Abstract
The family Macrosternodesmidae is redefined and recorded from western North America. Four small-bodied species in Arizona and California, USA, and Baja California Norté, Mexico, are assigned to Tidesmus Chamberlin 1943; Phreatodesmus and Oodedesmus, both authored by Loomis, 1960, are placed in synonymy. Phreatodesmus torreyanus Loomis, 1960 and O. variabilis Loomis, 1960, are transferred into Tidesmus as valid species; P. cooki Loomis, 1960, is a synonym of T. episcopus Chamberlin, 1943, the type species, and P. dentatus Loomis, 1960, is a synonym of P. torreyanus. Brachydesmus hastingsus Chamberlin, 1941, also is referable to Tidesmus; a topotypical male is needed to establish its identity in the absence of authentic type specimens. Tidesmus hubbsi Chamberlin, 1943, based on unidentifiable females, is geographically segregated and incompatible with the otherwise coherent generic distribution. A topotypical male is also necessary to determine its identity; for now, we remove hubbsi from Tidesmus and leave it unassigned.
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