Zoologie, 2 (Suppl.), 1–88 [1832, 1–16; 1833, 17–88; Direction 631/1957]
Lindberg, H. (1922) Verzeichnis der von John Sahlberg und Uunio Saalas in den Mittelmeergebieten gesammelten semiaquatilen und aquatilen Heteropteren. Notulae Entomologicae, 2, 15–19, 46–49.
Lindberg, H. (1959) Hemiptera Insularum Caboverdensium. Commentationes Biologicae, 19 (1) [1958], 1–246.
Linnavuori, R. (1954) A Palaearctic Heteropterous material collected by J. Sahlberg and U. Saalas. Annales Entomologici Fennici, 19 [1953], 147–167.
Linnavuori, R. (1960) Hemiptera of Israel. I. Annales Zoologici Societatis Zoologicæ Botanicæ Fennicæ ‘Vanamo’, 22 (1), 1–71.
Linnavuori, R. (1964) Hemiptera of Egypt, with remarks on some species of the adjacent Eremian region. Annales Zoologici Fennici, 1, 306–356.
Linnavuori, R. (1971) Hemiptera of the Sudan, with remarks on some species of the adjacent countries. 1. The aquatic and subaquatic families. Annales Zoologici Fennici, 8, 340–366.
Linnavuori, R. (1980) Hemiptera of the Sudan, with remarks on the species of adjacent countries. 8. Additions and corrections. Bibliography. Acta Entomologica Fennica, 36, 1–53.
Linnavuori, R. (1981) Hemiptera of Nigeria, with remarks on some species of the adjacent countries. 1. The aquatic and subaquatic families, Saldidae and Leptopodidae. Acta Entomologica Fennica, 37, 1–39.
Linnavuori, R.E. (1986) Heteroptera of Saudi Arabia. In: Büttiker, W. & Krupp, F. (Eds), Fauna of Saudi Arabia. Vol. 8. Pro Entomologia c/o Natural History Museum, Basel, pp. 31–197.
Linnavuori, R.E. (1994) Hemiptera of Iraq. IV. Heteroptera, the aquatic and subaquatic families, Saldidae and Leptopodidae. Entomologica Fennica, 5 (2), 87–95.
Linnavuori, R.E. & Hosseini, R. (2000) Heteroptera of Guilan with remarks on species of the adjacent areas. Part 1. Guilan University Press, Rasht, [1999], 94 pp.
Linnavuori, R.E. & Harten, A. van (1997) Notes on Heteroptera (Insecta: Hemiptera) of Yemen. In: Krupp, F. & Mahnert, V. (Eds.), Fauna of Saudi Arabia. Vol. 16. National Comission for Wildlife Conservation and Development (NCWCD), Riyadh & Pro Entomologia c/o Natural History Museum, Basel, pp. 169–236.
Linnavuori, R.E. & Harten, A. van (2002) Additional notes on the Heteroptera (Insecta: Hemiptera) of Yemen II, pp. 157–188. In: Hacker, H.H. & Peks, H. (Eds.), Esperiana, Buchreihe zur Entomologie. Vol. 9. Druck und Verlag, Schwanfeld, 606 pp.
Linnavuori, R.E., Kment, P. & Carapezza, A. (2011) Order Hemiptera, suborder Heteroptera. Infraorders Nepomorpha, Gerromorpha, and Leptopodomorpha. In: Harten, A. van (Ed.), Arthropod fauna of the United Arab Emirates. Vol. 4. Multiply Marketing Consultancy Services, Abu Dhabi, pp. 72–107.
Lukashuk, A.O. (1997) Annotated list of the Heteroptera of Belarus and Baltia. Zoological Institute, Russian Academy of Sciences, St. Petersburg, 45 pp.
Mancini, C. (1936) Emitteri raccolti dal Prof. G. Scortecci nel Fezzan. Atti della Socetà Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 75, 300–306.
Matthey, W. & Fiora, M. (1979) Contribution a l’étude entomologique de la tourbiere des pontins: Observations sur les Hétéropteres aquatiques de surface, et plus particulierement sur le genre Gerris. Bulletin de la Société Neuchateloise des Sciences Naturelles, 102, 23–30.
Nieser, N. (1995) Gerromorpha from Israel (Heteroptera semiaquatica). In: Nitzu, E. (Ed.), Soil Fauna of Israel 1. Editura Academiei Române, Bucureşti, pp. 153–155.
Nieser, N. & Montes, C. (1984) Lista faunistica y bibliografica de los Heteropteros acuaticos (Nepomorpha & Gerromorpha) de España y Portugal. Asociacion Española de Limnologia, Madrid, 69 pp.
Nieser, N. & Moubayed, Z. (1985) Les Hétéroptères aquatiques du Liban. I. Inventaire faunistique. Annales Limnologici, 21 (3), 247–252.
http://dx.doi.org/10.1051/limn/1985025Pagola-Carte, S. & Ribes, J. (2007) Registros interesantes de hetrópteros (Hemiptera: Heteroptera) en el País Vasco (norte de la Península Ibérica). Heteropterus Revista de Entomología, 7, 25–31.
Poisson, R. (1934) Contribution a l’étude de la faune du Mozambique. Voyage de M.P. Lesne (1928–1929) (15e note). Hémipteres aquatiques. Bulletin de la Société Zoologique de France, 59, 87–103.
Poisson, R. (1944) Contribution a la connaissance des especes africaines du genre Hebrus Curtis 1833 [Hemiptera Gymnocerata] (Missions de Ch. Alluaud et R. Jeannel en Afrique orientale). Revue Francaise d’Entomologie, Nouvelle Série, 10 (3–4) [1943], 89–112.
Poisson, R.A. (1952) Contribution à l’étude des Hydrocorises de Madagascar (Missions J. Milliot et R. Paulian 1949). 2e Note. Mémoires de lʼInstitut Scientifique de Madagascar, Série E, 1 (1), 23–70.
Poisson, R. (1953) Les Hydrocorises du Tassili des Ajjer (Sahara central). In: Bernard, F. (Ed.), Mission scientifique au Tassili des Ajjers (1949). I. Recherches Zoologiques et Medicales. Institut des Recherches sahariennes de l’Université d’Alger, Alger, pp. 65–85.
Poisson, R. (1954a) Deux Hebrides (Hem. het.) nouveaux des Canaries. Commentationes Biologicae, 14 (4), 1–3.
Poisson, R. (1954b) Sur la faune des Hémiptères-Hétéroptères aquatiques des îles Canaries. Ses affinités biogéographiques. Annales du Musée du Congo Belge, Tervuren, in-8°, Zoologie, 1, 506–510.
Poisson, R.A. (1956) Contribution a l’etude des Hydrocorises de Madagascar (Heteroptera). 4° Mémoire. Mémoires de l’Institut Scientifique de Madagascar, Série E, 7, 243–265.
Poisson, R. (1957a) Hétéroptères aquatiques. Faune de France, Vol. 61. Fédération Française des Sociétés de Sciences Naturelles, Paris, 263 pp.
Poisson, R. (1957b) Hemiptera Heteroptera: Hydrocorisae & Geocorisae-Gerroidea. In: Hanström, B., Brinck, P. & Rudebeck, G. (Eds), South African Animal Life. Vol. IV. Almqvist & Wiksell, Stockholm, pp. 327–373.
Poisson, R.A. (1964) Deux espèces nouvelles d’Hémiptères aquatiques de l’Afrique éthiopienne. Annales Entomologici Fennici, 30, 214–218.
Poisson, R.A. (1968) Contributions à la connaissance de la faune entomologique de la Côte d’Ivoire (J. Decelle 1961–1964). Annales du Musée du Congo Belge, Tervuren, in-8°, Zoologie, 165, 399–403.
Polhemus, J.T. & Polhemus, D.A. (1988) Family Hebridae Amyot and Serville, 1843. The velvet water bugs. In: Henry, J.T. & Froeschner, R.C. (Eds.), Catalog of the Heteroptera, or True Bugs, of Canada and the Continental United States. E.J. Brill, Leiden, New York, Københaven, Köln, pp. 152–155.
Prokin, A.A., Shapovalov, M.I. & Saprykin, M.A. (2009) New records of aquatic and semi-aquatic bugs (Heteroptera: Gerromorpha, Nepomorpha) from the North-West. Euroasian Entomological Journal, 8, 313–314.
Protić, Lj.D. (1998) Catalogue of the Heteroptera fauna of Yougoslav countries. Part one. Prirodnjački Muzej u Beogradu, Prosebna Izdanja, 38, 1–215.
Puton, A., 1884. Notes hémiptérologiques. Revue d’Entomologie, 3, 142–149.
Putshkov, V.G. & Putshkov, P.V. (1996) Heteroptera of the Ukraine: Check list and distribution. Zoological Institute, Russian Academy of Sciences, 109 pp.
Rabitsch, W. (2005) Heteroptera (Insecta). In: Schuster, R. (Ed.), Checklisten der Fauna Österreichs, No. 2. Österreichische Akademie der Wissenschaften, Wien, pp. 1–64.
Reduciendo Klementová, B., Kment, P. & Svitok, M. (2015) Checklist of water bugs (Hemiptera: Heteroptera: Nepomorpha, Gerromorpha) of Slovakia. Zootaxa, 4058 (2), 227–243.
http://dx.doi.org/10.11646/zootaxa.4058.2.5Reichling, L. & Gerend, R. (1994) Liste des Hétéroptères du Grand-Duché de Luxembourg. Bulletin de la Société des Naturalistes Luxembourgeois, 95, 273–286.
Rey, C. (1893) Remarques en passant. Échange, Revue Linnéenne, 9, 109.
Ribes, J. & Sauleda, N. (1979) Heteropteros de alicante y zonas adyacentes. Mediterránea, 3, 123–158.
http://dx.doi.org/10.14198/MDTRRA1979.3.05Rintala, T. & Rinne, V. (2010) Suomen Luteet. [Heteroptera of Finland]. Hyönteistarvike Tibiale Oy, Helsinki, 352 pp. [in Finnish]
Roth, S. & Coulianos, C.-C. (2014) A survey of aquatic and terrestrial Heteroptera in northern Europe with special regard to Finnmark, Norway (and adjacent regions). Norwegian Journal of Entomology, 61, 99–116.
Royer, M. (1924) Travaux scientifiques de l’Armée d’Orient (1916–1918). Hémiptéres Hétéropteres. (Troisiéme note). Bulletin du Muséum National d’Histoire Naturelle (Paris), 30, 193–200.
Saleh Ahmed, R. & Gadalla, S.M. (2005) Contribution to the study of Heteroptera, Gerromorpha (Semiaquatic bugs). VII. Check-list of Gerromorpha of Egypt, with a new record. Journal of Asia-Pacific Entomology, 8 (4), 339–344.
http://dx.doi.org/10.1016/S1226-8615(08)60255-1Sakenin, H., Samin, N., Rastegar, J., Havaskary, M., Imani, S. & Linnavuori, R.E. (2010) A faunistic survey on Heteroptera (Insecta) of Western Iran. Linzer Biologische Beiträge, 42, 861–872.
Samin, N., Sakenin, H., Linnavuori, R., Havaskary, M. & Mohebbi, H.R. (2011) A faunistic survey of Heteroptera (Insecta) from western Iran. Calodema, 146, 1–12.
Savage, A.A. (1989) Adults of the British aquatic Hemiptera Heteroptera. A key with ecological notes. Freshwater Biological Association, Ambleside, 173 pp.
Schilling, P.S. (1839) Der Zwerg-Wassertreter (Velia nana nov. spec.). Übersicht der Arbeiten und Veränderungen der Schlesischen Gesellschaft für Vaterländische Kultur, 1838, 56–57.
Scholtz, H. (1847) Prodromus zu einer Rhynchoten-Fauna von Schlesien. 1. Abhandlungen der Schlesischen Gesellschaft für Vaterländische Kultur, Naturwissenschaftlich-Medizinische Abteilung, 1846, 104–164.
Seabra, A.F. de (1926) Revisão dos Hemípteros Heterópteros da fauna paleárctica existentes no Museu Zoológico da Universidade de Coimbra. Memórias e Estudos do Museu Zoológico da Universidade de Coimbra, Série 1, 10, 1–234.
Seabra, A.F. de (1941) Contribuïções para o inventário da fauna lusitânica. Insecta. Heteroptera. Memórias e Estudos do Museu Zoológico da Universidade de Coimbra, Série 1, 123, 1–36.
Servadei, A. (1958) Considerazioni sugli Emitteri italiani (Heteroptera and Homoptera Auchenorhyncha [sic]). Atti dell’Accademia Nazionale Italiana di Entomologia Rendiconti, 6, 3–10.
Servadei, A. (1967) Rhynchota (Heteroptera, Homoptera Auchenorrhyncha). Fauna d’Italia. Calderini, Bologna, x + 851 pp.
Servadei, A. (1969) I Rincoti endemici d’Italia. Memorie della Società Entomologica Italiana, 48, 417–439.
Slimani, N., Chen, P.-P., Nieser, N., Moulet, P., Ribeiro, J.R.I., Boumaïza, M. & Guilbert, E. (2016) Annotated check-list of semi-aquatic bugs of Tunisia, with detailed faunistic survey of north Tunisia (Hemiptera: Heteroptera: Gerromorpha). Entomologica Americana, 122, 55–71.
http://dx.doi.org/10.1664/15-RA-059Spuņģis, V. (2008) Fauna and ecology of terrestrial invertebrates in raised bogs in Latvia. Latvijas Entomologijas Biedriba, Riga, 80 pp.
Stichel, W. (1955) Fam. Hebridae Fb. In: Stichel, W. (Ed.) (1955–1956) Illustrierte Bestimmungstabellen der Wanzen. II. Europa (Hemiptera-Heteroptera Europae). Vol. 1. W. Stichel, Berlin-Hermsdorf, pp. 150–155.
Stichel, W. (1956) Liste der paläarktischen Hemiptera-Heteroptera. In: Stichel, W. (Ed.), (1955–1956) Illustrierte Bestimmungstabellen der Wanzen. II. Europa (Hemiptera-Heteroptera Europae). Vol. 1. W. Stichel, Berlin-Hermsdorf, pp. 157–168.
Stoffelen, E., Henderickx, H., Vercauteren, T., Lock, K. & Bosmans, R. (2013) De water- en oppervlaktewantsen van België. Fauna van België. [Aquatic and semiaquatic Heteroptera of Belgium. Fauna of Belgium]. Koninklijk Belgisch Instituut voor Natuurwetenschappen, Brussels, 254 pp. [in Dutch]
Strauss, G. & Niedringhaus, R. (2014) Die Wasserwanzen Deutschlands. Bestimmungsschlüssel für alle Nepo- und Gerromorpha. WABW, Scheeßel, [iv] + 66 pp.
Tamanini, L. (1979) Eterotteri acquatici (Heteroptera: Gerromorpha, Nepomorpha). Consiglio Nazionale delle Ricerche, Verona, 106 pp.
Thomson, C.G. (1871) Bidrag till Sveriges insect·fauna (2). In: Thomson, C.G. (Ed.), Opuscula Entomologica. Vol. 4. Lund, pp. 361–397.
Van Duzee, E.P. (1921) Characters of some new species of North American hemipterous insects, with one new genus. Proceedings of the California Academy of Sciences, Series 4, 11, 111–134.
Vinokurov, N.N. (2005) Materialy po faune poluzhestkokrylykh Norskogo zapovednika Amurskoy oblasti. Zhivotnyy mir Dal’nego Vostoka. Vyp. 5. [Materials to the fauna of Heteroptera of Norskyy zapovednik Reserve in Amurskaya oblast’ Region. Animal world of the Far East. Vol. 5.]. Blagoveshchensk, pp. 15–26. [in Russian]
Vinokurov, N.N., Kanyukova, E.V. & Golub, V.B. (2010) Katalog poluzhestkokrylykh nasekomykh (Heteroptera) Aziatskoy chasti Rossii. (Catalogue of the Heteroptera of the Asian part of Russia). Nauka, Novosibirsk, 323 pp. [in Russian, English abstract]
Wachmann, E., Melber, A. & Deckert, J. (2006) Wanzen. Band 1. Dipsocoromorpha, Nepomorpha, Gerromorpha, Leptopodomorpha, Cimicomorpha (Teil 1). Die Tierwelt Deutschlands. 77. Teil. Goecke & Evers, Keltern, 264 pp.
Wagner, E. (1954) Une espece d’Hebrus du midi de la France jusqu’alors méconnu (Hemiptera Heteroptera, Hebridae). Cahiers des Naturalistes (Nouvelle Série), 9, 67–69.
Wagner, E. (1957) Neogaeus (= Hebrus) franzi n. sp. Memorie della Società Entomologica Italiana, 36, 201–202.
Wagner, E. (1962) Beitrag zur Heteropteren-Fauna Macedoniens (Hem. Het.). Fragmenta Balcanica, 4, 115–122.
Westwood, J.O. (1840) An introduction to the modern classification of insects; founded on the narural habits and corresponding organisation of the different families. Vol. 2. Longman, Orme, Brown, Green & Longmans, London, pp. 353–587. [pp. xi + 352 published in 1839]
White, F.B. (1877) Descriptions of new species of heteropterous Hemiptera collected in the Hawaiian Islands by the Rev. T. Blackburn. – No. 1. Annals and Magazine of Natural History, Series 4, 20, 110–117.
http://dx.doi.org/10.1080/00222937708682204Wróblewski A. (1980) Pluskwiaki (Heteroptera). Fauna słodkowodna polski. Vol. 8. [True bugs (Heteroptera). Freshwater fauna of Poland. Vol. 8]. Państwowe Wydawnictwo Naukowe, Warszawa, Poznań, 154 pp. [in Polish]
Zettel, H. (2006) Neue Arten der Gattung Hebrus Curtis, 1833 (Heteroptera: Hebridae) aus Südostasien – 2. Teil. Linzer Biologische Beiträge, 38, 1009–1040.
Zettel, H. (2011) A contribution to the knowledge of Gerromorpha (Insecta: Hemiptera) of Myanmar, with seven new species, eight new records, and a catalogue. Annales des Naturhistorischen Museums Wien B, 112 [2010], 89–114.
Zettel, H. (2014) Annotated catalogue of the semi-aquatic bugs (Hemiptera: Heteroptera: Gerromorpha) of Luzon Island, the Philippines, with descriptions of new species. Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen, 66, 85–140.
Zettel, H. & Tran, A.D. (2015) Hebridae. In: Tran, A.D. (Ed.), Water bugs of Singapore and Peninsular Malaysia. Lee Kong Chian Natural History Museum, Singapore, pp. 70–79.
Zettel, H. & Tran, A.D. (2016) First notes on Hebrus (Insecta: Heteroptera: Hebridae) in Vietnam. Annales des Naturhistorischen Museums Wien B, 118, 55–59.
Zimmermann, G. (1982) Beiträge zur Wasserwanzen-Fauna Griechenlands I (Heteroptera, Nepomorpha, Gerromorpha, Saldidae). Marburger Entomologische Publikationen, 1 (7), 183–216.