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Issue: Vol. 4387 No. 2: 27 Feb. 2018
Type: Article
Published: 2018-02-26
DOI: 10.11646/zootaxa.4387.2.3
Page range: 275–309
Abstract views: 280
PDF downloaded: 8

Taxonomic revision of the Cis-Andean species of Mylossoma Eigenmann & Kennedy, 1903 (Teleostei: Characiformes: Serrasalmidae)

Universidade Estadual Paulista, Departamento de Morfologia, Laboratório de Biologia e Genética de Peixes. Distrito de Rubião Júnior S/N, 18618-970 Botucatu, SP, Brazil.
Universidade Estadual Paulista, Departamento de Morfologia, Laboratório de Biologia e Genética de Peixes. Distrito de Rubião Júnior S/N, 18618-970 Botucatu, SP, Brazil.
Universidade Estadual de Maringá, Núcleo de Pesquisas em Limnologia, Ictiologia e Aquicultura. Av. Colombo 5790, 87020-900 Maringá, PR, Brazil. Universidade Estadual de Maringá, Programa de Pós-Graduação em Ecologia de Ambientes Aquáticos Continentais. Av. Colombo 5790, 87020-900 Maringá, PR, Brazil.
Pisces biodiversity freshwater fishes systematics taxonomy

Abstract

A revision of the cis-Andean species of Mylossoma is presented. Four species were recognized: M. albiscopum and M. aureum from the rio Amazonas and rio Orinoco basins; M. duriventre, from the rio Paraguai, lower rio Paraná and rio Uruguai basins; and M. unimaculatum, an endemic species from the Tocantins-Araguaia system. Both Mylossoma albiscopum and M. unimaculatum were removed from the synonymy of Mylossoma duriventre, which had its occurrence range restricted to the La Plata basin. The recognition of these four valid species corroborates the results of a previous DNA barcode study. Redescriptions of each species, and an identification key for the genus are provided.

 

References

  1. Ahl, E. (1928) Fischbestimmungsstelle. Das Aquarium: Die Fachzeitschrift des Naturfreundes, Berlin, 1, 192–194.

    Ahl, E. (1929) Beschreibung einer neuer sudamerikanischen characinidenart der gattung Mylossoma. Zoologischer Anzeiger, 81, 273–274.

    Baldwin, C.C. & Weigt, L.A. (2012) A new soapfish (Teleostei: Serranidae: Rypticus), with redescription of R. subbifrenatus and comments on the use of DNA barcoding in systematics studies. Copeia, 2012 (1), 23–36.
    https://doi.org/10.1643/CG-11-035

    Bemis, W.E., Hilton, E.J., Brown, B., Arrindell, R., Richmond, A.M, Little, C.D., Grande, L., Forey, P.L. & Nelson. G.J. (2004) Methods for preparing dry, partially articulated skeletons of osteichthyans, with notes on making ridewood dissections of the cranial skeleton. Copeia, 2004, 603–609.
    https://doi.org/10.1643/CI-03-054R1

    Benine, R.C., Mariguela, T.C. & Oliveira, C. (2009) New species of Moenkhausia Eigenmann, 1903 (Characiformes: Characidae) with comments on the Moenkhausia oligolepis species complex. Neotropical Ichthyology, 7 (2), 161–168.
    https://doi.org/10.1590/S1679-62252009000200005

    Bertin, L. (1948) Catalogue des types de poissons du Muséum National d'Histoire Naturelle, 3e partie: Ostariophysaires (Characiniformes, Gymnotiformes). Muséum national d'histoire naturelle, Paris, 49 pp.

    Bertoni, A.W. (1939) Catalogos sistematicos de los vertebrados del Paraguay. Revista de la Sociedad Cientifica del Paraguay, 4, 1–61.

    Böhlke, E.B. (1984) Catalog of type specimens in the Ichthyological Collection of the Academy of Natural Sciences of Philadelphia. Special Publication, Academy of Natural Sciences of Philadelphia, Philadelphia, 216 pp.

    Böhme, K. (1996) Briefe Franz Steindachners von der brasilien-expedition 1903. Annalen des Naturhistorisches Museums in Wien, 1996, 545–568.

    Brito, A., Lasso, C.A. & Sánchez-Duarte, P. (2011) Mylossoma duriventre. In: Lasso, C.A. & Sánchez-Duarte, P. (Eds.), Los peces del delta del Orinoco: diversidad, bioecología, uso y conservación. La Salle de Ciencias Naturales y Chevron, Caracas, pp. 129–130.

    Britski, H.A., Silimon, K.Z.S. & Lopes, B.S. (2007) Peixes do Pantanal: manual de identificação. 2nd Ed. Embrapa Informação Tecnológica, Brasília, 227 pp.

    Cabrera, V.H.G. & Vaca, H.C. (2006) Peces de Pando, Bolivia: especies de importancia comercial en mercados de la ciudad de Cobija. Environmental and Conservation Programs, División de peces the Field Museum y Universidad Amazónica de Pando, Chicago, 50 pp.

    Calcagnotto, D., Schaefer, S.A. & DeSalle, R. (2005) Relationships among Characiform fishes inferred from analysis of nuclear and mitochondrial gene sequences. Molecular Phylogenetics and Evolution, 36, 135–153.
    https://doi.org/10.1016/j.ympev.2005.01.004

    Cardoso, R.S. & Freitas, C.E.C. (2008) A pesca de pequena escala no rio Madeira pelos desembarques ocorridos em Manicoré (estado do Amazonas), Brasil. Acta Amazonica, 38, 781–788.
    https://doi.org/10.1590/S0044-59672008000400024

    Chernoff, B., Willink, P.W. & Montambault, J.R. (2001) A Biological assessment of the aquatic ecosystems of the río Paraguay basin, Alto Paraguay, Paraguay. In.: Chernoff, B., Willink, P.W. & Montambault, J.R. (Eds.), RAP Bulletin of Biological Assessment. Vol.19. Conservation International, Washington, pp. 1–156.

    Cope, E.D. (1872) On the fishes of the Ambyiacu River. Proceedings of the Academy of Natural Sciences of Philadelphia, 23 (3), 250–294.

    Costa-Silva, G.J., Rodriguez, M.S., Roxo, F.F., Foresti, F. & Oliveira, C. (2015) Using different methods to access the difficult task of delimiting species in a complex Neotropical hyperdiverse group. Plos ONE, 10 (9), e0135075.
    https://doi.org/10.1371/journal.pone.0135075

    Cuvier, G. (1818) Sur le poissons du sous-genre Myletes. Mémoires du Muséum d'Histoire Naturelle, 4, 444–456.

    Cuvier, M.G. & Valenciennes, A. (1850) Histoire naturelle des poissons. Tome Vingt- Deuxième. P. Bertrand éditeur, Paris, 532 pp.

    Devincenzi, G.J. (1925) El primer ensayo sobre ictyiología del Uruguay. La clase "Peces" de la zoología de Don Dámaso A. Larrañaga. Anales del Museo Nacional de Historia Natural de Montevideo, 2, 295–323.

    Eigenmann, C.H. (1903) New genera of South American fresh-water fishes, and new names for old genera. Smithsonian Miscellaneous Collections, 45, 144–148.

    Eigenmann, C.H. (1910) Catalogue of the fresh-water fishes of tropical and south temperate America. In: Reports of the Princeton University expeditions to Patagonia 1896–1899, Zoology, 3, 375–511.

    Eigenmann, C.H. (1915) The Serrasalminae and Myleinae. Annals of the Carnegie Museum, 9, 226–272, pls. XLIV–LVIII.

    Eigenmann, C.H. & Allen, W.R. (1942) Fishes of western South America. I – The intercordillera and Amazonian lowlands of Peru. II – The high pampas of Peru, Bolivia and northern Chile. The University of Kentucky, Lexington, xv + 494 pp.

    Eigenmann, C.H. & Kennedy, C.H. (1903) On a collection of fishes from Paraguay, with a synopsis of the American genera of Cichlids. Proceedings of the Academy of Natural Sciences of Philadelphia, 55, 497–537.

    Eschmeyer, W.N., Fricke, R. & van der Laan, R. (2017) Catalog of fishes. Genera, species, references. Available from: http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp (acessed 1 September 2017)

    Eschmeyer, W.N. & Fong, J.D. (2017) Catalog of fishes. Species by family/subfamily. Available from: http://research.calacademy.org/research/ichthyology/catalog/speciesbyfamily.asp (acessed 1 September 2017)

    Ferreira, E., Zuanon, J., Santos, G. & Amadio, S. (2011) A ictiofauna do Parque Estadual do Cantão, Estado do Tocantins, Brasil. Biota Neotropica, 11 (2), 277–284.
    https://doi.org/10.1590/S1676-06032011000200028

    Fowler, H.W. (1907) Further knowledge of some heterognathous fishes. Part II. Proceedings of the Academy of Natural Sciences of Philadelphia, 58, 431–483.

    Géry, J. (1972) Poissons characoïdes des Guyanes. I– Généralités. II– Famille des Serrasalmidae. Zoologische Verhandelingen, 122, 1–250.

    Géry, J. (1976) Les genres de Serrasalmidae (Pisces, Characoidei). Bulletin Zoölogisch Museum, Universiteit Van Amsterdam, 5, 47–54.

    Géry, J. (1977) The Characoids of the world. T.F.H. Publications, Neptune City, 672 pp.

    Géry, J., Mahnert, V. & Dlouhy, C. (1987) Poissons Characoïdes non Characidae du Paraguay (Pisces, Ostariophysi). Revue Suisse Zoologie, 94, 357–464.
    https://doi.org/10.5962/bhl.part.79720

    Gómez, S. & Chebez, J.C. (1996) Peces de la provincia de Misiones. In: Chebez, J.C. (Ed.), Fauna misionera: catálogo sistemático y zoogeográfico de los vertebrados de la provincia de Misiones (Argentina). L.O.L.A (Literature of Latin America), Buenos Aires, pp. 38–70.

    Gosline, W. (1951) Notes on the Characoid fishes of the subfamily Serrasalminae. Proceedings of the California Academy of Sciences, 27, 17–64.

    Goulding, M. (1980) The fishes and the forest: explorations in Amazonian natural history. University of California Press, Berkeley, 280 pp.

    Graça, W.J. & Pavanelli, C.S. (2007) Peixes da planície de inundação do alto rio Paraná e áreas adjacentes. Editora Universidade Estadual de Maringá, Maringá, 308 pp.

    Hebert, P.D.N., Cywinska, A., Ball, S.L. & Dewaard, J.R. (2003). Biological identifications through DNA barcodes. Proceedings of Royal Society, 270, 313–321.
    https://doi.org/10.1098/rspb.2002.2218

    Henn, A.W. (1928) List of types of fishes in the collection of the Carnegie Museum on september 1, 1928. Annals of the Carnegie Museum, 19, 51–99.

    Ibarra, M. & Stewart, D.J. (1987) Catalogue of type specimens of recent fishes in Field Museum of Natural History. Fieldiana Zoology, 35, 1–112.

    Janovetz, J. (2005) Functional morphology of feeding in the scale-eating specialist Catoprion mento. The journal of Experimental Biology, 208, 4757–4768.
    https://doi.org/10.1242/jeb.01938

    Jégu, M. (2003) Subfamily Serrasalminae: pacus and piranhas. In: Reis, R., Kullander, S.O. & Ferraris Jr., C.S. (Eds.), Check list of the freshwater fishes of South and Central America. Edipucrs, Porto Alegre, pp. 182–196.

    Jégu, M. (2004) Taxonomie des Serrasalminae phytophages et Phylogenie des Serrasalminae (Teleostei: Characiformes: Characidae). Museum National d'Histoire Naturelle, 421 pp.

    Jégu, M. & Santos, G.M. (1988). Une nouvelle espèce du genre Mylesinus (Pisces, Serrasalmidae), M. paucisquamatus, décrite du bassin du rio Tocantins (Amazonie, Brésil). Cybium, 12 (4), 331–341.

    Koerber, S., Vera-Alcaraz, H.S. & Reis, R.R. (2017) Checklist of the fishes of Paraguay (CLOFPY). Ichthyological Contributions of PecesCriollos, 53, 1–99.

    Kottelat, M. (1984) Catalogue des types du Musée d'Historie Naturelle de Neuchâtel. I– Pisces. Bulletin de la Societe Neuchâteloise des Sciences Naturelles, 107, 143–153.

    Kottelat, M. (1988) Authorship, dates of publication, status and types of Spix and Agassiz's brazilian fishes. Spixiana, 11, 69–93.

    Langeani, F., Castro, R.M.C., Oyakawa, O.T., Shibatta, O.A., Pavanelli, C.S. & Casatti, L. (2007) Diversidade da ictiofauna do Alto Rio Paraná: composição atual e perspectivas futuras. Biota Neotropica, 7 (3), 181–197.
    https://doi.org/10.1590/S1676-06032007000300020

    Larrañaga, D.A. (1923) Escritos de Don Dámaso Antonio Larrañaga. Edición Nacional 2. Instituto Histórico y Geográphico del Uruguay, Montevideo, 512 pp.

    Lima, F.C.T., Malabarba, L.R., Buckup, P.A., Silva, J.F.P., Vari, R.P., Harold, A., Benine, R., Oyakawa, O.T., Pavanelli, C.S., Menezes, N., Lucena, C.A.S., Malabarba, M.C., Lucena, Z.M.S., Reis, R.R., Langeani, F., Cassati, L., Bertaco, V., Moreira, C. & Lucinda, P.H.F. (2003) Genera Incertae Sedis in Characidae. In: Reis, R.R., Kullander, S.O. & Ferraris Jr., C.S. (Eds.), Check list of the freshwater fishes of South and Central America. Edipucrs, Porto Alegre, pp. 182–196.

    Lima, A.C. & Araújo-Lima, C.A.R.M. (2004) The distributions of larval and juvenile fishes in Amazonian rivers of different nutrient status. Freshwater Biology, 49, 787–800.

    Litz, T.O. & Koerber, S. (2014) Check List of the freshwater fishes of Uruguay (CLOFF-UY). Ichthyological Contributions of PecesCriollos, 28, 1–40.

    López, H. & Nass, P. (1989) Etapas del desarrollo de Mylossoma duriventris (Characiformes, Characidae) de los Llanos de Venezuela. Acta Biologica Venezuelica, 12, 121–126.

    López, H.L., Miquelarena, A.M. & Menni, R.C. (2003) Lista comentada de los peces continentales de la Argentina. Serie técnica y didáctica, Probiota, 85 pp.

    Lucinda, P.H.F., Freitas, I.S., Soares, A.B., Marques, E.E., Agostinho, C.S. & Oliveira, R.J. (2007) Fish, Lajeado Reservoir, rio Tocantins drainage, State of Tocantins, Brazil. Check list, 3 (2), 70–83.
    https://doi.org/10.15560/3.2.70

    Machado-Allison, A. (2005) Los Peces de los Llanos de Venezuela: Un ensayo sobre su historia natural. Universidad Central de Venezuela: Consejo de Desarrollo Cientifico y Humanistico, Caracas, 231 pp.

    Machado-Allison, A. & Castillo, O. (1992) Estudios sobre la sistematica de la subfamília Serrasalminae. IV. El genero Mylossoma: bases para la revision del grupo en America del Sur. Acta Biologica Venezuelica, 13, 1–34.

    Machado-Allison, A. & Fink, W. (1996) Los peces caribes de Venezuela: diagnosis, claves, aspectos ecologicos y evolutivos. Collection Monograph. Universidad Central de Venezuela, Caracas, 151 pp.

    MacDonagh, E. & Fuster, M. (1944) Sobre dos palometas (pirañas) y un pacucito (Pisces: Characinidae). Notas del Museo de La Plata, Zoología, 9, 349–362.

    Mateussi, N.T.B., Pavanelli, C.S. & Oliveira, C. (2016) Molecular identification of cryptic diversity in species of cis-Andean Mylossoma (Characiformes: Serrasalmidae). Mitochondrial DNA, 3 pp.
    https://doi.org/10.1080/24701394.2016.1180515

    Melo, B.F., Benine, R.C., Mariguela, T.C. & Oliveira, C. (2011) A new specis of Tetragonopterus Cuvier, 1816 (Characiformes: Characidae: Tetragonopterinae) from the rio Jari, Amapá, northern Brazil. Neotropical Ichthyology, 9 (1), 49–56.
    https://doi.org/10.1590/S1679-62252011000100002

    Menni, R.C. (2004) Peces y ambientes en la Argentina continental. Monografías del Museo Argentina de Ciencias Naturales, 5, 1–316.

    Merckx, A., Jégu, M. & Santos, G.M. (2000) Une nouvelle espèce de Serrasalmus (Teleostei: Characidae: Serrasalminae), S. altispinis n. sp., decrite du rio Uatumã (Amazonas, Brésil), avec une description complementaire de S. rhombeus (Linnaeus, 1766) du Plateau Guyanais. Cybium, 24 (2), 181–201.

    Mérona, B., Juras, A.A., Anastácio, A., Santos, G.M. & Cintra, I.H.A. (2010) Os peixes e a pesca no baixo rio Tocantins: vinte anos depois da UHE Tucuruí. Eletrobrás Eletronorte, Brasília, 208 pp.

    Mirande, J.M. (2010) Phylogeny of the family Characidae (Teleostei: Characiformes): from characters to taxonomy. Neotropical Ichthyology, 8, 385–568.
    https://doi.org/10.1590/S1679-62252010000300001

    Mirande, J.M. & Koerber, S. (2015) Checklist of the freshwater fishes of Argentina (CLOFFAR). Ichthyological Contributions of PecesCriollos, 36, 1–68.

    Miquelarena, A. (1986) Estudio de la dentición en peces caracoideos de la república Argentina. Biología Acuática, 8, 1–51.

    Myers, G.S. (1950) On the Characid fishes called Hydrocynus and Hydrocyon by Cuvier. Proceedings of the California Zoological Club, 1, 45–47.

    Nelson, E. (1961) The swimbladder in the Serrasalmidae. Fieldiana Zoology, 39, 603–624.

    Nico, L & Taphorn, D.C. (1988) Food habits of piranhas in the low Llanos of Venezuela. Biotropica, 20 (4), 311–321.
    https://doi.org/10.2307/2388321

    Norman, J.R. (1929) The South American Characid fishes of the subfamily Serrasalmoninae with a revision of the genus Serrasalmus Lacepede. Proceedings of the Zoological Society of London, 52, 781–829.

    Oldani, N.O. (1983) Identificación y morfología de larvas, juveniles y adultos de Mylossoma paraguayensis Norman, 1929. (Pisces, Characidae). Studies on Neotropical Fauna and Environment, 18, 89–100.
    https://doi.org/10.1080/01650528309360622

    Oliveira, E.C. & Araújo-Lima, C.A.R.M. (1998) Distribuição das larvas de Mylossoma aureum e M. duriventre (Pisces: Serrasalmidae) nas margens do rio Solimões, AM. Revista Brasileira de Biologia, 58, 349–358.
    https://doi.org/10.1590/S0034-71081998000300002

    Oliveira, C., Avelino, G.S., Abe, K.T., Mariguela, T.C., Benine, R., Ortí, G., Vari, R.P. & Castro, R.M.C. (2011) Phylogenetic relationships within the speciose family Characidae (Teleostei: Ostariophysi: Characiformes) based on multilocus analysis and extensive ingroup sampling. BMC Evolutionary Biology, 11, 1–25.
    https://doi.org/10.1186/1471-2148-11-275

    Ortega, H. & Vari, R.P. (1986) Annotated checklist of the freshwater fishes of Peru. Smithsonian Contributions to Zoology, 437, 1–25.
    https://doi.org/10.5479/si.00810282.437

    Ortí, G., Petry, P., Porto, J.I.R., Jégu, M. & Meyer, A. (1996) Patterns of nucleotide change in mitochondrial ribosomal RNA genes and the phylogeny of piranhas. Journal of Molecular Evolution, 42, 169–182.
    https://doi.org/10.1007/BF02198843

    Ortí, G., Sivasundar, A., Dietz, K. & Jégu, M. (2008) Phylogeny of the Serrasalmidae (Characiformes) based on mitochondrial DNA sequences. Genetics and Molecular Biology, 31, 343–351.
    https://doi.org/10.1590/S1415-47572008000200030

    Ota, R.P., Röpke, C.P., Zuanon, J. & Jégu, M. (2013) Serrasalmidae. In: Queiroz, L.J., Torrente-Vilara, G., Ohara, W.M., Pires, T.HS., Zuanon, J. & Doria, C.R.C. (Eds.), Peixes do rio Madeira. Vol. II. Santo Antônio Energia, São Paulo, pp. 14–47.

    Ramos, T.P.A., Ramos, R.T.C. & Ramos, S.A.Q.A. (2014) Ichthyofauna of the Parnaiba river basin, northeastern Brazil. Biota Neotropica, 14, 1–8.
    https://doi.org/10.1590/S1676-06020140039

    Resende, E.K., Pereira, R.A.C & Almeida, V.L.L. (1998) Peixes herbívoros da planície inundável do rio Miranda, Pantanal, Mato Grosso do Sul, Brasil. Embrapa–CPAP, Corumbá, 24 pp.

    Rodriguez, M., Cramer, C., Bonatto, S.L. & Reis, R. (2008) Taxonomy of Ixinandria Isbrücker & Nijssen (Loricariidae: Loricariinae) based on morphological and molecular data. Neotropical Ichthyology, 6, 367–378.
    https://doi.org/10.1590/S1679-62252008000300010

    Sabaj, M.H. (2016) Standard symbolic codes for institutional resource collections in herpetology and ichthyology: an Online Reference. Version 6.5 (16 august 2016). Available from: http://www.asih.org/resources (acessed 1 September 2017)

    Santos, G.M., Mérona, B., Juras, A.A. & Jégu, M. (2004) Peixes do baixo rio Tocantins: 20 anos depois da usina hidrelétrica de Tucuruí. Brasília, Eletronorte, 216 pp.

    Santos, G.M., Ferreira, E. & Zuanon, J. (2006) Peixes comerciais de Manaus. Manaus, Ibama/AM, Provárzea, 144 pp.

    Sazima, I. (1983) Scale-eating in characoids and other fishes. Enviromental Biology of Fishes, 9, 87–101.
    https://doi.org/10.1007/BF00690855

    Sazima, I. & Machado, F.A. (1990) Underwater observations of the piranhas in western Brazil. Environmental Biology of Fishes, 28, 17–31.
    https://doi.org/10.1007/BF00751026

    Silva, G.S.C., Melo, B.F., Oliveira, C. & Benine, R.C. (2013) Morphological and molecular evidence for two new species of Tetragonopterus (Characiformes: Characidae) from central Brazil. Journal of Fish Biology, 82 (5), 1613–1631.

    https://doi.org/10.1111/jfb.12094

    Spix, J.B. & Agassiz, L. (1829–31) Selecta genera et species piscium quos in itinere per brasiliam annos mdcccxvii–mdcccxx jussu et auspiciis maximiliani josephi i. colleget et pingendso curavit dr J. B. de Spix. Part 1 & 2. Typis C. Wolf, Monachii, xvi + ii + 82 pp., 48 pls. & 56 pp., 53 pls. [Part 1: pp. i–xvi + i–ii + 1–82, pls. 1–48; Part 2: pp. 83–138, pls. 49–101]

    Steindachner, F. (1908) Über eine neue Metynnis–art (fam. Caracidae) aus einer lagune am rio Medonho. Anzeiger der Akademie der Wissenschaften in Wien, 45, 326–327.

    Sverlij, S.B., Schenke, R.L.D., López, H.L. & Ros, A.E. (1998) Peces del río Uruguay. Guia ilustrada de las especies mas comúnes del río Uruguay inferior y el embalse de Salto Grande. Comision administradora del rio Uruguay, Paysandú, 89 pp.

    Taphorn, P. (1992) The Characiform fishes of the Apure river drainage, Venezuela. Biollania edición especial. No. 4. Monografias Cientificas del Museo de Ciencias Naturales, Guanara, 537 pp.

    Thompson, A.W., Betancur-R, R., López-Fernández, H. & Ortí, G. (2014) A time-calibrated, multi-locus phylogeny of piranhas and pacus (Characiformes: Serrasalmidae) and a comparison of species tree methods. Molecular Phylogenetics and Evolution, 81, 242–257.

    https://doi.org/10.1016/j.ympev.2014.06.018

    Vanzolini, P.E. (2004) Episódios da zoologia brasílica. São Paulo, Editora Hucitec, 212 pp.

    Vitule, J.R.S., Skóra, F. & Abilhoa, V. (2012) Homogenization of freshwater fish faunas after the elimination of a natural barrier by a dam in neotropics. Diversity and Distributions, 18, 111–120.
    https://doi.org/10.1111/j.1472-4642.2011.00821.x

    Zarske, A. & Géry, J. (1999) Revision der neotropischen Gattung Metynnis Cope, 1878. 1. Evaluation der Typusexemplare der nominellen Arten (Teleostei: Characiformes: Serrasalmidae). Zoologische Abhandlungen, 13, 169–216.