Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2017-09-04
Page range: 201–224
Abstract views: 804
PDF downloaded: 228

The Wayward Dog: Is the Australian native dog or Dingo a distinct species?

Animal Biosecurity & Food Safety, NSW Department of Primary Industries, Orange, New South Wales 2800, Australia. School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, NSW 2052. Division of Mammals, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013-7012, USA.
School of Archaeology & Anthropology, Australian National University, Canberra, ACT 0200, Australia.
Vertebrate Pest Research Unit, Biosecurity NSW, NSW Department of Primary Industries, Orange, New South Wales 2800, Australia. School of Environmental & Rural Science, University of New England, Armidale, NSW 2351, Australia.
Division of Mammals, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013-7012, USA.
Australian Museum Research Institute, Australian Museum, 1 William St. Sydney, NSW 2010, Australia.
School of Archaeology & Anthropology, Australian National University, Canberra, ACT 0200, Australia.
School of Biological Sciences, Environment Institute, and ARC (Australian Research Council) Centre for Australian Biodiversity and Heritage, University of Adelaide, Adelaide, SA 5005, Australia.
Mammalia behaviour Dingo dog domesticate free-roaming dog genetics hybridisation morphology New Guinea Singing Dog reproduction species concept taxonomy

Abstract

The taxonomic identity and status of the Australian Dingo has been unsettled and controversial since its initial description in 1792. Since that time it has been referred to by various names including Canis dingo, Canis lupus dingo, Canis familiaris and Canis familiaris dingo. Of these names C. l. dingo and C. f. dingo have been most often used, but it has recently been proposed that the Australian Dingo should be once again recognized as a full species—Canis dingo. There is an urgent need to address the instability of the names referring to the Dingo because of the consequences for management and policy. Therefore, the objective of this study was to assess the morphological, genetic, ecological and biological data to determine the taxonomic relationships of the Dingo with the aim of confirming the correct scientific name. The recent proposal for Canis dingo as the most appropriate name is not sustainable under zoological nomenclature protocols nor based on the genetic and morphological evidence. Instead we proffer the name C. familiaris for all free-ranging dogs, regardless of breed and location throughout the world, including the Australian Dingo. The suggested nomenclature also provides a framework for managing free-ranging dogs including Dingoes, under Australian legislation and policy. The broad principles of nomenclature we discuss here apply to all free-roaming dogs that coexist with their hybrids, including the New Guinea Singing Dog.

 

References

  1. Alberch, P. (1985) Developmental constraints: why St. Bernard often have extra digit and poodles never do. American Naturalist, 126 (3), 430–433.

    https://doi.org/10.1086/284428

    Allen, B., Fleming, P.J.S., Allen, L.R., Engeman, R.M., Ballard, G. & Leung, L.K.-P. (2013) As clear as mud: a critical review of the evidence for the ecological role of the Australian dingo. Biological Conservation, 159, 158–174.

    https://doi.org/10.1016/j.biocon.2012.12.004

    Allen, B.L., Higginbottom, K., Bracks, J.H., Davies, N. & Baxter, G.S. (2015a) Balancing dingo conservation with human safety on Fraser Island: the numerical and demographic effects of humane destruction of dingoes. Australasian Journal of Environmental Management, 22 (2), 197–215.

    https://doi.org/10.1080/14486563.2014.999134

    Allen, B.L., Allen, L.R. & Leung, L.K-P. (2015b) Interactions between two naturalised invasive predators in Australia: are feral cats suppressed by dingoes? Biological Invasions, 17, 761–776.

    https://doi.org/10.1007/s10530-014-0767-1

    Aplin, K.P., Suzuki, H., Chinen, A.A., Chesser, R.T., ten Have, J., Donnellan, S.C., Austin, J., Frost, A., Gonzalez, J.-P., Herbreteau, V., Catzeflis, F., Soubrier, J., Fang, Y.-P., Robins, J., Matisoo-Smith, E., Bastos, A.D.S., Maryanto, I., Sinaga, M.H., Denys, C., Yap, G., Van Den Bussche, R.A., Conroy, C., Rowe, K. & Cooper, A. (2011) Multiple geographic origins of commensalism and complex dispersal history of black rats. PLoS ONE, 6 (11), e26357.

    https://doi.org/10.1371/journal.pone.0026357

    Archer-Lean, C., Wardell-Johnson, A., Conroy, G. & Carter, J. (2015) Representations of the dingo: contextualising iconicity. Australasian Journal of Environmental Management, 22 (2), 181–196.

    https://doi.org/10.1080/14486563.2014.985268

    Archibald, A.L., Cockett, N.E., Dalrymple, B.P., Faraut, T., Kijas, J.W., Maddox, J.F., McEwan, J.C., Oddy, H.V., Raadsma, H.W. & Wade, C. (2010) The sheep genome reference sequence: a work in progress. Animal Genetics, 41 (5), 449–453.

    https://doi.org/10.1111/j.1365-2052.2010.02100.x

    Ardalan, A., Oskarsson, M., Natanaelsson, C., Wilton, A., Ahmadian, A. & Savolainen, P. (2012) Narrow genetic basis for the Australian dingo confirmed through analysis of paternal ancestry. Genetica, 140 (1–3), 65–73.

    https://doi.org/10.1007/s10709-012-9658-5

    Atkinson, J. (1826) An Account of the State of Agriculture and Grazing in New South Wales. J Cross, London, 146 pp.

    Axelsson, E., Ratnakumar, A., Arendt, M.-J., Maqbool, K., Webster, M.T., Perloski, M., Liberg, O., Arnemo, J.M., Hedhammar, A. & Lindblad-Toh, K. (2013) The genomic signature of dog domestication reveals adaptation to a starch-rich diet. Nature, 495 (7441), 360–364.

    https://doi.org/10.1038/nature11837

    Baker, R.J. & Bradley, R.D. (2006) Speciation in mammals and the genetic species concept. Journal of Mammalogy, 87 (4), 643–662.

    https://doi.org/10.1644/06-MAMM-F-038R2.1

    Balme, J. & O'Connor, S. (2016) Dingoes and Aboriginal social organization in Holocene Australia. Journal of Archaeological Science: Reports, 7, 775–781.

    Bannasch, D., Bannasch, M., Ryun, J., Famula, T. & Pedersen, N. (2005) Y chromosome haplotype analysis in purebred dogs. Mammalian Genome, 16 (4), 273–280.

    https://doi.org/10.1007/s00335-004-2435-8

    Barker, B.C.W. & Macintosh, A. (1979) The dingo-a review. Archaeology and Physical Anthropology in Oceania, 14 (1), 27–53.

    Bellwood, P. (1997) Prehistory of the Indo-Malaysian Archipelago. University of Hawaii Press, Honolulu, 384 pp.

    Bino, R. (1996) Notes on behaviour of New Guinea singing dogs (Canis lupus dingo). Science in New Guinea, 22, 43–47.

    Boddaert, P. (1785) Elenchus Animalium. Volume 1. Sistens Quadrupedia huc usque nota eorumque varietates. C.R. Hake, Rotterdam, 174 pp.

    Bohlken, H. (1958) Vergleichende Untersuchungen an Wildrindern (Tribus Bovini Simpson, 1945). Zoologische Jahrbücher. Abteilung für Allgemeine Zoologie und Physiologie der Tiere, 68, 113–202.

    Bohlken, H. (1961) Der Kouprey, Bos (Bibos) sauveli Urbain 1937. Zeitschrift für Saugetierkunde, 26, 193–254.

    Brown, S.K., Pedersen, N.C., Jafarishorijeh S., Bannasch, D.L., Ahrens, K.D., Wu, J.T., Okon, M. & Sacks, B.N. (2011) Phylogenetic distinctiveness of Middle Eastern and Southeast Asian village dog Y chromosomes illuminates dog origins. PLoS ONE, 6, e28496.

    https://doi.org/10.1371/journal.pone.0028496

    Butler, J.R.A., du Toit, J.T. & Bingham, J. (2004) Free-ranging Domestic Dogs (Canis familiaris) as predators and prey in rural Zimbabwe: threats of competition and disease to large wild carnivores. Biological Conservation, 115 (3), 369–378.

    https://doi.org/10.1016/S0006-3207(03)00152-6

    Cairns, K.M., Wilton, A.N. & Ballard, J.W.O. (2011) The identification of dingoes in a background of hybrids. In: Urbano, K.V. (Ed.), Advances in Genetics Research. Nova Science Publishers, New York, pp. 309–327.

    Cairns, K.M. & Wilton, A.N. (2016) New insights on the history of canids in Oceania based on mitochondrial and nuclear data. Genetica, 144, 553–565.

    https://doi.org/10.1007/s10709-016-9924-z

    Catling, P.C. (1979) Seasonal variation in plasma testosterone and the testis in captive male dingoes, Canis familiaris dingo. Australian Journal of Zoology, 27 (6), 939–944.

    https://doi.org/10.1071/zo9790939

    Catling, P.C., Corbett, L.K. & Newsome, A.E. (1992) Reproduction in captive and wild dingoes (Canis familiaris dingo). Wildlife Research, 19 (2), 195–205.
    https://doi.org/10.1071/WR9920195

    Christie, D.W. & Bell, E.T. (1971) Some observations on the seasonal incidence and frequency of oestrus in breeding bitches in Britain. Journal of Small Animal Practice, 12 (3), 159–167.

    https://doi.org/10.1111/j.1748-5827.1971.tb06213.x

    Ciucci, P., Lucchini, V., Boitani, L. & Randi, E. (2003) Dewclaws in wolves as evidence of admixed ancestry with dogs. Canadian Journal of Zoology, 81 (12), 2077–2081.

    https://doi.org/10.1139/Z03-183

    Clutton-Brock, J., Corbet, G.B. & Hills, M. (1976) A review of the family Canidae, with a classification by numerical methods. Bulletin of the British Museum (Natural History), Zoology Series, 29 (3), 117–199.

    https://doi.org/10.5962/bhl.part.6922

    Clutton-Brock, J., Corbett, L.K., Harden, B. & Robertshaw, J. (1990) Dingo (Canis familiaris dingo). In: Ginsberg, J.R. & Macdonald, D.W. (Eds.), Foxes, Wolves, Jackals and Dogs: An Action Plan for the Conservation of Canids. International Union for Conservation of Nature and Natural Resources, Gland, pp. 53–54.

    Clutton-Brock, J. (1995) Origins of the dog: domestication and early history. In: Serpell, J. (Ed.), Domestic Dog: Its Evolution, Behavior and Interactions with People. Cambridge University Press, Cambridge, pp. 2–19.

    Clutton-Brock, J. (2015) Naming the scale of nature. In: Behie, A.M. & Oxenham, M.F. (Eds.), Taxonomic Tapestries: the Threads of Evolutionary, Behavioural and Conservation Research. Australian National University Press, Canberra, pp.171–182.

    Cole, S.R., Baverstock, P.R. & Green, B. (1977) Lack of genetic differentiation between Domestic Dogs and dingoes at a further 16 loci. Australian Journal of Experimental Biology and Medical Science, 55 (2), 229–232.

    https://doi.org/10.1038/icb.1977.17

    Corbet, G.B. & Clutton-Brock, J. (1984) Appendix: Taxonomy and nomenclature. In: Mason, I.L. (Ed.), Evolution of Domesticated Animals. Longman, London, pp. 434–438.

    Corbett, L.K. (1985) Morphological comparisons of Australian and Thai dingoes: a reappraisal of dingo status, distribution and ancestry. Proceedings of the Ecological Society of Australia, 13, 277–291.

    Corbett, L. (1995) The Dingo in Australia and Asia. 1st Edition. J.B. Books, Marleston, 200 pp.

    Corbett, L. (2001) The conservation status of the dingo Canis lupus dingo in Australia, with particular reference to New South Wales. In: Dickman, C.R. & Lunney, D. (Eds.), A Symposium on the Dingo. Royal Zoological Society of New South Wales, Sydney, pp. 10–19.

    Corbett, L. (2004) Dingo. In: Sillero-Zubiri, C., Hoffmann, M. & Macdonald, D.W. (Eds.), Canids: Foxes, Wolves, Jackals and Dogs. International Union for Conservation of Nature and Natural Resources, Gland, pp. 223–230.

    Corbett, L. (2006) The Australasian Dingo. In: Merrick, J.R., Archer, M., Hickey, G.M. & Lee, M.S.Y. (Eds.), Evolution and Biogeography in Australia. Auscipub Pty Ltd., Sydney, pp. 745–751.

    Corbett, L. (2008) Dingo Canis lupus. In: Van Dyck, S. & Strahan, R. (Eds.), The Mammals of Australia. New Holland, Sydney, pp. 737–739.

    Coyne, J.A. & Orr, H.A. (2004) Speciation. Sinauer Associates, Inc., Publishers, Sunderland, Massachusetts, 545 pp.

    Cracraft, J. (1983) Species concepts and speciation analysis. Current Ornithology, 1, 159–187.

    https://doi.org/10.1007/978-1-4615-6781-3_6

    Crowther, M.S., Fillios, M., Colman, N. & Letnic, M. (2014) An updated description of the Australian dingo (Canis dingo Meyer, 1793). Journal of Zoology, 293 (3), 192–203.

    https://doi.org/10.1111/jzo.12134

    Daniels, M.J. & Corbett, L. (2003) Redefining introgressed protected mammals: when is a wildcat a wildcat and a dingo a wild dog? Wildlife Research, 30 (3), 213–218.

    https://doi.org/10.1071/WR02045

    Darwin, C. (1868) The Variation of Animals and Plants Under Domestication. 2nd Edition. John Murray, London, 411 pp.

    Dennler de La Tour, G. (1968) Zur Frage der Haustier-Nomenklaturm. Säugetierkundliche Mitteilungen, 16, 1–20.

    Desmarest, A.G. (1821) Mammalogie, ou description des espèces de mammifères. Premier Partie, contenant les orders des bimanes, des quadrumanes et des carnassiers. V. Agasse, Paris, 547 pp.

    https://doi.org/10.5962/bhl.title.59887

    de Queiroz, K. (2007) Species concepts and species delimitation. Systematic Biology, 56 (6), 879–886.

    https://doi.org/10.1080/10635150701701083

    Ding, Z.-L., Oskarsson, M., Ardalan, A., Angleby, H., Dahlgren, L.-G., Tepeli, C., Kirkness, E., Savolainen, P. & Zhang, Y.-P. (2012) Origins of Domestic Dog in Southern East Asia is supported by analysis of Y-chromosome DNA. Heredity, 108 (5), 507–514.

    https://doi.org/10.1038/hdy.2011.114

    Dinets, V. (2015) The Canis tangle: a systematics overview and taxonomic recommendations. Vavilovskii Zhurnal Genetiki i SelektsiiVavilov Journal of Genetics and Breeding, 19, 286–291.
    https://doi.org/10.18699/VJ15.036

    Druzhkova, A.S., Thalmann, O., Trifonov, V.A., Leonard, J.A., Vorobieva, N.V., Ovodov, N.D., Graphodatsky, A.S. & Wayne, R.K. (2013 Ancient DNA analysis affirms the canid from Altai as a primitive dog. PLoS ONE, 8 (3), e57754.

    https://doi.org/10.1371/journal.pone.0057754

    Dwyer, P.D. & Minnegal, M. (2016) Wild dogs and village dogs in New Guinea: were they different? Australian Mammalogy, 38, 1–11.

    https://doi.org/10.1071/AM15011

    Elledge, A.E., Leung, L.K.P., Allen, L.R., Firestone, K. & Wilton, A.N. (2006) Assessing the taxonomic status of dingoes Canis familiaris dingo for conservation. Mammal Review, 36 (2), 142–156.

    https://doi.org/10.1111/j.1365-2907.2006.00086.x

    Elledge, A.E., Allen, L.R., Carlsson, B.L., Wilton, A.N. & Leung, L.K.P. (2008) An evaluation of genetic analyses, skull morphology and visual appearance for assessing dingo purity: implications for dingo conservation. Wildlife Research, 35 (8), 812–820.

    https://doi.org/10.1071/WR07056

    Etheridge, R. Jr. (1916) The cylindro-conical and cornute stone implements of western New South Wales; The warrigal, or 'Dingo' introduced or indigenous? Memoirs of the Geological Survey of New South Wales, Ethnological Series, 2, 1–70.

    Fiedel, S.J. (2000) The peopling of the New World: Present evidence, new theories, and future directions. Journal of Archaeological Research, 8 (1), 39–103.
    https://doi.org/10.1023/A:1009400309773

    Fillios, M.A. & Taçon, P.S.C. (2016) Who let the dogs in? A review of the recent genetic evidence for the introduction of the dingo to Australia and implications for the movement of people. Journal of Archaeological Science: Reports, 7, 782–792.
    https://doi.org/10.1016/j.jasrep.2016.03.001

    Finlayson, H.H. (1939) On mammals from the Lake Eyre Basin. Part IV. The Monodelphia. Transactions of the Royal Society of South Australia, 63, 88–118.

    Fleming, P., Corbett, L., Harden, R. & Thompson, P. (2001) Managing the Impacts of Dingoes and Other Wild Dogs. Bureau of Rural Sciences, Canberra, 186 pp.

    Fleming, P.J.S., Allen, B.L., Allen, L.R., Ballard, G., Bengsen, A.J., Gentle, M.N., McLeod, L.J., Meek, P.D. & Saunders, G.R. (2014) Management of wild canids in Australia: free-ranging dogs and red foxes. In: Glen, A.S. & Dickman, C.R. (Ed.), Carnivores of Australia: Past, Present and Future. CSIRO Publishing, Melbourne, pp. 105–149.

    Frankham, R, Ballou, J.D., Eldridge, M.D.B., Lacy, R.C., Ralls, K., Dudash, M.R. & Fenster, C.B. (2011) Predicting the probability of outbreeding depression. Conservation Biology, 25, 465–475.

    https://doi.org/10.1111/j.1523-1739.2011.01662.x

    Frankham, R., Ballou, J.D., Dudash, M.R., Eldridge, M.D.B., Fenster, C.B., Lacy, R.C., Mendelson III, J.R., Porton, I.J., Ralls, K. & Ryder, O.A. (2012) Implications of different species concepts for conserving biodiversity. Biological Conservation, 153, 25–31.

    https://doi.org/10.1016/j.biocon.2012.04.034

    Freedman, A.H., Gronau, I., Schweizer, R.M., Ortega-Del Vecchyo, D., Han, E., Silva, P.M., Galaverni, M., Fan, Z., Marx, P., Lorente-Galdos, B., Beale, H., Ramirez, O., Hormozdiari, F., Alkan, C., Vilà, C., Squire, K., Geffen, E., Kusak, J., Boyko, A.R., Parker, H.G., Lee, C., Tadigotla, V., Siepel, A., Bustamante, C.D., Harkins, T.T., Nelson, S.F., Ostrander, E.A., Marques-Bonet, T., Wayne, R.K. & Novembre, J. (2014) Genome sequencing highlights the dynamic early history of dogs. PLoS Genetics, 10, e1004016.

    https://doi.org/10.1371/journal.pgen.1004016

    French, D.D., Corbett, L.K. & Easterbee, N. (1988) Morphological discriminants of Scottish wildcats (Felis silvestris), domestic cats (F. catus) and their hybrids. Journal of Zoology, 214, 235–259.

    https://doi.org/10.1111/j.1469-7998.1988.tb04719.x

    García-Moreno, J., Matocq, M.D., Roy, M.S., Geffen, E. & Wayne, R.K. (1996) Relationships and genetic purity of the endangered Mexican Wolf based in analysis of microsatellite loci. Conservation Biology, 10 (2), 376–389.

    https://doi.org/10.1046/j.1523-1739.1996.10020376.x

    Gentry, A., Clutton-Brock, J. & Groves, C.P. (1996) Case 3010. Proposed conservation of usage of mammal specific names based on wild species which are antedated by or contemporary with those based on domestic animals. Bulletin of Zoological Nomenclature, 53, 28–37.

    https://doi.org/10.5962/bhl.part.14102

    Gentry, A., Clutton-Brock, J. & Groves, C.P. (2004) The naming of wild animal species and their domestic derivatives. Journal of Archaeological Science, 31 (5), 645–651.

    https://doi.org/10.1016/j.jas.2003.10.006

    Geraldes, A., Ferrand, N. & Nachman, M.W. (2006) Contrasting patterns of introgression at X-linked loci across the hybrid zone between subspecies of the European rabbit (Oryctolagus cuniculus). Genetics, 173 (2), 919–933.

    https://doi.org/10.1534/genetics.105.054106

    Germonpré, M., Sablin, M.V., Stevens, R.E., Hedges, R.E.M., Hofreiter, M., Stiller, M. & Deprés, V.R. (2009) Fossil dogs and wolves from Palaeolithic sites in Belgium, the Ukraine and Russia: osteometry, ancient DNA and stable isotopes. Journal of Archaeological Science, 36 (2), 473–490.

    https://doi.org/10.1016/j.jas.2008.09.033

    Germonpré, M., Sablin, M.V., Lázničková-Galetová, M., Deprés, V., Stevens, R.E., Stiller, M. & Hofreiter, M. (2015) Palaeolithic dogs and Pleistocene wolves revisited: a reply to Morey (2014). Journal of Archaeological Science, 54, 210–216.

    https://doi.org/10.1016/j.jas.2014.11.035

    Goebel, T. (1999) Pleistocene human colonization of Siberia and peopling of the Americas: An ecological approach. Evolutionary Anthropology, 8 (6), 208–227.

    https://doi.org/10.1002/(sici)1520-6505(1999)8:6<208::aid-evan2>3.0.co;2-m

    Gollan, J.K. (1982) Prehistoric dingo. Unpublished PhD thesis, Australian National University, Canberra, 463 pp.

    Gompper, M. (2014) The dog-human-wildlife interface: assessing the scope of the problem. In: Gompper, M. (Ed.), Free-ranging Dogs and Wildlife Conservation. Oxford University Press, Oxford, pp. 9–54.

    Gonzalez, A. (2012) The Pariah case: Some comments on the origin and evolution of primitive dogs and on the taxonomy of related species. PhD Thesis, Australian National University, Canberra, 362 pp.

    Gould, J. (1859 [1845–1863]) The Mammals of Australia. Vol. III. The Author, London, 2 pls. [text to pls 51–52]

    Gray, J.E. (1826) Appendix B. Vertebrata. Mammalia. In: King, P.P. (Ed.), Narrative of a Survey of the Intertropical and Western Coasts of Australia. Performed between the years 1818 and 1822. With an Appendix, containing various subjects relating to hydrography and natural history. Vol. 2. J. Murray, London, pp. 412–415.

    Gray, M.M., Granka, J., Bustamante, C.D., Sutter, N., Boyko, A., Zhu, L., Ostrander, E.A. & Wayne, R.K. (2009) Linkage disequilibrium and demographic history of wild and domestic canids. Genetics, 181 (4), 1493–1505.

    https://doi.org/10.1534/genetics.108.098830

    Grayson, D.K. (1988) Danger Cave, Last Supper Cave, and Hanging Rock Shelter: The Faunas. Anthropological Papers of the American Museum Natural History, 66 (1), 1–130.

    Gregory, J.W. (1906) The Dead Heart of Australia. John Murray, London, 371 pp.

    Groves, C.P., Ziccardi, F. & Toschi, A. (1966) Sull'asino selvatico africano. Ricerche di Zoologia Applicata alla Caccia, Bologna Supplement, 5, 1–30.

    Groves, C.P. (1995) On the nomenclature of domestic animals. The Bulletin of Zoological Nomenclature, 52, 137–141.

    https://doi.org/10.5962/bhl.part.6749

    Groves, C.P. (2001) Primate Taxonomy. Smithsonian Institution, Washington, D.C., 350 pp.

    Groves, C. (2004) The what, why and how of primate taxonomy. International Journal of Primatology, 25 (5), 1105–1126.

    https://doi.org/10.1023/b:ijop.0000043354.36778.55

    Groves, C. & Grubb, P. (2011) Ungulate Taxonomy. Johns Hopkins University Press, Baltimore, 317 pp.

    Harrop, A.E. (1960) The physiology of reproduction in the dog and bitch. In: Harrop, A.E. (Ed.), Reproduction in the Dog. Balliére, Tindall and Cox, London, pp. 64–86.

    Hedrick, P.W., Miller, P.S., Geffen, E. & Wayne, R.K. (1997) Genetic evaluation of the three captive Mexican Wolf lineages. Zoo Biology, 16 (1), 47–69.

    https://doi.org/10.1002/(SICI)1098-2361

    Howard, A.J. (1990) Hall’s heelers. In: Warner, R.M. (Ed.), Over-Halling the Colony: George Hall Pioneer. Southwood Press, Sydney, pp. 26–28.

    Hunter, J. (1793) An Historical Journal of the Transactions at Port Jackson and Norfolk Island, with the Discoveries which have been made in New South Wales and in the Southern Ocean. J. Stockdale, London, 525 pp.

    International Commission of Zoological Nomenclature (ICZN) (1957) Opinion 451. Use of the plenary powers to secure that the specific name dingo Meyer, 1793, as published in the combination Canis dingo shall be the oldest available name for the dingo of Australia (Class Mammalia). Opinions and Declarations Rendered by the International Commission of Zoological Nomenclature, 15, 329–338.

    International Commission of Zoological Nomenclature (ICZN) (2003) Opinion 2027 (Case 3010). Usage of 17 specific names based on wild species which are predated by or contemporary with those based on domestic animals (Lepidoptera, Osteichthyes, Mammalia). Bulletin of Zoological Nomenclature, 60, 81–84.

    Iredale, T. (1947) The scientific name of the dingo. Proceedings of the Royal Zoological Society of New South Wales, 67 (1946–1947), 35–36.

    Iredale, T. & Troughton, E. le G. (1934) A check-list of the mammals recorded from Australia. Memoirs of the Australian Museum, 6, 1–122.

    https://doi.org/10.3853/j.0067-1967.6.1934.516

    Jackson, S.M. (2003) Dingoes. In: Jackson, S.M. (Ed.), Australian Mammals: Biology and Captive Management. CSIRO Publishing, Melbourne, pp. 381–407.

    Jackson, S.M. & Groves, C.P. (2015) Taxonomy of Australian Mammals. CSIRO Publishing, Melbourne, 529 pp.

    Johnson, C.N., Isaac, J.L. & Fisher, D.O. (2007) Rarity of a top predator triggers continent-wide collapse of mammal prey: dingoes and marsupials in Australia. Proceedings of the Royal Society, Series B Biological Sciences, 274 (1608), 341–346.
    https://doi.org/10.1098/rspb.2006.3711

    Johnson, I. (1979) The getting of data. A case study from the recent industries of Australia. PhD Thesis, Australian National University, Canberra, 186 pp.

    Jones, E. & Stevens, P.L. (1988) Reproduction in wild canids, Canis familiaris, from the eastern highlands of Victoria. Wildlife Research, 15 (4), 385–397.

    https://doi.org/10.1071/WR9880385

    Jones, E. (1990) Physical characteristics and taxonomic status of wild canids Canis familiaris, from the eastern highlands of Victoria. Australian Wildlife Research, 17 (1), 69–81.

    https://doi.org/10.1071/wr9900069

    Jones, E. (2009) Hybridisation between the dingo, Canis lupus dingo, and the Domestic Dog, Canis lupus familiaris, in Victoria: a critical review. Australian Mammalogy, 31 (1), 1–7.

    https://doi.org/10.1071/AM08102

    Kadletz, M. (1932) Anatomischer Atlas der Extremitätengelenke von Pferd und Hund. Urban & Schwarzenberg, Berlin, 101 pp.

    Kerr, R. (1792) The Animal Kingdom, or zoological system, of the celebrated Sir Charles Linnaeus; Class1. Mammalia: containing a complete systematic description, arrangement, and nomenclature, of all the known species and varieties of the Mammalia, or animals which give suck to their young; being a translation of that part of the Systema Naturae, as lately published, with great improvements, by Professor Gmelin of Goettingen. Together with numerous additions from more recent zoological writers, and illustrated with copper plates. Vol. 1. J. Murray & R. Faulder, London, 644 pp.

    Kleiman, D.G. (1968) Reproduction in the Canidae. International Zoo Yearbook, 8, 3–8.

    https://doi.org/10.1111/j.1748-1090.1968.tb00419.x

    Koler-Matznick, J., Brisbin, I.L. Jr & McIntyre, J. (2000) The New Guinea singing dog: a living primitive dog. In: Crockford, S.J. (Ed.), Dogs through time: an archaeological perspective. Proceedings of the 1st ICAZ Symposium on the history of the Domestic Dog. British Archaeological Press, Oxford, pp. 239–247.

    Koler-Matznick, J., Brisbin, I.L., Feinstein, M. & Bulmer, S. (2003) An updated description of the New Guinea singing dog (Canis hallstromi, Troughton 1957). Journal of Zoology, 261 (2), 109–118.

    https://doi.org/10.1017/S0952836903004060

    Koler-Matznick, J., Yates, B.C., Bulmer, S. & Brisbin, I.L. (2007) The New Guinea singing dog: its status and scientific importance. Australian Mammalogy, 29 (1), 47–56.

    https://doi.org/10.1071/AM07005

    Koop, B.F., Burbidge, M., Byun, A., Rink, U. & Crockford, S.J. (2000) Ancient DNA evidence of a separate origin for North American indigenous dogs. In: Crockford, S.J. (Ed.), Dogs Through Time: An Archaeological Perspective. British Archaeological Reports, Oxford, pp. 271–285.

    Krefft, G. (1866) On the vertebrated animals of the lower Murray and Darling: their habits, economy, and geographical distribution. Transactions of the Philosophical Society of New South Wales, 1862–1865, 1–33.

    Krefft, G. (1868) Notes on the fauna of Tasmania. Papers and Proceedings of the Royal Society of Tasmania, 1868, 93–105.

    Lampert, R.J. (1971) Burrill Lake and Currarong. Terra Australia 1. Australian National University Press, Canberra, 86 pp.

    Larson, G., Karlsson, E.K., Perri, A., Webster, M.T., Ho, S.Y., Peters, J., Stahl, P.W., Piper, P.J., Lingaas, F., Fredholm, M., Comstock, K.E., Modiano, J.F., Schelling, C., Agoulnik, A.I., Leegwater, P.A., Dobney, K., Vigne, J.D., Vilà, C., Andersson, L. & Lindblad-Toh, K. (2012) Rethinking dog domestication by integrating genetics, archeology, and biogeography. Proceedings of the National Academy of Science, 109, 8878–8883.

    https://doi.org/10.1073/pnas.1203005109

    Leonard, J.A., Wayne, R.K., Wheeler, J., Valadez, R., Guillén, S. & Vilà, C. (2002) Ancient DNA evidence for old world origin of new world dogs. Science, 298 (5598), 1613–1616.

    https://doi.org/10.1126/science.1076980

    Leonard, J.A., Vilà, C., Fox-Dobbs, K., Koch, P.L., Wayne, R.K. & Van Valkenburgh, B. (2007) Megafaunal extinctions and the disappearance of a specialised wolf ectomorph. Current Biology, 17 (13), 1146–1150.

    https://doi.org/10.1016/j.cub.2007.05.072

    Lesèble, L. (1890) Les dingos au chenil du Jardin Zoologique d’Acclimatisation. Revue des Sciences Naturelles Appliquées: Bulletin Bimensuel de la Société National d’Acclimatisation de France, 37, 681–684.

    Letnic, M., Koch, F., Gordon, C., Crowther, M.S. & Dickman, C.R. (2009) Keystone effects of an alien top-predator stem extinctions of native mammals. Proceedings of the Royal Society of London, Series B Biological Sciences, 276 (1671), 3249–3256.

    https://doi.org/10.1098/rspb.2009.0574

    Letnic, M., Ritchie, E.G. & Dickman, C.R. (2012) Top predators as biodiversity regulators: the dingo Canis lupus dingo as a case study. Biological Reviews, 87 (2), 390–413.

    https://doi.org/10.1111/j.1469-185X.2011.00203.x

    Lindblad-Toh, K., Wade, C.M., Mikkelsen, T.S., Karlsson, E.K., Jaffe, D.B., Kamal, M., Clamp, M., Chang, J.L., Kulbokas, E.J., Zody, M.C., Mauceli, E., Xie, X., Breen, M., Wayne, R.K., Ostrander, E.A., Ponting, C.P., Galibert, F., Smith, D.R., deJong, P.J., Kirkness, E., Alvarez, P., Biagi, T., Brockman, W., Butler, J., Chin, C-W., Cook, A., Cuff, J., Daly, M.J., DeCaprio, D., Gnerre, S., Grabherr, M., Kellis, M., Kleber, M., Bardeleben, C., Goodstadt, L., Heger, A., Hitte, C., Kim, L., Koepfli, K.-P., Parker, H.G., Pollinger, J.P., Searle, S.M.J., Sutter, N.B., Thomas, R., Webber, C. & Lander, E.S. (2005) Genome sequence, comparative analysis and haplotype structure of the Domestic Dog. Nature, 438 (7069), 803–819.
    https://doi.org/10.1038/nature04338

    Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Vol. 1. Laurentii Salvii, Holmiae, 823 pp.

    Lorenzini, R. & Fico, R. (1995) A genetic investigation of enzyme polymorphisms shared by wolf and dog: suggestions for conservation of the wolf in Italy. Acta Theriologica, 40 (Supplement 3), 101–110.

    Lord, K., Feinstein, M., Smith, B. & Coppinger, C. (2013) Variation in reproductive traits of members of the genus Canis with special attention to the Domestic Dog (Canis familiaris). Behavioural Processes, 92, 131–142.

    https://doi.org/10.1016/j.beproc.2012.10.009

    Lowry, J.W.J. & Merrilees, D. (1969) Age of the desiccated carcase of a thylacine (Marsupialia, Dasyuridae) from Thylacine Hole, Nullarbor Region, Western Australia. Helictite, 7 (1), 15–16.

    Lucas, A.H.S. & Le Souëf, W.H.D. (1909) The Animals of Australia - Mammals, Reptiles, and Amphibians. Whitcombe & Tombs, Melbourne, 327 pp.

    https://doi.org/10.5962/bhl.title.69495

    Macintosh, N.W.G. (1964) A 3,000 year old dingo from shelter 6 (Fromm’s Landing, South Australia). Proceedings of the Royal Society of Victoria, 77, 498–507.

    Macintosh, N.W.G. (1975) The origin of the dingo: an enigma. In: Fox, M.W. (Ed.), The Wild Canids: Their Systematics, Behavioural Ecology and Evolution. Van Nostrand Reinhold, New York, pp. 87–106.

    Mahoney, J.A. & Richardson, B.J. (1988) Canidae. In: Walton, D.W. (Ed.), Zoological Catalogue of Australia. 5. Mammalia. Australian Government Publishing Service, Canberra, pp. 217–220.

    Major, R. (2009) Predation and hybridisation by feral dogs (Canis lupus familiaris) - Key threatening process listing. New South Wales Department of Environment, Climate Change, and Water, Sydney. Available from: http://www.environment.nsw.gov.au/determinations/feraldogsFD.htm (accessed 17 July 2017)

    Manor, R. & Saltz, D. (2004) The impact of free-roaming dogs on gazelle kid/female ratio in a fragmented area. Biological Conservation, 119 (2), 231–236.

    https://doi.org/10.1016/j.biocon.2003.11.005

    Mayden, R. (1997) A hierarchy of species concepts: the denouement in the saga of the species problem. In: Oaridge, M.F., Dawah, H.A. & Wilson, M.R. (Eds.), Species: The Units of Biodiversity. Chapman & Hall, London, pp. 381–424.

    McCoy, F. (1862) Note on the ancient and recent natural history of Victoria. Annals and Magazine of Natural History, Series 3, 9, 137–150.

    McCoy, F. (1882) Prodromus of the Palaeontology of Victoria; or Figures and Descriptions of Victorian organic remains. Decade VII. Geological Survey of Victoria. J. Ferres, Melbourne, 30 pp.

    McGreevy, P.D., Georgevsky, D., Carrasco, J., Valenzuela, M., Duffy, D.L. & Serpell, J.A. (2013) Dog behavior co-varies with height, bodyweight and skull shape. PLoS ONE, 8 (12), e80529.

    https://doi.org/10.1371/journal.pone.0080529

    Mech, L.D. (1970) The Wolf: The Ecology and Behavior of an Endangered Species. Natural History Press, New York, 384 pp.

    Meyer, F.A.A. (1793) Systematisch-summarische Uebersicht der neuesten zoologischen Entdeckungen in Neuholland und Afrika: nebst zwey andern zoologischen Abhandlungen. Dykische Buchhandlung, Leipzig, 178 pp.
    https://doi.org/10.5962/bhl.title.39685

    Milham, P. & Thompson, P. (1976) Relative antiquity of human occupation and extinct fauna at Madura Cave, southeastern Western Australia. Mankind, 10 (3), 175–180.

    https://doi.org/10.1111/j.1835-9310.1976.tb01149.x

    Morrison-Scott, T.C.S. (1955) Support for Tate’s proposed use of the plenary powers to validate the specific name “dingo” Meyer, 1793, as published in the combination “Canis dingo” as the name for the dingo (Class Mammalia). The Bulletin of Zoological Nomenclature, 11, 168.

    Mulvaney, D.J. (1975) The Prehistory of Australia. Penguin Books, Melbourne, 327 pp.

    Newsome, A.E., Corbett, K.L., Best, L.W. & Green, B. (1973) The dingo. Australian Meat Research Committee Review, 14, 1–11.

    Newsome, A.E., Corbett, L.K. & Carpenter, S.M. (1980) The identity of the dingo. I. Morphological discriminants of dingo and dog skulls. Australian Journal of Zoology, 28 (4), 615–625.

    https://doi.org/10.1071/ZO9800615

    Newsome, A.E. & Corbett, L.K. (1982) The identity of the dingo. II. Hybridisation with Domestic Dogs in captivity and in the wild. Australian Journal of Zoology, 30 (2), 365–374.

    https://doi.org/10.1071/ZO9820365

    Newsome, A.E. & Corbett, L.K. (1985) The identity of the dingo. III. The incidence of dingoes, dogs and hybrids and their coat colours in remote and settled regions of Australia. Australian Journal of Zoology, 33 (3), 363–375.

    https://doi.org/10.1071/ZO9850363

    Newsome, T.M., Stephens, D., Ballard, G.-A., Dickman, C.R. & Fleming, P.J.S. (2013a) Genetic profile of dingoes (Canis lupus dingo) and free-roaming Domestic Dogs (C. l. familiaris) in the Tanami Desert, Australia. Wildlife Research, 40 (3), 196–206.

    https://doi.org/10.1071/WR12128

    Newsome, T.M., Ballard, G.-A., Dickman, C.R., Fleming, P.J.S. & Howden, C. (2013b) Anthropogenic resource subsidies determine space use by Australian arid zone Dingoes: An improved resource selection modelling approach. PLoS ONE, 8, e63931.

    https://doi.org/10.1371/journal.pone.0063931

    Nixon, K.C. & Wheeler, Q.C. (1990) Amplification of the phylogenetic species concept. Cladistics, 6 (3), 211–223.

    https://doi.org/10.1111/j.1096-0031.1990.tb00541.x

    Nosil, P., Crespi, B.J. & Sandoval, C.P. (2003) Reproductive isolation driven by the combined effects of ecological adaptation and reinforcement. Proceedings of the Royal Society of London, Series B Biological Sciences, 270 (1527), 1911–1918. https://doi.org/10.1098/rspb.2003.2457

    Olsen, S.J. & Olsen, J.W. (1977) The Chinese wolf, ancestor of New World dogs. Science, 197 (4303), 533–535.

    https://doi.org/10.1126/science.197.4303.533

    Oppenheimer, E.C. & Oppenheimer, J.R. (1975) Certain behavioural features in the pariah dog (Canis familiaris) in West Bengal. Applied Animal Ethology, 2 (1), 81–92.

    https://doi.org/10.1016/0304-3762(75)90067-X

    Oskarsson, M.C.R., Klütsch, C.F.C., Boonyaprakob, U., Wilton, A., Tanabe, Y. & Savolainen, P. (2012) Mitochondrial DNA data indicate an introduction through mainland Southeast Asia for Australian dingoes and Polynesian Domestic Dogs. Proceedings of the Royal Society of London, Series B Biological Sciences, 279 (1730), 967–974.

    https://doi.org/10.1098/rspb.2011.1395

    Ovodov, N.D., Crockford, S.J., Kuzmin, Y.V., Higham, T.F.G., Hodgins, G.W.L. & van der Plicht, J. (2011) A 33,000-year-old incipient dog from the Altai Mountains of Siberia: Evidence of the earliest domestication disrupted by the Last Glacial Maximum. PLoS ONE, 6 (7), e22821.

    https://doi.org/10.1371/journal.pone.0022821

    Pal, S.K. (2001) Population ecology of free-ranging urban dogs in West Bengal, India. Acta Theriologica, 46 (1), 69–78.

    https://doi.org/10.1007/BF03192418

    Pal, S.K. (2008) Maturation and development of social behaviour during early ontogeny in free-ranging dog puppies in West Bengal, India. Applied Animal Behaviour Science, 111 (1–2), 95–107.

    https://doi.org/10.1016/j.applanim.2007.05.016

    Pang, J.-F., Kluetsch, C., Zou, X.-J., Zhang, A.-B., Luo, L.Y., Angleby, H., Ardalan, A., Ekström, C., Sköllermo, A., Lundeberg, J., Matsumura, S., Leitner, T., Zhang, Y.-P. & Savolainen, P. (2009) mtDNA data indicate a single origin for dogs South of Yangtze River, less than 16,300 years ago, from numerous wolves. Molecular Biology and Evolution, 26 (12), 2849–2864.

    https://doi.org/10.1093/molbev/msp195

    Parr, W.C.H., Wilson, L.A.B., Wroe, S., Colman, N.J., Crowther, M.S. & Letnic, M. (2016) Cranial shape and the modularity of hybridization in dingoes and dogs; hybridization does not spell the end for native morphology. Evolutionary Biology, 43, 171–187.
    https://doi.org/10.1007/s11692-016-9371-x

    Phillip, A. (1789) The Voyage of Governor Phillip to Botany Bay, with an account of the establishment of the colonies of Port Jackson & Norfolk Island compiled from authentic papers, which have been obtained from the several departments, to which are added the journals of Lieuts. Shortland, Watts, Ball & Capt. Marshall; with an account of their discoveries, embellished with LV copper plates, the maps and charts taken from actual surveys, & the plans and views drawn on the spot, by Capt. Hunter, Lieuts. Shortland, Watts, Dawes, Bradley, Capt. Marshall. John Stockdale, London, 298 pp.

    Pilot, M., Branicki, W., Jędrzejewski, W., Goszczyński, J., Jędrzejewska, B., Dykyy, I., Shkvyrya, M. & Tsingarska, E. (2010) Phylogenetic history of grey wolves in Europe. BMC Evolutionary Biology, 10 (104), 1–11.

    https://doi.org/10.1186/1471-2148-10-104

    Purcell, B. (2010) Dingo. CSIRO Publishing, Melbourne, 166 pp.

    Radford, C.G., Letnic, M., Fillios, M. & Crowther, M.S. (2012) An assessment of the taxonomic status of wild canids in south-eastern New South Wales: phenotypic variation in dingoes. Australian Journal of Zoology, 60 (2), 73–80.

    https://doi.org/10.1071/ZO12006

    Ritchie, E.G. & Johnson, C.N. (2009) Predator interactions, mesopredator release and biodiversity conservation. Ecology Letters, 12 (9), 982–998.

    https://doi.org/10.1111/j.1461-0248.2009.01347.x

    Sacks, B.N., Bannasch, D.L., Chomel, B.B. & Ernest, H.B. (2008) Coyotes demonstrate how habitat specialization by individuals of a generalist species can diversify populations in a heterogeneous ecoregion. Molecular Biology and Evolution, 25 (7), 1384–1394.

    https://doi.org/10.1093/molbev/msn082

    Sacks, B.N., Brown, S.K., Stephens, D., Pedersen, N.C., Wu, J.-T. & Berry, O. (2013) Y chromosome analysis of dingoes and Southeast Asian village dogs suggests a Neolithic continental expansion from Southeast Asia followed by multiple Austronesian dispersals. Molecular Biology and Evolution, 30 (5), 1103–1118.

    https://doi.org/10.1093/molbev/mst027

    Savolainen, P., Zhang, Y., Luo, J., Lundeberg, J. & Leitner, T. (2002) Genetic evidence for an east Asian origin of Domestic Dogs. Science, 298 (5598), 1610–1613.

    https://doi.org/10.1126/science.1073906

    Savolainen, P., Leitner, T., Wilton, A.N., Matisoo-Smith, E. & Lundeberg, J. (2004) A detailed picture of the origin of the Australian dingo, obtained from the study of mitochondrial DNA. Proceedings of the National Academy of Sciences, USA, 101 (33), 12387–12390.

    https://doi.org/10.1073/pnas.0401814101

    Scott, J.P. & Fuller, J.L. (1965) Genetics and Social Behavior of the Dog. University of Chicago Press, Chicago, 468 pp.

    Sheldon, J. (1992) Wild Dogs: The Natural History of the Nondomestic Canidae. Academic Press, Inc, San Diego, 248 pp.

    Sillero-Zubiri, C. (2009) Family Canidae (Dogs). In: Wilson, D.E. & Mittermeier, R.A. (Eds.), Handbook of the Mammals of the World. Vol. 1. Carnivores. Lynx Editions, Barcelona, pp. 352–466.

    Skoglund, P., Ersmark, E., Palkopoulou, E. & Dalén, L. (2015) Ancient wolf genome reveals an early divergence of Domestic Dog ancestors and admixture into high-latitude breeds. Current Biology, 25 (11), 1515–1519.

    https://doi.org/10.1016/j.cub.2015.04.019

    Smith, B. (2015) The role of dingoes in Indigenous Australian lifestyle, culture, and spirituality. In: Smith, B. (Ed.), The dingo debate: origins, behaviour and conservation CSIRO Publishing, Melbourne, pp. 81–101.

    Smith, M.A. (1980) Saltbush, Sampling Strategy and Settlement Pattern. MA Thesis, Australian National University, Canberra, 295 pp. [unpublished]

    Stephens, D., Wilton, A.N., Fleming, P.J.S. & Berry, O. (2015) Death by sex in an Australian icon: a continent-wide survey reveals extensive hybridisation between dingoes and Domestic Dogs. Molecular Ecology, 24 (22), 5643–5656.
    https://doi.org/10.1111/mec.13416

    Strahan, R. (Ed.) (1983) The Complete Book of Australian Mammals. Angus and Robertson, Sydney, 530 pp.

    Strahan, R. (1992) Encyclopedia of Australian Animals: Mammals. Angus & Robertson, Sydney, 184 pp.

    Strahan, R. (Ed.) (1995) Mammals of Australia. Reed Books, Sydney, 756 pp.

    Suzuki, H., Nunome, M., Kinoshita, G., Aplin, K.P., Vogel, P., Kryukov, A.P., Lin, M.-L., Han, S.-H., Maryanto, I., Tsuchiya, K., Ikeda, H., Shiroishi, T., Yonekawa, H. & Moriwaki, K. (2013) Evolutionary and dispersal history of Eurasian house mice Mus musculus clarified by more extensive geographic sampling of mitochondrial DNA. Heredity, 111 (5), 375–390.

    https://doi.org/10.1038/hdy.2013.60

    Tao, Y., Chen, S., Hartl, D.L. & Laurie, C.C. (2003) Genetic dissection of hybrid incompatibilities between Drosophila simulans and D. mauritiana. I. Differential accumulation of hybrid male sterility effects on the X and autosomes. Genetics, 164 (4), 1383–1398.

    Tate, G.H.H. (1955) Proposed use of the plenary powers to validate the specific name “dingo” Meyer, 1793, as published in the combination “Canis dingo” as the name for the dingo (Class Mammalia). The Bulletin of Zoological Nomenclature, 11, 121.

    Teeter, K.C., Payseur, B.A., Harris, L.W., Bakewell, M.A., Thibodeau, L.M., O’Brien, J.E., Krenz, J.G., Sans-Fuentes, M.A., Nachman, M.W & Tucker, P.K. (2008). Genome-wide patterns of gene flow across a house mouse hybrid zone. Genome Research, 18 (1), 67–76.

    https://doi.org/10.1101/gr.6757907

    Temminck, C.J. (1838) Over de kennis en de verbreiing der zoogdiern van Japan. Tijdschrift voor Natuurlijke Geschiedenis en Physiologie, 5, 273–293.

    Tench, W. (1789) A narrative of the expedition to Botany Bay. J. Debrett, London, 64 pp. Avaliable from: http://setis.library.usyd.edu.au/ozlit/pdf/p00039.pdf (Accessed 1 Sept. 2017)

    Thalmann, O., Shapiro, B., Cui, P., Schuenemann, V.J., Sawyer, S.K., Greenfield, D.L., Germonpré, M.B., Sablin, M.V., López-Giráldez, F., Domingo-Roura, X., Napierala, H., Uerpmann, H.P., Loponte, D.M., Acosta, A.A., Giemsch, L., Schmitz, R.W., Worthington, B., Buikstra, J.E., Druzhkova, A., Graphodatsky, A.S., Ovodov, N.D., Wahlberg, N., Freedman, A.H., Schweizer, R.M., Koepfli, K.P., Leonard, J.A., Meyer, M., Krause, J., Pääbo, S., Green, R.E. & Wayne, R.K. (2013) Complete mitochondrial genomes of ancient canids suggest a European origin of Domestic Dogs. Science, 342 (6160), 871–874.

    https://doi.org/10.1126/science.1243650

    The Bovine Genome Sequencing Analysis Consortium, Elsik, C.G., Tellam, R.L. & Worley, K.C. (2009) The Genome Sequence of Taurine Cattle: A Window to Ruminant Biology and Evolution. Science, 324 (5926), 522–528.

    https://doi.org/10.1126/science.1169588

    Thompson, P.C. (1992a) The behavioural ecology of dingoes in north-western Australia. IV. Social and spatial organisation, and movements. Wildlife Research, 19 (5), 543–564.

    https://doi.org/10.1071/WR9920543

    Thompson, P.C. (1992b) The behavioural ecology of dingoes in North-Western Australia. II. Activity patterns, breeding season, and pup rearing. Wildlife Research, 19, 519–530.
    https://doi.org/10.1071/WR9920519

    Troughton, E. le G. (1957) A new native dog from the Papuan highlands. Proceedings of the Royal Zoological Society of New South Wales, 76 (1955–1956), 93–94.

    Troughton, E. le G. (1967) Furred Animals of Australia. 9th Edition. Angus and Robertson, Sydney, 314 pp.

    True, J.R., Weir, B.S. & Laurie, C.C. (1996) A genome-wide survey of hybrid incompatibility factors by the introgression of marked segments of Drosophila mauritiana chromosomes into Drosophila simulans. Genetics, 142 (3), 819–837.

    Turnbull, P.F. & Reed, C.A. (1974) The fauna of the terminal Pleistocene of Palegawra Cave, a Zarazian occupational site in northeastern Iraq. Fieldiana Anthropology, 63, 81–146.

    Uerpmann, H.-P. (1982) Faunal remains from Shams ed-Din Tannira, a Halafian Site in Northern Syria. Berytus Archaeological Studies (supplement), 30, 3–52.

    Vaysse, A., Ratnakumar, A., Derrien, T., Axelsson, E., Rosengren Pielberg, G., Sigurdsson, S., Fall, T., Seppälä, E.H., Hansen, M.S., Lawley, C.T., Karlsson, E.K., The LUPA Consortium, Bannasch, D., Vilà, C., Lohi, H., Galibert, F., Fredholm, M., Häggström, J., Hedhammar, A., André, C., Lindblad-Toh, K., Hitte, C. & Webster, M.T. (2011) Identification of genomic regions associated with phenotypic variation between dog breeds using selection mapping. PLoS Genetics, 7 (10), e1002316.
    https://doi.org/10.1371/journal.pgen.1002316

    Van Dyck, S. & Strahan, R. (2008) The Mammals of Australia. 3rd Edition. Reed New Holland, Sydney, 887 pp.

    Venter, J.C., Adams, M.D., Myers, E.W., Li, P.W., Mural, R.J., Sutton, G.G., Smith, H.O., Yandell, M., Evans, C.A., Holt, R.A., Gocayne, J.D., Amanatides, P., Ballew, R.M., Huson, D.H., Wortman, J.R., Zhang, Q., Kodira, C.D., Zheng, X.H., Chen, L., Skupski, M., Subramanian, G., Thomas, P.D., Zhang, J., Gabor Miklos, G.L., Nelson, C., Broder, S., Clark, A.G., Nadeau, J., McKusick, V.A., Zinder, N., Levine, A.J., Roberts, R.J., Simon, M., Slayman, C., Hunkapiller, M., Bolanos, R., Delcher, A., Dew, I., Fasulo, D., Flanigan, M., Florea, L., Halpern, A., Hannenhalli, S., Kravitz, S., Levy, S., Mobarry, C., Reinert, K., Remington, K., Abu-Threideh, J., Beasley, E., Biddick, K., Bonazzi, V., Brandon, R., Cargill, M., Chandramouliswaran, I., Charlab, R., Chaturvedi, K., Deng, Z., Di Francesco, V., Dunn, P., Eilbeck, K., Evangelista, C., Gabrielian, A.E., Gan, W., Ge, W., Gong, F., Gu, Z., Guan, P., Heiman, T.J., Higgins, M.E., Ji, R.R., Ke, Z., Ketchum, K.A., Lai, Z., Lei, Y., Li, Z., Li, J., Liang, Y., Lin, X., Lu, F., Merkulov, G.V., Milshina, N., Moore, H.M., Naik, A.K., Narayan, V.A., Neelam, B., Nusskern, D., Rusch, D.B., Salzberg, S., Shao, W., Shue, B., Sun, J., Wang, Z., Wang, A., Wang, X., Wang, J., Wei, M., Wides, R., Xiao, C., Yan, C., Yao, A., Ye, J., Zhan, M., Zhang, W., Zhang, H., Zhao, Q., Zheng, L., Zhong, F., Zhong, W., Zhu, S., Zhao, S., Gilbert, D., Baumhueter, S., Spier, G., Carter, C., Cravchik, A., Woodage, T., Ali, F., An, H., Awe, A., Baldwin, D., Baden, H., Barnstead, M., Barrow, I., Beeson, K., Busam, D., Carver, A., Center, A., Cheng, M.L., Curry, L., Danaher, S., Davenport, L., Desilets, R., Dietz, S., Dodson, K., Doup, L., Ferriera, S., Garg, N., Gluecksmann, A., Hart, B., Haynes, J., Haynes, C., Heiner, C., Hladun, S., Hostin, D., Houck, J., Howland, T., Ibegwam, C., Johnson, J., Kalush, F., Kline, L., Koduru, S., Love, A., Mann, F., May, D., McCawley, S., McIntosh, T., McMullen, I., Moy, M., Moy, L., Murphy, B., Nelson, K., Pfannkoch, C., Pratts, E., Puri, V., Qureshi, H., Reardon, M., Rodriguez, R., Rogers, Y.H., Romblad, D., Ruhfel, B., Scott, R., Sitter, C., Smallwood, M., Stewart, E., Strong, R., Suh, E., Thomas, R., Tint, N.N., Tse, S., Vech, C., Wang, G., Wetter, J., Williams, S., Williams, M., Windsor, S., Winn-Deen, E., Wolfe, K., Zaveri, J., Zaveri, K., Abril, J.F., Guigo, R., Campbell, M.J., Sjolander, K.V., Karlak, B., Kejariwal, A., Mi, H., Lazareva, B., Hatton, T., Narechania, A., Diemer, K., Muruganujan, A., Guo, N., Sato, S., Bafna, V., Istrail, S., Lippert, R., Schwartz, R., Walenz, B., Yooseph, S., Allen, D., Basu, A., Baxendale, J., Blick, L., Caminha, M., Carnes-Stine, J., Caulk, P., Chiang, Y.H., Coyne, M., Dahlke, C., Mays, A., Dombroski, M., Donnelly, M., Ely, D., Esparham, S., Fosler, C., Gire, H., Glanowski, S., Glasser, K., Glodek, A., Gorokhov, M., Graham, K., Gropman, B., Harris, M., Heil, J., Henderson, S., Hoover, J., Jennings, D., Jordan, C., Jordan, J., Kasha, J., Kagan, L., Kraft, C., Levitsky, A., Lewis, M., Liu, X., Lopez, J., Ma, D., Majoros, W., McDaniel, J., Murphy, S., Newman, M., Nguyen, T., Nguyen, N., Nodell, M., Pan, S., Peck, J., Peterson, M., Rowe, W., Sanders, R., Scott, J., Simpson, M., Smith, T., Sprague, A., Stockwell, T., Turner, R., Venter, E., Wang, M., Wen, M., Wu, D., Wu, M., Xia, A., Zandieh, A. & Zhu, X. (2001) The Sequence of the Human Genome. Science, 291 (5507), 1304–1351.

    https://doi.org/10.1126/science.1058040

    Vernes, K., Dennis, A. & Winter, J. (2001) Mammalian diet and broad hunting strategy of the dingo (Canis familiaris dingo) in the wet tropical rainforests of northeastern Australia. Biotropica, 33 (2), 339–345.

    https://doi.org/10.1646/0006-3606(2001)033[0339:mdabhs]2.0.co;2

    Vilà, C.P., Savolainen, P., Maldonado, J.E., Amorim, I.R., Rice, J.E., Honeycutt, R.L., Crandall, K.A., Lundenberg, J. & Wayne, R.K. (1997) Multiple and ancient origins of the Domestic Dog. Science, 276 (5319), 1687–1689.

    https://doi.org/10.1126/science.276.5319.1687

    Vilà, C., Amorim, I.R., Leonard, J.A., Posada, D., Castroviejo, J., Petrucci-Fonseca, F., Crandall, K.A., Ellegren, H. & Wayne, R.K. (1999) Mitochondrial DNA phylogeography and population history of the grey wolf Canis lupus. Molecular Ecology, 8 (12), 2089–2103.

    https://doi.org/10.1046/j.1365-294x.1999.00825.x

    Voigt, F.S. (1831) G. Cuvier, Das Thierreich, geordnet nach seiner organisation : Als grundlage der naturgeschichte der thiere und einleitung in die vergleichende anatomie. Vol. 1. Brockhaus, Leipzig, 974 pp.

    vonHoldt, B.M., Polliger, J.P., Lohmueller, K.E., Han, E., Parker, H.G., Quignon, P., Degenhardt, J.D., Boyko, A.R., Earl, D.A., Auton, A., Reynolds, A., Bryc, K., Brisbin, A., Knowles, J.C., Mosher, D.S., Spady, T.C., Elkahloun, A., Geffen, E., Pilot, M., Jedrzejewski, W., Greco, C., Randi, E., Bannasch, D., Wilton, A., Shearman, J., Musiani, M., Cargill, M., Jones, P.G., Qian, Z., Huang, W., Ding, Z.-L., Zhang, Y.-P., Bustamante, C.D., Ostrander, E.A., Novembre, J. & Wayne, R.K. (2010) Genome-wide SNP and haplotype analyses reveal a rich history underlying dog domestication. Nature, 464 (7290), 898–903.

    https://doi.org/10.1038/nature08837

    vonHoldt, B.M., Pollinger, J.P., Earl, D.A., Knowles, J.C., Boyko, A.R., Parker, H., Geffen, E., Pilot, M., Jedrzejewski, W., Jedrzejewska, B., Sidorovich, V., Greco, C., Randi, E., Musiani, M., Kays, R., Bustamante, C.D., Ostrander, E.A., Novembre, J. & Wayne, R.K. (2011) A genome-wide perspective on the evolutionary history of enigmatic wolf-like canids. Genome Research, 21 (8), 1294–1305.

    https://doi.org/10.1101/gr.116301.110

    Wakefield, N.A. (1966) Mammals of the Blandowski Expedition to north western Victoria, 1856-57. Proceedings of the Royal Society of Victoria, 79 (2), 371–379.

    Walters, B. (1995) The Company of Dingoes: Two Decades with our Native Dog. Australian Native Dog Conservation Society, Bargo, NSW, 109 pp.

    Wang, G.-D., Zhai, W.W., Yang, H.-C., Fan, R.-X., Cao, X., Zhong, L., Wang, L., Liu, F., Wu, H., Cheng, L.G., Poyarkov, A.D., Poyarkov, N.A., Tang, S.S., Zhao, W.M., Gao, Y., Lv, X.M., Irwin, D.M., Savolainen, P., Wu, C.I. & Zhang, Y.P. (2013) The genomics of selection in dogs and the parallel evolution between dogs and humans. Nature Communications, 4 (1860), 1–9.

    https://doi.org/10.1038/ncomms2814

    Wayne, R.K. (1993) Molecular evolution of the dog family. Trends in Genetics, 9 (6), 218–224.

    https://doi.org/10.1016/0168-9525(93)90122-x

    Wayne, R.K. & O’Brien, S.J. (1987) Allozyme divergence within the Canidae. Systematic Zoology, 36 (4), 339–355.

    https://doi.org/10.2307/2413399

    Wayne, R.K. & Ostranger, E.A. (1999) Origin, genetic diversity, and genome structure of the Domestic Dog. Bioessays, 21 (3), 247–257.

    https://doi.org/10.1002/(sici)1521-1878(199903)21:3<247::aid-bies9>3.0.co;2-z

    Wilson, D.E. & Reeder, D.M. (1993) Mammal Species of the World - A Taxonomic and Geographic Reference. Smithsonian Institution Press, Washington D.C., 1206 pp.

    Wilton, A.N., Steward, D.J. & Zafiris, K. (1999) Microsatellite variation in the Australian dingo. Journal of Heredity, 90 (1), 108–111.

    https://doi.org/10.1093/jhered/90.1.108

    Wilton, A.N. (2001) DNA methods of assessing dingo purity. In: Dickman C.R. & Lunney, D. (Eds.), A Symposium on the Dingo. Royal Zoological Society of New South Wales, Sydney, pp. 49–56.

    https://doi.org/10.7882/fs.2001.008

    Wood Jones, F.W. (1921) The status of the dingo. Transactions of the Royal Society of South Australia, 45, 254–263.

    Wood-Jones, F. (1925) The Mammals of South Australia. Part III. The Monodelphia. R.E.E. Rogers Government Printers, Adelaide, 458 pp.

    Woodall, P.F., Pavlov, P. & Twyford, K.L. (1996) Dingoes in Queensland, Australia: skull dimensions and the identity of wild canids. Wildlife Research, 23 (5), 581–587.

    https://doi.org/10.1071/WR9960581

    Wozencraft, W.C. (2005) Order Carnivora. In: Wilson, D.E. & Reeder, D.A. (Eds.), Mammal Species of the World. A Taxonomic and Geographic Reference. 3rd Edition. Johns Hopkins University Press, Baltimore, pp. 532–628.

    Zachos, F.E. & Lovari, S. (2013) Taxonomic inflation and the poverty of the Phylogenetic Species Concept—a reply to Gippoliti and Groves. Hystrix. Italian Journal of Mammalogy, 24, 142–144.

    Zimen, E. (1975) Social dynamics of the wolf pack. In: Fox, M.W. (Ed.), The Wild Canids: Their Systematics, Behavioral Ecology and Evolution. Robert E. Krieger Publishing Co., Florida, pp. 336–362.