Skip to main content Skip to main navigation menu Skip to site footer
Responsive image
Issue: Vol. 4341 No. 1: 30 Oct. 2017
Type: Article
Published: 2017-10-30
DOI: 10.11646/zootaxa.4341.1.2
Page range: 25–40
Abstract views: 96
PDF downloaded: 2

A new species of Cyrtodactylus (Squamata: Gekkonidae) and the first record of C. otai from Son La Province, Vietnam

Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam. Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam
Faculty of Biology and Chemistry, Tay Bac University, Son La City, Son La Province, Vietnam.
AG Zoologischer Garten Köln, Riehler Strasse 173, D–50735 Cologne, Germany. Institute of Zoology, University of Cologne, Zülpicher Strasse 47b, D–50674 Cologne, Germany.
Faculty of Biology, Hanoi University of Science, Vietnam National University, 334 Nguyen Trai Road, Hanoi, Vietnam.
Faculty of Environmental Sciences, Hanoi University of Science, Vietnam National University, 334 Nguyen Trai Road, Hanoi, Vietnam. Central Institute for Natural Resources and Environmental Studies, Vietnam National University, 19 Le Thanh Tong, Hanoi, Vietnam Department of Herpetology, American Museum of Natural History, Central Park West at 79th Street, New York, New York 10024
Reptilia Cyrtodactylus sonlaensis sp. nov. C. otai molecular phylogeny new record taxonomy Phu Yen Van Ho

Abstract

We describe a new species of Cyrtodactylus on the basis of four specimens collected from the limestone karst forest of Phu Yen District, Son La Province, Vietnam. Cyrtodactylus sonlaensis sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: maximum SVL of 83.2 mm; dorsal tubercles in 13–15 irregular rows; ventral scales in 34–42 rows; ventrolateral folds prominent without interspersed tubercles; enlarged femoral scales 15–17 on each thigh; femoral pores 14–15 on each thigh in males, absent in females; precloacal pores 8, in a continuous row in males, absent in females; postcloacal tubercles 2 or 3; lamellae under toe IV 18–21; dorsal head with dark brown markings, in oval and arched shapes; nuchal loop discontinuous; dorsum with five brown bands between limb insertions, third and fourth bands discontinuous; subcaudal scales distinctly enlarged. In phylogenetic analyses, the new species is nested in a clade consisting of C. huongsonensis and C. soni from northern Vietnam and C. cf. pulchellus from Malaysia based on maximum likelihood and Bayesian analyses. In addition, we record Cyrtodactylus otai Nguyen, Le, Pham, Ngo, Hoang, Pham & Ziegler for the first time from Son La Province based on specimens collected from Van Ho District.

 

References

  1. Bauer, A.M. (2002) Two new species of Cyrtodactylus (Squamata: Gekkonidae) from Myanmar. Proceedings of the California Academy of Sciences, 53 (7), 75–88.

    Bauer, A.M. (2003) Descriptions of seven new Cyrtodactylus (Squamata: Gekkonidae) with a key to the species of Myanmar (Burma). Proceedings of the California Academy of Sciences, 54 (25), 463–498.

    Bauer, A.M., Kunya, K., Sumontha, M., Niyomwan, P., Panitvong, N., Pauwels, O.S.G., Chanhome, L. & Kunya, T. (2009) Cyrtodactylus erythrops (Squamata: Gekkonidae), a new cave-dwelling gecko from Mae Hong Son Province, Thailand. Zootaxa, 2124, 51–62.

    Bauer, A., Kunya, K., Sumontha, M., Niyomwan, P., Pauwels, O.S.G., Chanhome, L. & Kunya, T. (2010) Cyrtodactylus dumnuii (Squamata: Gekkonidae), a new cave–dwelling gecko from Chiang Mai Province, Thailand. Zootaxa, 2570, 41–50.

    Bauer, A.M., Pauwels, O.S.G. & Chanhome, L. (2002) A new species of cave-dwelling Cyrtodactylus (Squamata: Gekkonidae) from Thailand. The Natural History Journal of Chulalongkorn University, 2, 19–29.

    Bauer, A.M., Sumontha, M. & Pauwels, O.S.G. (2003) Two new species of Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from Thailand. Zootaxa, 376, 1–18.
    https://doi.org/10.11646/zootaxa.376.1.1

    Bergsten, J. (2005) A review of long-branch attraction. Cladistics, 21, 163–193.
    https://doi.org/10.1111/j.1096-0031.2005.00059.x

    Chan-ard, T. & Makchai, S. (2011) A new insular species of Cyrtodactylus Gray, 1827 (Squamata, Gekkonidae), from the Surin Islands, Phang-nga Province, southern Thailand. The Thailand Natural History Museum Journal, 5 (1), 7–15.

    Chung, K.-F., Leong, W.-C., Rubite, R.R., Repin, R., Kiew, R., Liu, Y. & Peng, C.-I. (2014) Phylogenetic analyses of Begonia sect. Coelocentrum and allied limestone species of China shed light on the evolution of Sino-Vietnamese karst flora. Botanical Studies, 55, 1

    Clements, R., Sodhi, N.S., Schilthuizen, M. & Ng, P.K.L. (2006) Limestone karsts of Southeast Asia: imperiled arks of biodiversity. BioScience, 56, 733–742.
    https://doi.org/10.1641/0006-3568(2006)56[733:LKOSAI]2.0.CO;2

    Connette, G.M., Oswald, P., Thura, M.K., Connette, K.J.L., Grindley, M.E., Songer, M., Zug, R.G. & Mulcahy, G.D. (2017) Rapid forest clearing in a Myanmar proposed national park threatens two newly discovered species of geckos (Gekkonidae: Cyrtodactylus). PLOS ONE, 12 (4), e0174432.
    https://doi.org/10.1371/journal.pone.0174432

    David, P., Nguyen, T.Q., Schneider, N. & Ziegler, T. (2011) A new species of the genus Cyrtodactylus Gray, 1827 from central Laos (Squamata: Gekkonidae). Zootaxa, 2833, 29–40.

    Grismer, L.L. & Ahmad, N. (2008) A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia. Zootaxa, 1924, 53–68.

    Grismer, L.L., Wood, P.L. Jr., Anuar, S., Davis, H.R., Burch, B.T., Cobos, A.J. & Murdoch, M.L. (2016) A new species of karst forest Bent-toed Gecko (genus Cyrtodactylus Gray) not yet threatened by foreign cement companies and a summary of Peninsular Malaysia’s endemic karst forest herpetofauna and the need for its conservation, Zootaxa, 4061 (1), 1–17.

    https://doi.org/10.11646/zootaxa.4061.1.1

    Grismer, L.L., Wood, P.L. Jr., Quah, E.S.H., Anuar, S., Muin, A., Sumontha, M., Ahmad, N., Bauer, A.M., Wangkulangkul, S., Grismer, J.L. & Pauwels, O.S.G. (2012) A phylogeny and taxonomy of the Thai-Malay Peninsula Bent-toed Geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species. Zootaxa, 3520, 1–55.

    Hoang, Q.X., Orlov, N.L., Ananjeva, N.B., Johns, A.G., Hoang, T.N. & Dau, V.Q. (2007) Description of a new species of the genus Cyrtodactylus Gray, 1827 (Squamata: Sauria: Gekkonidae) from the karst of North Central Vietnam. Russian Journal of Herpetology, 14, 98–106.

    Ivanova, N.V., de Waard, J. & Hebert, P.D.N. (2006) An inexpensive, automation-friendly protocol for recovering high-quality DNA. Molecular Ecology Notes, 6, 998–1002.
    https://doi.org/10.1111/j.1471-8286.2006.01428.x

    Kunya, K., Panmongkol, A., Pauwels, O.G.S., Sumontha, M., Meewasana, J., Bunkhwamdi, W. & Dangsri, S. (2014) A new forest-dwelling Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Doi Suthep, Chiang Mai Province, northern Thailand. Zootaxa, 3811 (2), 251–261.
    https://doi.org/10.11646/zootaxa.3811.2.6

    Kunya, K., Sumontha, M., Panitvong, N., Dongkumfu, W., Sirisamphan, T. & Pauwels, O.S.G. (2015) A new forest-dwelling Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Doi Inthanon, Chiang Mai Province, northern Thailand. Zootaxa, 3905 (4), 573–584.

    https://doi.org/10.11646/zootaxa.3905.4.9

    Le, D.T., Nguyen, T.Q., Le, M.D. & Ziegler, T. (2016) A new species of Cyrtodactylus (Squamata: Gekkonidae) from Ninh Binh Province, Vietnam. Zootaxa, 4162 (2), 268–282.

    https://doi.org/10.11646/ zootaxa.4162.2.4

    Le, T.D., Nguyen, T.T., Nishikawa, K., Nguyen, L.H.S., Pham, V.A., Matsui, M., Bernardes, M., & Nguyen, Q.T. (2015a) A New Species of Tylototriton Anderson, 1871 (Amphibia: Salamandridae) from Northern Indochina. Current Herpetology, 34, 38–50.

    Le, T.D., Pham, V.A., Nguyen, L.H.S., Ziegler, T. & Nguyen, Q.T. (2014) Babina lini (Chou, 1999) and Hylarana menglaensis Fei, Ye, and Xie, 2008, two additional anuran species for the herpetofauna of Vietnam. Russian Journal of Herpetology, 21, 315–321.

    Le, T.D., Pham, V.A., Nguyen, L.H.S., Ziegler, T. & Nguyen, Q.T. (2015b) First records of Megophrys daweimontis Rao and Yang, 1997 and Amolops vitreus (Bain, Stuart and Orlov, 2006) (Anura: Megophryidae, Ranidae) from Vietnam. Asian Herpetological Research, 6, 66–72.

    Le, T.D., Pham, V.A., Pham, T.C., Nguyen, L.H.S., Ziegler, T. & Nguyen, Q.T. (2015c) Review of the genus Sinonatrix in Vietnam with a new country record of Sinonatrix yunnanensis Rao and Yang, 1998. Russian Journal of Herpetology, 22, 84–88.

    Le, M., Raxworthy, C.J., McCord, W.P. & Mertz, L. (2006) A molecular phylogeny of tortoises (Testudines: Testudinidae) based on mitochondrial and nuclear genes. Molecular Phylogenetics and Evolution, 40, 517–531.
    https://doi.org/10.1016/j.ympev.2006.03.003

    Luu, V.Q., Bonkowski, M., Nguyen, T.Q., Le, M.D., Schneider, N., Ngo, H.T. & Ziegler, T. (2016a) Evolution in karst massifs: Cryptic diversity among bent-toed geckos along the Truong Son Range with descriptions of three new species and one new country record from Laos. Zootaxa, 4107 (2), 101–140.

    https://doi.org/10.11646/zootaxa.4107.2.1

    Luu, V.Q., Calame, T., Bonkowski, M., Nguyen, T.Q. & Ziegler, T. (2014) A new species of Cyrtodactylus (Squamata: Gekkonidae) from Khammouane Province, Laos. Zootaxa, 3760 (1), 54–66.

    https://doi.org/10.11646/zootaxa.3760.1.3

    Luu, V.Q., Calame, T., Nguyen, T.Q., Bonkowski, M. & Ziegler, T. (2015) A new species of Cyrtodactylus (Squamata: Gekkonidae) from the limestone forest of Khammouane Province, central Laos. Zootaxa, 4058 (3), 388–402.

    https://doi.org/10.11646/zootaxa.4058.3.6

    Luu, V.Q., Calame, T., Nguyen, T.Q., Le, M.D., Bonkowski, M. & Ziegler, T. (2016b) Cyrtodactylus rufford, a new cave-dwelling Bent-toed Gecko (Squamata: Gekkonidae) from Khammouane Province, central Laos. Zootaxa, 4067 (2), 185–199.

    https://doi.org/10.11646/zootaxa.4067.2.4

    Luu, V.Q., Nguyen, T.Q., Do, H.Q. & Ziegler, T. (2011) A new Cyrtodactylus (Squamata: Gekkonidae) from Huong Son limestone forest, Hanoi, northern Vietnam. Zootaxa, 3129, 39–50.

    Luu, V.Q., Nguyen, T.Q., Le, M.D., Bonkowski, M. & Ziegler, T. (2016c) A new species of karst-dwelling Bent-toed Gecko (Squamata: Gekkonidae) from Khammouane Province, central Laos. Zootaxa, 4079 (1), 87–102.

    https://doi.org/10.11646/zootaxa.4079.1.6

    Nazarov, R.A., Poyarkov, N.A., Orlov, N.L., Nguyen, S.N., Milto, K.D., Martynov, A.A., Konstantinov, E.L. & Chulisov, A.S. (2014) A review of genus Cyrtodactylus (Reptilia: Sauria: Gekkonidae) in fauna of Laos with description of four new species. Proceedings of the Zoological Institute RAS, 318, 391–423.

    Ngo, T.V. (2008) Two new cave-dwelling species of Cyrtodactylus Gray (Squamata: Gekkonidae) from Southwestern Vietnam. Zootaxa, 1909, 37–51.

    Ngo, T.V. & Bauer, A.M. (2008) Descriptions of two new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) endemic to southern Vietnam. Zootaxa, 715, 27–42.

    Ngo, T.V. & Chan, K.O. (2010) A new species of Cyrtodactylus Gray, 1826 (Squamata: Gekkonidae) from Khanh Hoa province, southern Vietnam. Zootaxa, 2504, 47–60.

    Ngo, T.V. & Chan, K.O. (2011) A new karstic cave-dwelling Cyrtodactylus Gray (Squamata: Gekkonidae) from northern Vietnam. Zootaxa, 3125, 51–63.

    Ngo, T.V. & Grismer, L.L. (2010) A new karst dwelling Cyrtodactylus (Squamata: Gekkonidae) from Son La Province, northwestern Vietnam. Hamadryad, 35, 84–95.

    Ngo, V.T. & Grismer, L.L. (2012) A new endemic species of Cyrtodactylus Gray (Squamata: Gekkonidae) from Tho Chu Island, southwestern Vietnam. Zootaxa, 3228 (1), 48–60.
    https://doi.org/10.11646/zootaxa.3228.1.2

    Ngo, V.T., Grismer, L.L. & Grismer, J.L. (2008) A new endemic cave dwelling species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) in Kien Giang Biosphere Reserve, southwestern Vietnam. Zootaxa, 1967, 53–62.

    Ngo, V.T., Grismer, L.L. & Grismer, J.L. (2010) A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) in Phu Quoc National Park, Kien Giang Biosphere Reserve, southwestern Vietnam. Zootaxa, 2604, 37–51.

    Ngo, T.V. & Pauwels, O.S.G. (2010) A new cave-dwelling species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Khammouane Province, southern Laos. Zootaxa, 2730 (1), 44–56.

    https://doi.org/10.11646/zootaxa.2730.1.3

    Nguyen, S.N., Orlov, N.L. & Darevsky, S.I. (2006) Descriptions of two new species of the genus Cyrtodactylus Gray, 1827 (Squamata: Sauria: Gekkonidae) from Southern Vietnam. Russian Journal of Herpetology, 13, 215–226.

    Nguyen, S.N., Yang, J-X., Le, N.T.T., Nguyen, L.T., Orlov, N.L., Hoang, C.V., Nguyen, T.Q., Jin, J-Q., Rao, D-Q., Hoang, T.N., Che, J., Murphy, R.W. & Zhang, Y-P. (2014) DNA barcoding of Vietnamese bent-toed geckos (Squamata: Gekkonidae: Cyrtodactylus) and the description of a new species. Zootaxa, 3784 (1), 48–66.

    https://doi.org/10.11646/zootaxa.3784.1.2

    Nguyen, T.Q., Le, M.D., Pham, A.V., Ngo, H.N., Hoang, C.V., Pham, C.T. & Ziegler, T. (2015a) Two new species of Cyrtodactylus (Squamata: Gekkonidae) from the karst forest of Hoa Binh Province, Vietnam. Zootaxa, 3985 (3), 375–390. https://doi.org/10.11646/zootaxa.3985.3.3

    Nguyen, T.Q, Pham, V.A., Nguyen, L.H.S., Le, D.M. & Ziegler, T. (2015b) First record of Parafimbrios lao Teynié, David, Lottier, Le, Vidal & Nguyen, 2015 (Squamata: Xenodermatidae) from Vietnam. Russian Journal of Herpetology, 22, 297–300.

    Nicolas, V., Herbreteau, V., Couloux, A., Keovichit, K., Douangboupha, B. & Hugot, J.-P. (2012) A remarkable case of micro-endemism in Laonastes aenigmamus (Diatomyidae, Rodentia) revealed by nuclear and mitochondrial DNA sequence data. PLOS ONE, 7, e48145.
    https://doi.org/10.1371/journal.pone.0048145

    Orlov, N.L., Nguyen, Nazarov, R.A., Anajeva, N.B. & Nguyen, S.N. (2007) A new species of the genus Cyrtodactylus Gray, 1827 and redescription of Cyrtodactylus paradoxus (Darevsky et Szczerbak, 1997) [Squamata: Sauria: Gekkonidae] from south Vietnam. Russian Journal of Herpetology, 14, 145–152.

    Panitvong, N., Sumontha, M., Tunprasert, J. & Pauwels, O.S.G. (2014) Cyrtodactylus saiyok sp. nov., a new dry evergreen forest-dwelling Bent-toed Gecko (Squamata: Gekkonidae) from Kanchanaburi Province, western Thailand. Zootaxa, 3869 (1), 64–74.
    https://doi.org/10.11646/zootaxa.3869.1.6

    Pauwels, O.S.G., Bauer, A.M., Sumontha, M. & Chanhome, L. (2004) Cyrtodactylus thirakhupti (Squamata: Gekkonidae), a new cave-dwelling gecko from southern Thailand. Zootaxa, 772, 1–11.

    Pauwels, O.S.G. & Sumontha, M. (2014) Cyrtodactylus samroiyot, a new limestone-dwelling Bent-toed Gecko (Squamata: Gekkonidae) from Prachuap Khiri Khan Province, peninsular Thailand. Zootaxa, 3755 (6), 573–583.
    https://doi.org/10.11646/zootaxa.3755.6.4

    Pauwels, O.S.G., Sumontha, M. & Bauer, A.M. (2016) A new Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Phetchaburi Province, Thailand. Zootaxa, 4088 (3), 409–419.

    https://doi.org/10.11646/zootaxa.4088.3.6

    Pauwels, O.S.G., Sumontha, M., Keeratikiat, K. & Phanamphon, E. (2014a) Cyrtodactylus kunyai (Squamata: Gekkonidae), a new cave-dwelling Bent-toed Gecko from Loei Province, northeastern Thailand. Zootaxa, 3821 (2), 253–264.

    Pauwels, O.S.G., Sumontha, M., Latinne, A. & Grismer, L.L. (2013) Cyrtodactylus sanook (Squamata: Gekkonidae), a new cave-dwelling gecko from Chumphon Province, southern Thailand. Zootaxa, 3635 (3), 275–285.

    https://doi.org/10.11646/zootaxa.3635.3.7

    Pauwels, O.S.G., Sumontha, M., Panitvong, N. & Varaguttanonda, V. (2014b) Cyrtodactylus khelangensis, a new cave-dwelling Bent-toed Gecko (Squamata: Gekkonidae) from Lampang Province, northern Thailand. Zootaxa, 3755 (6), 584–594.

    Pham, V.A., Le, T.D., Nguyen, L.H.S., Ziegler, T. & Nguyen, Q.T. (2014) First records of Leptolalax eos Ohler, Wollenberg, Grosjean, Hendrix, Vences, Ziegler & Dubois, 2011 and Hylarana cubitalis (Smith, 1917) (Anura: Megophryidae, Ranidae) from Viet Nam. Russian Journal of Herpetology, 21, 195–200.

    Pham, V.A., Le, T.D., Pham, T.C., Nguyen, L.H.S., Ziegler, T. & Nguyen, Q.T. (2016) Two additional records of megophryid species, Leptolalax minimus (Taylor, 1962) and Leptobrachium masatakasatoi Matsui, 2013, for the herpetofauna of Vietnam. Revue suisse de Zoologie, 123, 35–43.

    Posada, D. & Crandall, K.A. (1998) MODELTEST: testing the model of DNA substitution. Bioinformatics, 14, 817–818.
    https://doi.org/10.1093/bioinformatics/14.9.817

    Ronquist, F., Teslenko, M., van der Mark, P., Ayres, D.L., Darling, A., Höhna, S., Larget, B., Liu, L., Suchard, M.A. & Huelsenbeck, J.P. (2012) MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space. Systematic Biology, 61, 539–542.

    Schneider, N., Nguyen, T.Q., Schmitz, A., Kingsada, P., Auer, M. & Ziegler, T. (2011) A new species of karst dwelling Cyrtodactylus (Squamata: Gekkonidae) from northwestern Laos. Zootaxa, 2930, 1–21.

    Simmons, J.E. (2002) Herpetological collecting and collections management. Revised edition. Society for the Study of Amphibians and Reptiles. Herpetological Circular, 31, 1-153.

    Smith, M.A. (1917) Descriptions of new reptiles and a new batrachian from Siam. Journal of Natural History Society of Siam, 2, 221–225.

    Smith, M.A. (1921a) New or little-known reptiles and batrachians from southern Annam (Indo-China). Proceedings of the Zoological Society of London, 1921, 423–440.

    Smith, M.A. (1921b) Reptiles and Batrachians collected on Pulo Condore. Journal Natural History Society, 4, 93–97. [1920]

    Smith, M.A. (1935) The fauna of British India, including Ceylon and Burma. Vol. II. Sauria. Taylor and Francis, London, 8 + 440 pp.

    Sumontha, M., Panitvong, N. & Deein, G. (2010) Cyrtodactylus auribalteatus (Squamata: Gekkonidae), a new cave-dwelling gecko from Phitsanulok Province, Thailand. Zootaxa, 2370, 53–64.

    Sumontha, M., Pauwels, O.S.G., Kunya, K., Nitikul, A., Samphanthamit, P. & Grismer, L.L. (2012) A new forest-dwelling gecko from Phuket Island, Southern Thailand, related to Cyrtodactylus macrotuberculatus (Squamata: Gekkonidae). Zootaxa, 3522, 61–72.

    Sumontha, M., Pauwels, O.S.G., Suwannakarn, N., Nutatheera, N. & Sodob, W. (2014) Cyrtodactylus wangkulangkulae (Squamata: Gekkonidae), a new Bent-toed Gecko from Satun Province, southern Thailand. Zootaxa, 3821 (1), 116–124.
    https://doi.org/10.11646/zootaxa.3821.1.8

    Swofford, D.L. (2001) PAUP*. Phylogenetic Analysis Using Parsimony (* and Other Methods). Version 4. Sinauer Associates, Sunderland, Massachusetts.

    Taylor, E.H. (1963) Lizards of Thailand. The Universtiy of Kansas Science Bulletin, 55, 687–1077.

    The People's Committee of Son La Province (2007) Son La Province Portal. Available from: http://sonla.gov.vn/gioi-thieu (accessed 25 March 2017)

    Thompson, J.D., Gibson, T.J., Plewniak, F., Jeanmougin, F. & Higgins, D.G. (1997) The ClustalX windows interface: Xexible strategies for multiple sequence alignment aided by quality analysis tools. Nucleic Acids Research, 25, 4876–4882.

    Ulber, T. (1993) Bemerkungen über cyrtodactyline Geckos aus Thailand nebst Beschreibungen von zwei neuen Arten (Reptilia: Gekkonidae). Mitteilungen aus dem Zoologischen Museum in Berlin, 69, 187–200.
    https://doi.org/10.1002/mmnz.19930690202

    Ziegler, T., Nazarov, R., Orlov, N., Nguyen, T.Q., Vu, T.N., Dang, K.N., Dinh, T.H. & Schmitz, A. (2010) A third new Cyrtodactylus (Squamata: Gekkonidae) from Phong Nha – Ke Bang National Park, Vietnam. Zootaxa, 2413, 20–36.

    Ziegler, T., Phung, T.M., Le, M.D. & Nguyen, T.Q. (2013) A new Cyrtodactylus (Squamata: Gekkonidae) from Phu Yen Province, southern Vietnam. Zootaxa, 3686 (4), 432–446.

    https://doi.org/10.11646/zootaxa.3686.4.2

    Ziegler, T., Rösler, H., Herrmann, H.-W. & Vu, T.N. (2002) Cyrtodactylus phongnhakebangensis sp. n., ein neuer Bogenfingergecko aus dem annamitischen Karstwaldmassiv, Vietnam. Herpetofauna, 24, 11–25.