Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2017-06-01
Page range: 1–18
Abstract views: 149
PDF downloaded: 1

An annotated catalogue of the gamasid mites associated with small mammals in Asiatic Russia. The family Haemogamasidae (Acari: Mesostigmata: Gamasina)

Omsk State University, 28 Adrianova Str., Omsk, Russian Federation, 644077 Saint-Petersburg State University, 7/9 Universitetskaya Emb., Saint-Petersburg, Russian Federation, 199034.
Laboratory of Arthropod-Borne Viral Infections, Omsk Research Institute of Natural Foci Infections, 7 Mira Str. Omsk, Russian Federation, 644080 Omsk State Pedagogical University, 14 Tukhachevskogo Emb., Omsk, Russian Federation, 644099
Acari Haemogamasidae Northern Asia parasitic gamasid mites small mammals host-parasite relationships species diversity

Abstract

We provide a list of the species of the family Haemogamasidae Oudemans, 1926 living in Asiatic Russia, with data on their synonymy, distribution, and relationships with mammal hosts. In total, 23 species of mites distributed between two genera (Eulaelaps Berlese, 1903, and Haemogamasus Berlese, 1889) are covered.

 

References

  1. Airoldi, J.-P., Solomon, L. & Duca, A.-V. (1989) Les Gamasides (Acari) des nids de la forme fousseuse du campagnol terrestre Arvicola terrestris (L.). Revue Suisse de Zoologie, 96, 161–189.
    https://doi.org/10.5962/bhl.part.117762

    Allred, D.M. (1969) Haemogamasid mites of Eastern Asia and Western Pacific with a key to species. Journal of Medical Entomology, 6, 103–108.
    https://doi.org/10.1093/jmedent/6.2.103

    Allred, D.M. & Beck, D.E. (1966) Mites of Utah mammals. Brigham Young University Science Bulletin, 8, 1–123.
    https://doi.org/10.5962/bhl.part.7441

    Asanuma, K. (1948) Notes on parasitic Acari collected from the Manchurian tarabagan (Marmota bobac sibirica). Seibutu, 3, 171–176.

    Asanuma, K. (1951a) Redescription of Haemogamasus kitanoi Asanuma, 1948, together with remarks on Acari parasitic on a Mongolian Pika, Ochotona daurica (Pallas), from Northwest Manchuria. Miscellaneous Reports of the Research Institute for Natural Resources, Tokyo, 22, 4–8.

    Asanuma, K. (1951b) Critical notes on the genera Euhaemogamasus Ewing and Haemogamasus Berlese, with description of a new species from Manchuria (Acari, Laelaptidae, Haeogamasinae). Miscellaneous Reports of the Research Institute for Natural Resources, Tokyo, 24, 15–21.

    Asanuma, K. (1952) Two new species of blood-sucking mites parasitic on the striped mouse, Apodemus agrarius, from Manchuria. Miscellaneous Reports of Research Institute of Natural Resources, 25, 86–92.

    Balashov, Yu. S. (2009) Acari and Insect Parasitism on Terrestrial Vertebrates. Nauka Publishers, Saint-Petersburg, 358 pp. [in Russian]

    Banks, N. (1909) New Canadian mites. Proceedings of the Entomological Society of Washington, 11, 133–143.

    Beaulieu, F., Dowling, A.P.G., Klompen, H., de Moraes, G.J. & Walter, D.E. (2011) Superorder Parasitiformes Reuter, 1909. In: Zhang, Z.-Q. (Ed.), Animal Biodiversity: An Outline of Higher Classification and Survey of Taxonomic Richness. Zootaxa, 3148, pp. 123–128.

    Berlese, A. (1889) Acari, Myriopoda et Scorpiones hucusque in Italia reperta. Tipografia del Seminario, Padova, 52 pp.

    Berlese, A. (1903a) Acari nuovi, manipulus IIus. Redia, 1, 258–280.

    Berlese, A. (1903b) Diagnosi di alcune nouve specie di Acari Italiani, mirmecofili et liberi. Zoologischer Anzeiger, 27, 12–28.

    Bondarchuk, A.S. (1976) A new species of mite of the genus Haemogamasus (Parasitiformes, Gamasoidea). Parazitologiya, 10, 191–193. [in Russian]

    Bregetova, N.G. (1949) Materials on the fauna of mites of the family Haemogamasidae. Parazitologicheskiy sbornik Zoologicheskogo Instituta Akademii Nauk SSSR, 11, 161–185. [in Russian]

    Bregetova, N.G. (1950) New species of gamasid mites (Acarina, Gamasoidea). Parazitologicheskiy sbornik Zoologicheskogo Instituta Akademii Nauk SSSR, 12, 13–16. [in Russian]

    Bregetova, N.G. (1955) The family Haemogamasidae Ouds. Opredeliteli po faune SSSR, izdavaemye Zoologicheskim Institutom Akademii Nauk SSSR, 59, 258–287. [in Russian]

    Bregetova, N.G. (1956a) Gamasid mites (Gamasoidea). Opredeliteli po faune SSSR, izdavaemye Zoologicheskim Institutom Akademii Nauk SSSR, 61, 1–247. [in Russian]

    Bregetova, N.G. (1956b) New data about mites of the family Haemogamasidae (Parasitiformes, Gamasoidea) of the USSR fauna. Zoologicheskiy Zhurnal, 1956, 1647–1656. [in Russian]

    Bregetova, N.G. (1977) Fam. Haemogamasidae Oudemans, 1926. In: Ghilyarov, M.S. & Bregetova, N.G. (Eds.), Key to the Soil Inhabiting Mites. Mesostigmata, Nauka, Leningrad, pp. 569–572. [in Russian]

    Bregetova, N.G. & Nel’zina, E.N. (1952) The suslik mite, Haemogamasus citelli Bregetova et Nelzina, sp. nov. (Gamasoidea, Haemogamasidae). Parazitologicheskiy sbornik Zoologicheskogo Instituta Akademii Nauk SSSR, 14, 71–74. [in Russian]

    Buyakova, T.G. (1982) A new species of mite of the genus Haemogamasus (Parasitiformes, Gamasina). In: Pronin, N.M. (Ed.), Zooparazitologiya Zabaikalya. Buryat Branch of the Soviet Academy of Sciences, Ulan-Ude, pp. 21–26. [in Russian]

    Buyakova, T.G. & Goncharova, A.A. (1962) A new species of mite of the genus Haemogamasus (Parasitiformes, Gamasoidea). Zoologicheskiy Zhurnal, 41, 760–763. [in Russian]

    Buyakova, T.G. & Goncharova, A.A. (1964) New species of gamasid mites (Parasitiformes, Gamasoidea). Zoologicheskiy Zhurnal, 43, 268–271. [in Russian]

    Canestrini, G (1884) Prospetto dell’acarofauna Italiana. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, Series 6, 2, 1563–1607.

    Casanueva, M.E., Berrios, A.M., Peredo, A.A.A. & Martinez, R.I. (1994) Mites associated on horses stables I. Androlaelaps casalis (Berlese), Haemogamasus horridus Michael and Proctolaelaps pygmaeus (Müller), the first record for Chile and redescription of the male (Acari: Mesostigmata). Gayana Zoologia, 58, 61–69.

    Cicek, H., Stanyukovich, M., Yağci, S., Aktaş, M. & Karaer, Z. (2008) Gamasine mite (Parasitiformes: Mesostigmata) infestations of small mammals (Mammalia: Rodentia, Insectivora) in Turkey. Türkiye Parazitoloji Dergisi, 32, 65–70.

    Costa, M. (1961) Mites associated with rodents in Israel. Bulletin of the British Museum (Natural History) Zoology, 8, 3–70.

    Davydova, M.S. (1966) Zoogeographical characteristics and morphological peculiarities of gamasid mites of the Altai. In: Cherepanov, A.I. (Ed.), Fauna i ekologia chlenistonogikh Sibiri. Nauka, Novosibirsk, pp. 136–147. [in Russian]

    Davydova, M.S., & Nikolsky, V.V. (1986) Gamasid Mites of Western Siberia. Nauka, Novosibirsk, 124 pp. [in Russian]

    Dowling, A.P.G. & OConnor, B.M. (2010) Phylogeny of Dermanyssoidea (Acari: Parasitiformes) suggests multiple origin of parasitism. Acarologia, 50, 113–129.
    https://doi.org/10.1051/acarologia/20101957

    Dugès, A.L. (1834) Recherches sur l’ordre des Acariens en général et la famille des Trombidiés en particulier. Annales des Sciences Naturelles. Zoologie, 2 (1), 5–46.

    Dusbabek, F. (1966) A contribution to the knowledge of parasitic mites from Mongolia (Acarina: Gamasides): results of the Mongolian-German biological expeditions since 1962, No. 9. Mitteilungen aus dem Zoologischen Museum in Berlin, 42, 43–58.

    Edler, A. & Mehl, R. (1972) Mites (Acari, Gamasina) from small mammals in Norway. Norsk Entomologisk Tidsskrift, 19, 133–147.

    Evans, G.O. & Till, W.M. (1966) Studies on the British Dermanyssidae (Acari: Mesostigmata). Part II. Classification. Bulletin of the British Museum (Natural History) Zoology, 14, 107–370.

    Evans, G.O. & Till, W.M. (1979) Mesostigmatic mites of Britain and Ireland. Transactions of the Zoological Society of London, 35, 139–270.
    https://doi.org/10.1111/j.1096-3642.1979.tb00059.x

    Ewing, H.E. (1925) New mites of the parasitic genus Haemogamasus. Proceedings of the Biological Society of Washington, 38, 137–144.

    Ewing, H.E. (1933) New genera and species of parasitic mites of the superfamily Parasitoidea. Proceedings of the United States National Museum, 2971, 1–14.
    https://doi.org/10.5479/si.00963801.82-2971.1

    Fang, W., Yao, L., Cao, X., Sun, Y., Zhang, X., Shao, L. & Shen, T. (2015) First molecular detection of Rickettsia felis-like organism in Eulaelaps stabularis from the Changbai Mountain Area of China. Journal of Parasitology, 101, 514–519. https://doi.org/10.1645/14-695.1

    Fonseca, F.d. (1960) Acarology notes. XLV. Investigation in Peru. Acarologia, 2, 1–34.

    Fyodorova, S.Zh. & Kharadov, A.V. (2012) A new gamasid mite species, Haemogamasus limneticus sp. n. (Parasitiformes: Gamasoidea), from Kyrgyzstan. Parazitologiya, 46, 272–278. [in Russian]
    https://doi.org/10.1134/s0013873812090126

    Garrett, D.A. & Allred, D.M. (1971) Mesostigmatid mites from Turkey, with keys to genera and species. Journal of Medical Entomology, 8, 292–298.
    https://doi.org/10.1093/jmedent/8.3.292

    Goncharova, A.A., Bondarchuk, A.S. & Vershinina, O.N. (1991) Gamasid mitesEctoparasites of Mammals in Transbaikalia. Chita State Medical University, Chita, 121 pp. [in Russian]

    Goncharova, A.A. & Buyakova, T.G. (1960) Bionomics of a gamasid mite Haemogamasus mandshuricus Vitzth. under conditions of Transbaikalia. Parazitologicheskiy sbornik Zoologicheskogo Instituta Akademii Nauk SSSR, 19, 155–163. [in Russian]

    Goncharova, A.A. & Buyakova, T.G. (1961) To the study of mites of the family Haemogamasidae (Parasitiformes, Gamasoidea) in the USSR. Zoologicheskiy Zhurnal, 40 (2), 276–280. [in Russian]

    Goncharova, A.A. & Buyakova, T.G. (1962) Bionomics of a gamasid mite Eulaelaps cricetuli Vitzthum in Transbaikalia. Zoologicheskiy Zhurnal, 41 (1), 139–143. [in Russian]

    Gwiazdowicz, D.J., Błoszyk, J., Bajerlein, D., Halliday, R.B. & Mizera, T. (2006) Mites (Acari: Mesostigmata) inhabiting nests of the white-tailed sea eagle Haliaeetus albicilla (L.) in Poland. Entomologica Fennica, 17, 366–372.

    Haitlinger, R. (1988) Haemogamasidae Oudemans, 1926 (Acari, Mesostigmata) of Poland. Polskie Pismo Entomologiczne, 58, 635–662.

    Halliday, B. (2011) Occurrence of the predatory mite Haemogamasus pontiger (Berlese) (Acari: Laelapidae) in Australia, with a review of its biology. Australian Journal of Entomology, 50, 61–64.
    https://doi.org/10.1111/j.1440-6055.2010.00776.x

    Harris, Ph.D., Paziewska, A., Zwolińska, I. & Siński, E. (2009) Seasonality of the ectoparasite community of woodland rodents in a Mazurian forest, Poland. Wiadomości Parazytologiczne, 55 (4), 377–388.

    Herrin, C.S. & Sage, R.D. (2012) Description of a new species of Haemogamasus (Mesostigmata, Laelapidae, Haemogamasinae) from Chubut, Río Negro and Neuquén Provinces, Argentina. Zookeys, 173, 51–77.
    https://doi.org/10.3897/zookeys.173.1592

    Hirst, S. (1914) Preliminary list of the Acari occurring on the brown rat (Mus norvegicus) in Great Britain, with the description of a new species (Haemogamasus oudemansi). Bulletin of Entomological Research, 5, 119–124.
    https://doi.org/10.1017/S0007485300033964

    Hull, J.E. (1918) Terrestrial Acari of the Tyne Province. Transactions of the Natural History Society of Northumberland, Durham, and Newcastle-upon-Tyne, 5, 13–89.

    Karg, W. (1971) Acari (Acarina), Milben. Unterordnung Anactinochaeta (Parasitiformes). Die freilebenden Gamasina (Gamasides), Raubmilben. Die Tierwelt Deutschlands, 59, 1–475.

    Karg, W. (1993) Acari (Acarina), Milben Parasitiformes (Anactinochaeta), Cohors Gamasina Leach, Raubmilben. Die Tierwelt Deutschlands, 59, 1–523.

    Keegan, H.L. (1946) Six new mites of the superfamily Parasitoidea. Transactions of the American Microscopical Society, 65, 69–77.
    https://doi.org/10.2307/3223286

    Keegan, H.L. (1951) The mites of the subfamily Haemogamasinae (Acari: Laelaptidae). Proceedings of the United States National Museum, 101, 203–268.
    https://doi.org/10.5479/si.00963801.101-3275.203

    Kim, M.H., Lee, H.P. & Noh, J.T. (1987) Study of laelaptid mites parasitising rodents in Korea. III. Mites of the genus Haemogamasus Berlese, 1889 (Acari: Mesostigmata). Korean Arachnology, 3 (1), 35–44.

    Koch, C.L. (1836) Deutschlands Crustaceen, Myriapoden und Arachniden. Ein Beitrag zur deutschen Fauna. Herrig-Schäffer, Regensburg, 4, 1–24 pls.

    Koch, C.L. (1839) Deutschlands Crustaceen, Myriapoden und Arachniden. Ein Beitrag zur deutschen Fauna. Herrig-Schäffer, Regensburg, 27, 1–24 pls.

    Koch, L. (1879) Arachniden aus Sibirien und Novaya Semlya, eingesammelt von der Schwedischen Expedition im Jahre 1875. Kungliga Svenska Vetenskapsakademiens handlingar, Series 3, 16, 3–136.

    Kozlowski, S. (1960) Contribution to study of the natural system of Acarina of the genus Haemogamasus Berlese, 1889 (Gamasides). Acta Parasitologica Polonica, 8 (26), 403–418.

    Lange, A.B. (1958) Superfamily Gamasoidea. In: Beklemishev, V.N. (Ed.), Key to arthropods injuring human health. Medgiz, Moscow, pp. 195–217. [in Russian]

    Li, W.X. (1991) The ecogeographical distribution of gamasides in Liaoning Province. Acta Entomologica Sinica, 34, 108–116.

    Lindquist, E.E., Krantz, G.W. & Walter, D.E. (2009) Order Mesostigmata. In: Krantz, G.W. & Walter, D.E. (Eds.), A Manual of Acarology. 3rd Edition. Texas Tech University Press, Lubbock, TX, pp. 124–232.

    Lopatina, Yu. V., Petrova-Nikitina, A.D. & Tamarina, N.A. (1991) Structure and changes in populations of Acari and Siphonaptera in burrows of Citellus musicus in the Central Caucasus. Entomological Review, 70 (2), 123–132.

    Lundquist, L. & Edler, A. (1979) Differential taxonomy of Haemogamasus nidi Michael, 1892 and H. nidiformis Bregetova, 1955 (Acari, Mesostigmata, Laelaptidae). Entomologia Scandinavia, 10, 73–78.
    https://doi.org/10.1163/187631279X00420

    Lundquist, L. (1990) Taxonomic and functional significance of the presternal area in Haemogamasus mites (Acari-Mesostigmata: Laelaptidae), with a revised key to Fennoscandian species. Entomologia Scandinavia, 21, 329–337.
    https://doi.org/10.1163/187631290X00256

    Mašán, P. & Fenda, P. (2010) A Review of the Laelapid Mites Associated with Terrestrial Mammals in Slovakia, with a Key to the European Species (Acari: Mesostigmata: Dermanyssoidea). Institute of Zoology, Slovakian Academy of Sciences, Bratislava, 187 pp.

    Michael, A.D. (1892) On the variations in the internal anatomy of the Gamasinae especially in that of the genital organs and their mode of coition. Transactions of the Linnean Society of London, Series 2, 5, 281–324.
    https://doi.org/10.1111/j.1096-3642.1892.tb00175.x

    Mitchell, R.M. (1977) Accounts of Nepalese Mammals and Analysis of the Host-ectoparasite Data by Computer Techniques. PhD Thesis. Iowa State University, Ames, 558 pp.

    Mit’kova, K., Berthová, L., Kalúz, S., Kazimírová, M., Burdová, L. & Kocianová, E. (2015) First detections of Rickettsia helvetica and R. monacensis in ectoparasitic mites (Laelapidae and Trombiculidae) infesting rodents in south-western Slovakia. Parasitology Research, 114, 2465–2472.
    https://doi.org/10.1007/s00436-015-4443-x

    Mosquera, S. (1988) Eucheyletia hardyi Baker, 1949 y Haemogamasus pontiger (Berlese, 1904), dos acaros nuevos para la fauna Argentina. Neotropica, 34, 73–76.

    Mrciak, M. (1958) Roztoce z radu Parasitiformes (Acari) z drobných cicavcov Vysokých Tatier. Zoologické listy (Brno), 7, 65–89.

    Nikulina, N.A. (1987) Gamasid mites (cohort Gamasina). In: Soboleva, R.G. (Ed.), Insects and mites of the Far East of Medical and Veterinary Importance. Nauka, Leningrad, pp. 216–234. [in Russian]

    Nikulina, N.A. (2004) A catalogue of parasitic Gamasina mites of mammals of Northern Eurasia (Russia). Aktsioner & Co, Saint-Petersburg, 170 pp. [in Russian]

    Oudemans, A.C. (1903) Notes on Acari. Eighth series. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 8, 69–92.

    Oudemans, A.C. (1913) Acarologisches aus Maulwurfsnestern. Archiv für Naturgeschichte, Abteilung A, 79, 108–200.

    Oudemans, A.C. (1926) Acarologisches Aanteekeningen LXXXII. Entomologische Berichten, 7, 119–126.

    Petrova, A.D. & Taskayeva, E.Z. (1974) Parasitic gamasid mites of rodents of the southeastern part of the Peoples Republic of China. Byulleten’ Moskovskogo Obshchestva Ispytateley Prirody, otdel biologicheskiy, 79, 44–50. [in Russian]

    Piryanik, G.I. (1962) Gamasid Mites of Mouse-like Rodents of the Ukrainian Forest-steppe. Kiev University Press, Kiev, 176 pp. [in Russian]

    Prasad, V. (1974) Parasitic mesostigmatic mites from Nepal (Acarina: Mesostigmata). Oriental Insects, 8, 63–70.
    https://doi.org/10.1080/00305316.1974.10434444

    Přívora, M. & Samšiňák, K. (1957) Zástupce čeledě Haemogamasidae jako parasit člověka a několik noznámek k systematice této čeledě. Československá Parasitologie, 4, 267–274.

    Punda, H. (1972) On variation in Haemogamasus nidi (Acari, Gamasoidae). Annales Zoologici, 29, 83–89.

    Radovsky, F.J. (1960) Haemogamasus liponyssoides hesperus, n. ssp., with a discussion of the H. liponyssoides complex (Acarina: Haemogamasidae). The Journal of Parasitology, 46, 401–409.
    https://doi.org/10.2307/3275129

    Radovsky, F.J. & Gettinger, D. (1999) Acanthochelinae, new subfamily, (Acari: Parasitiformes, Laelapidae), with redescription of Acanthochela chilensis Ewing and descriptions of a new genus and species from Argentina. International Journal of Acarology, 25, 77–90.
    https://doi.org/10.1080/01647959908683619

    Redington, B.C. (1970) Studies on the morphology and taxonomy of Haemogamasus reidi Ewing, 1925 (Acari: Mesostigmata). Acarologia, 12, 643–667.

    Rounsevell, D. (1978) Populations of introduced arthropods of Australasian antarctic stations. Pacific Insects, 18, 199–202.

    Saaristo, M.I. & Marusik, Yu.M. (2003) Revision of the Holarctic spider genus Oreoneta Kulczyński, 1894 (Arachnida: Aranei: Linyphiidae). Arthropoda Selecta, 12, 207–249.

    Sellnick, M. (1974) Einige neue Milbenarten (Acari) von Island und Bemerkungen über bekannte. Entomologica Scandinavica, 8, 209–216.
    https://doi.org/10.1163/187631274X00245

    Senotrusova, V.G. (1972) On morphology of the pre-imaginal and imaginal stages of development in the mite Haemogamasus kitanoi Asan., 1948. Parazitologiya, 6, 246–251. [in Russian]

    Senotrusova, V.G. (1973) Biology and distribution of the mite Haemogamasus kitanoi Asan. in Kazakhstan. Parazitologiya, 7, 307–311. [in Russian]

    Senotrusova, V.G. (1976) Eulaelaps novus Vitzth., 1925, a separate species of mites. Trudy Instituta Zoologii Akademii Nauk Kazakhskoi SSR, 36, 86–91. [in Russian]

    Senotrusova, V.G. (1987) Gamasid mitesparasites of wild animals in Kazakhstan. Nauka, Alma-Ata, 224 pp. [in Russian]

    Sludsky, A.A. (2014) The epizootology of the plague (a review of studies and hypotheses). Saratov, 1, 1–313. [in Russian]

    Strandtmann, R.W. & Wharton, G.W. (1958) A Manual of Mesostigmatid Mites Parasitic on Vertebrates. University of Maryland, College Park, 330 pp.

    Thorell, T. (1872) Om några Arachnider från Grönland. Öfversigt af Kongliga Svenska Vetenskaps-akademiens Förhandlingar, 29, 151–166.

    Trägårdh, I. (1902) Revision der von Thorell aus Grönland, Spitzbergen und der Bären-Insel und von L. Koch aus Sibirien und Novaja Semlja beschrieben Acariden. Zoologischer Anzeiger, 25, 56–62.

    Trågärdh, I. (1904) Monographie der arktischen Acariden. Gustav Fischer, Jena, 78 pp.
    https://doi.org/10.5962/bhl.title.5373

    Trägårdh, I. (1912) Biospeleologica, XXII. Acari (first series). Archives de Zoologie Expérimentale et Générale, Séries 5, 8, 519–620.

    Troitsky, V.A. (1973) Some morphological peculiarities in the structure of chelicerae of bloodsucking mites of the fam. Haemogamasidae (Gamasoidea). Parazitologiya, 7, 526–530. [in Russian]

    Turk, F.A. (1945) Studies of Acari. Second series: descriptions of new species and notes on established forms of parasitic mites. Parasitology, 36, 133–141.
    https://doi.org/10.1017/S0031182000012099

    Uchikawa, K. & Rack, G. (1979) Eulaelaps stabularis (Koch, 1839) and Eulaelaps oudemansi Turk, 1945 (Mesostigmata: Haemogamasidae). Acarologia, 20, 163–172.

    Valiente Moro, C., Chauve, C. & Zenner, L. (2005) Vectorial role of some dermanyssoid mites (Acari, Mesostigmata, Dermanyssoidea). Parasite, 12, 99–109.
    https://doi.org/10.1051/parasite/2005122099

    Vinarski, M.V. & Korallo-Vinarskaya, N.P. (2016) An annotated catalogue of the gamasid mites associated with small mammals in Asiatic Russia. The family Laelapidae s. str. (Acari: Mesostigmata: Gamasina). Zootaxa, 4411 (3), 223–245. https://doi.org/10.11646/zootaxa.4111.3.2

    Vitzthum, H. (1930) Milben als Pestträger? Zoologischer Jahrbücher. Abteilung für Systematik, Ökologie und Geographie der Tiere, 60, 381–428.

    Wen, T.-W. (1976) A new subfamily Eulaelapinae Wen, and three new species of the genus Eulaelaps Berlese (Gamasides, Haemogamasidae). Acta Entomologica Sinica, 19, 348–356. [in Chinese]

    Whitaker, J.O. Jr., Walters, B.L., Caster, L.K., Ritzi, C.M. & Wilson, N. (2007) Host and distribution lists of mites (Acari), parasitic and phoretic, in the hair or on the skin of North American wild mammals north of Mexico: records since 1974. Faculty Publications from the Harold W. Manter Laboratory of Parasitology, 1–173.

    Whitaker, J.O. Jr. & Wilson, N. (1974) Host and distribution lists of mites (Acari), parasitic and phoretic, in the hair of wild mammals of North America, north of Mexico. American Midland Naturalist, 91, 1–67.
    https://doi.org/10.2307/2424511

    Williams, G.L., Smiley, R.L. & Redington, B.C. (1978) A taxonomic study of the genus Haemogamasus in North America, with descriptions of two new species (Acari: Mesostigmata, Laelaptidae). International Journal of Acarology, 4, 235–273.
    https://doi.org/10.1080/01647957808683122

    Willmann, C. (1952) Parasitische Milben an Kleinsäugern. Zeitschrift für Parasitenkunde, 15, 392–428.
    https://doi.org/10.1007/BF00259346

    Womersley, H. (1937) Acarina. Scientific reports of the Australasian Antarctic Expedition, 1911–1914, Series C, 10, 1–24.

    Yakimenko, V.V., Dekonenko, A.Ye., Mal’kova, M.G., Kuz’min, I.V., Tantsev, A.K., Dzagurova, T.K. & Tkachenko, Ye.A. (2000) Spread of hantaviruses in West Siberia. Meditsinskaya Parazitologiya i Parazitarnye Bolezni, 3, 21–28. [in Russian]

    Žákovská, A., Janouškovcová, E., Pejchalová, K., Halouzka, J. & Dendis, M. (2008) Identification and characterization of 31 isolates of Borrelia burgdorferi (Spirochaetales, Spirochaetaceae) obtained from various hosts and vectors using PCR-RFLP and SDS-PAGE analysis. Acta Parasitologica, 53, 186–192.
    https://doi.org/10.2478/s11686-008-0028-5

    Zemskaya, A.A. (1973) Parasitic Gamasid Mites and Their Medical Importance. Meditsina Publishing House, Moscow, 168 pp. [in Russian]

    Zemskaya, A.A. & Piontkovskaya, S.P. (1957) Gamasid mites of the Altai zokor from areas of newly developed lands in Eastern Kazakhstan region. Parazitologicheskiy sbornik Zoologicheskogo Instituta Akademii Nauk SSSR, 17, 38–44. [in Russian]

    Zhou, M.-S., Gu, Y.M. & Wen, T.H. (1995) Mites of the Haemogamasidae in China with description of a new genus (Acari: Parasitiformes). Acta Zootaxonomica Sinica, 20 (2), 172–175.