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Type: Monograph
Published: 2018-07-20
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The upper Miocene chitons of northwest France (Mollusca: Polyplacophora)

Via Briscata 16/6, I-16154 Genova, Italy.
Naturalis Biodiversity Center, P.O. Box 9517, 2300 RA Leiden, The Netherlands;
Cambeenboslaan A 11, B-2960 Brecht, Belgium.
Avenue Général Naessens de Loncin 1, B-1330 Rixensart, Belgium
Mollusca Polyplacophora northwestern France upper Miocene new taxa

Abstract

This study describes the chiton fauna (Mollusca, Polyplacophora) from the upper Miocene Tortonian deposits of northwest France previously known as the ‘Redonian’. Twenty-two species were identified (represented by 9,380 valves), of which seven were already known, one is assigned at generic level (Leptochiton sp.) and 14 are described as new (Lepidopleurus pseudobenoisti sp. nov., L. gallicus sp. nov., Leptochiton parvus sp. nov., Leptochiton lateropustulosus sp. nov., Leptochiton renauleauensis sp. nov., Hanleya sancticlementensis sp. nov., H. sossoi sp. nov., Ischnochiton nitidum sp. nov., Callochiton pouweri sp. nov., Rhyssoplax assurrectum sp. nov., Tonicella redoniensis sp. nov., Acanthochitona globosa sp. nov., A. andegavensis sp. nov. and A. chauvereauensis sp. nov.). Regarding the stratigraphic distribution of the 21 taxa identified, 15 are restricted to the Miocene, the remaining six are extant and are distributed in the Atlantic and Mediterranean (Lepidopleurus cajetanus, Leptochiton algesirensis, Hanleya hanleyi, Ischnochiton rissoi, Callochiton doriae and Rhyssoplax corallinus. The number of new species is high (14 out of 22), possibly explained in part by the fact that this is the first description of the northwestern French upper Miocene chiton fauna - true endemicity is less likely in the light of scarce sampling.

 

References

  1. Adams, H. & Adams, A. (1853–1858) The genera of recent Mol-lusca; arranged according to their organization. Vol. 1 & 2. John van Voorst, London, 484 pp. & 660 pp. [Vol. 1: pp. 1–256, pls. 1–32 (1853), 257–484 (1854); Vol. 2. pp. 1–284, pls. 33–96 (1855), 285–412, pls. 97–112 (1856), 413–540, pls. 113–128 (1857), 541–660, pls. 129–138 (1858)]

    Antevs, E. (1917) Post-glacial marine shell-beds in Bohuslan. Geologiska föreningens i Stockholm förhandlingar, 39 (4), 247–425.
    https://doi.org/10.1080/11035891709444845

    Antevs, E. (1928) Shell beds in the Skagerack. Geologiska föreningens i Stockholm förhandlingar, 50 (4), 479–749.
    https://doi.org/10.1080/11035897.1928.9626355

    Arnaud, P.M. (1977) Revision des taxa malacologiques mediterraneens introduits par Antoine Risso. Annales du Muséum d’Histoire Naturelle de Nice, 5, 101–150, pls. 8–12.

    Ashby, E. (1925) Monograph on Australian Fossil Polyplacophora (chitons). Proceedings of the Royal Society of Victoria, 37, 170–205, pls. 18–22.

    Ashby, E. (1929) New Zealand fossil Polyplacophora (chitons). Transactions and Proceedings of the New Zealand Institute, 60, 366–369, pl. 32.

    Ashby, E. (1939) Fossil chitons from Mornington, Victoria. Proceedings of the Linnean Society of London, 3, 186–189, pl. 3.

    Ashby, E. & Cotton, B.C. (1936) South Australian fossil chitons. Records of the South Australian Museum, 5, 509–512.

    Ashby, E. & Cotton, B.C. (1939) New fossil chitons from the Miocene and Pliocene of Victoria. Records of the South Australian Museum, 6, 209–242, pls. 19–21.

    Bałuk, W. (1965) Chitony z piaskow tortonskich kotliny Sadeckiej [Chitons from the Tortonian of the Nowy Sacz depression, Poland]. Acta Palaeontologica Polonica, 10, 365–378, 1 pl.

    Bałuk, W. (1971) Lower Tortonian chitons from the Korytnica clays, southern slopes of the Holy Cross Mts. Acta geologica Polonica, 21, 449–472, pls. 1–6.

    Bałuk, W. (1984) Additional data on chitons and cuttlefish from the Korytnica clays (Middle Miocene; Holy Cross Mountains, Central Poland). Acta geologica Polonica, 34, 281–297, pls. 1–17.

    Beets, C. (1941) Eine jungmiocane Mollusken-Fauna von der Halbinsel Mangkalihat, Ost–Borneo. Geologisch–Mijnbouwkundig Genootschap voor Nederland en Kolonien Verhandelingen, 13, 1–219, pls. 1–9.

    Bergenhayn, J.R.M. (1930) Kurze Bemerkungen zur Kenntnis der Schalenstruktur und Systematik der Loricaten. Kungliga Svenska vetenskapsakademiens handlingar, Series 3, 9, 1–54, pls. 1–10.

    Bergenhayn, J.R.M. (1955) Die Fossilen Schwedischen Loricaten nebst einer vorlaufigen revision des systems der ganzen klasse Loricata. Acta Universitatis Lundensis, New Series, Series 2, 51, 1–44, pls. 1–2.

    Berry, C.T. (1940) Some fossil Amphineura from the Atlantic coastal plain of North America. Proceedings of the Academy of Natural Sciences of Philadelphia, 91, 207–217, pls. 9–12

    Berry, S.S. (1922) Fossil chitons of western North America. Proceedings of the California Academy of Sciences, Series 4, 11, 399–526.

    Beu, A.G., Dell, R.K. & Fleming, C.A. (1969) Rhyssoplax Thiele, 1893 (Mollusca, Amphineura): request for designation of a type-species under the plenary powers. Bulletin of Zoological Nomenclature, 25, 184–185.

    Beu, A.G. & Maxwell, P.A. (1990) Cenozoic Mollusca of New Zealand. New Zealand Geological Survey Paleontological Bulletin, 58, 1–518.

    Bielokrys, L.S. (1999) Late Eocene Chitonids (Class Polyplacophora) from Ukraine. Paleontological Journal, 33, 339–349 (translated from Paleontologicheskii Zhurnal, 4, 5–15).

    Boettger, O. (1905) Zur Kenntniss der Fauna der mittelmiocänen Schichten von Kostej im Krassó-Szörényer Komitat. Verhandlungen und Mittheilungen des siebenbürgischen Vereins für Naturwissenschaften zu Hermannstadt, 55, 102–217.

    Bonfitto, A., Dell'Angelo, B., Evangelisti, F. & Sabelli, B. (2011) The genus Acanthochitona (Mollusca: Polyplacophora) in the Mediterranean Sea: morphological and molecular data. Scientia Marina, 75, 171–180.
    https://doi.org/10.3989/scimar.2011.75n1171

    Brébion, P. (1964) Les gastéropodes du Redonien et leur signification, 1–2. Thèse de doctorat, Faculté des Sciences de l’Université de Paris, 775 pp., 15 pls.

    Broderip, W.J. & Sowerby, G.B. (1832–1833) Characters of new species of Mollusca and Conchifera, collected by Mr. Cuming. Proceedings of the Zoological Society of London for 1832, 25–33, 50–61, 104–108, 124–126, 173–179 (1832), 194–202 (1833).

    Capellini, J. (1859) Catalogue des oscabrions de la Méditerranée suivi de la description de quelques espèces nouvelles. Journal de Conchyliologie, 7, 321–328, pl. 12.

    Carmona Zalvide, M.P. & García, F.J. (2000) Moluscos poliplacóforos del litoral atlántico del sur de la península Ibérica. Graellsia, 56, 5–14.
    https://doi.org/10.3989/graellsia.2000.v56.i0.304

    Carmona Zalvide, P. & García, F.J. (1999) Consideraciones morfológicas y taxonómicas sobre Ischnochiton (Ischnochiton) rissoi (Payraudeau, 1826) (Mollusca Polyplacophora). Graellsia, 55, 177–186.
    https://doi.org/10.3989/graellsia.1999.v55.i0.326

    Carmona Zalvide, P., García, F.J. & Urgorri, V. (2002) Consideraciones taxonómicos y zoogeográficas del género Callochiton Gray, 1847 en la península ibérica. Archiv für Molluskenkunde, 131, 185–199.
    https://doi.org/10.1127/arch.moll/131/2002/185

    Carpenter, P.P. (1873) On the generic affinities of the New England chitons. Bulletin of the Essex Institute, 5, 152–155.

    Castany, G., Gobert, G. & Harson, L. (1956) Le Quaternaire marin de Monastir. Annales Mines and Geology, 19, 1–58, 2 pls.

    Cavet, P., Arnaud, A., Blaise, J., Brossé, R., Chauris, L., Gruet, M., Lardeux, H. & Service géologique régional Bretagne-Pays de la Loire (1976) Carte Géologique de la France à 1/50000 (N° 454). Feuille Angers (map & explanatory note). Bureau de Recherches Géologiques et Minières, Orléans, 454, 1–43, map.

    Ceulemans, L., Landau, B.M. & Van Dingenen, F. (2014) Carinofusus gen. nov. from the Mio–Pliocene transition of western France. Vita Malacologica, 12, 23–30.

    Ceulemans, L., Van Dingenen, F. & Landau, B.M. (2016a) The lower Pliocene gastropods of Le Pigeon Blanc (Loire–Atlantique, Northwest France). Patellogastropoda and Vetigastropoda. Cainozoic Research, 16 (1), 51–100.

    Ceulemans, L., Van Dingenen, F., Merle, D. & Landau, B.M. (2016b) The lower Pliocene gastropods of Le Pigeon Blanc (Loire-Atlantique, northwest France). Part 3. Muricidae. Vita Malacologica, 15, 35–55.

    Chirli, C. (2004) Malacofauna Pliocenica Toscana. Vol. 4°. Polyplacophora Gray J.E., 1821. Monoplacophora Odhner, 1940. Archaeogastropoda Thiele, 1925. B.B.M., Firenze, 113 pp.

    Costa, O.G. (1830) Catalogo sistematico e ragionato de’ testacei delle Due Sicilie. Tipografia della Minerva, Napoli, 8 + cxxxii pp., 2 pls. [1829]

    Cotton, B.C. & Godfrey, F.K. (1940) Australian Fossil Chitons. In: Handbook, Molluscs of South Australia. Part 2. South Australian Museum, Adelaide, pp. 569–590.

    Couffon, O. (1915) Contribution à l’étude des faluns de l’Anjou, 4. Miocène supérieur, gisement de Saint-Michel-et-Chanveaux. Bulletin de la Société d’Études Scientifiques d’Angers, New Series, 44, 31–56.

    Dall, W.H. (1878) Descriptions of new forms of mollusks from Alaska contained in the collection of the National Museum. Proceedings of United States National Museum, 1, 1–3.
    https://doi.org/10.5479/si.00963801.1-2.3

    Dall, W.H. (1879) Report on the limpets and chitons of the Alaskan and Arctic regions, with descriptions of genera and species believed to be new. Proceedings of United States National Museum, 1, 63–126, pls. 1–5.
    https://doi.org/10.5479/si.00963801.1-48.281

    Dall, W.H. (1889) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877-78) and in the Caribbean Sea (1879-80), by the U.S. Coast Survey Steamer “Blake”, Lieut.-Commander C.D. Sigsbee, U.S.N., and Commander J.R. Bartlett, U.S.N., commanding, 29: Report on the Mollusca, 2. Gastropoda and Scaphopoda. Bulletin of the Museum of Comparative Zoology, 18, 1–492.

    Davis, A.G. (1954) Lower Miocene Chitons from Mafia Island, East Africa. Proceedings of the Malacological Society of London, 31, 17–20.

    De Lamothe (1911) Les anciennes lignes de rivage du Sahel d'Alger et d'une part de la cote algèrienne. Mémoires de la Société géologique de France, Series 4, 1 (6), 288 pp.

    Dell’Angelo, B. & Bonfitto, A. (2005) Note on Fossil Chitons. 1. A new species of Lepidopleurus (Mollusca: Polyplacophora) from the Pleistocene of Salice (Sicily, Italy). Zootaxa, 821 (1), 1–6.
    https://doi.org/10.11646/zootaxa.821.1.1

    Dell’Angelo, B., Bonfitto, A. & Taviani, M. (2011a) Chitons (Polyplacophora) from Paleogene strata in Western Washington State, U.S.A. Journal of Paleontology, 85, 936–954.
    https://doi.org/10.1666/10-114.1

    Dell'Angelo, B. & Forli, M. (1995) I Polyplacophora del Pleistocene inferiore di Riparbella (Pisa) con elenco dei molluschi rinvenuti. Bollettino Malacologico, 30, 221–252.

    Dell'Angelo, B., Forli, M. & Lombardi, C. (2001) I Polyplacophora Plio-Pleistocenici della Toscana. Bollettino Malacologico, 36, 143–154.

    Dell'Angelo, B., Garilli, V., Germanà, A., Reitano, A., Sosso, M. & Bonfitto, A. (2012) Notes on fossil chitons. 4. Polyplacophora from the Pliocene of Altavilla (NW Sicily). Bollettino Malacologico, 48, 51–68.

    Dell’Angelo, B., Giuntelli, P., Sosso, M. & Zunino, M. (2014) Notes on Fossil Chitons. 6. A new species of Stenoplax (Mollusca: Polyplacophora) from the Miocene of NW Italy. Bollettino della Società Paleontologica Italiana, 53, 49–54.

    Dell’Angelo, B., Giuntelli, P., Sosso, M. & Zunino, M. (2015) Polyplacophora from the Miocene of North Italy. Part 1: Leptochitonidae, Hanleyidae, Ischnochitonidae and Callistoplacidae. Rivista Italiana di Paleontologia e Stratigrafia, 121, 217–242.

    Dell’Angelo, B., Giuntelli, P., Sosso, M. & Zunino, M. (2016) Polyplacophora from the Miocene of North Italy. Part 2: Callochitonidae, Chitonidae, Lepidochitonidae, Acanthochitonidae and Cryptoplacidae. Rivista Italiana di Paleontologia e Stratigrafia, 122, 23–54.

    Dell’Angelo, B. & Giusti, F. (1997) I Polyplacophora di una tafocenosi profonda. La Conchiglia, 29 (283), 51–58.

    Dell'Angelo, B., Giusti, .F., Paolini, P., Sosso, M. & Bonfitto, A. (2009) New data on Polyplacophora from Tuscan Archipelago. I. Leptochiton sarsi Kaas, 1981 and Leptochiton pepezamorai Carmona Zalvide, Urgorri & Garcia, 2004, two species new to the Mediterranean Sea. Italian Journal of Zoology, 76, 76–82.
    https://doi.org/10.1080/11250000802137061

    Dell'Angelo, B., Grigis, M. & Bonfitto, A. (2007) Notes on fossil chitons. 2. Polyplacophora from the Middle Miocene of Lăpugiu (Romania). Bollettino Malacologico, 43, 39–50.

    Dell'Angelo, B., Landau, B. & Marquet, R. (2004) Polyplacophora from the Early Pliocene of Estepona (Málaga, southwest Spain). Bollettino Malacologico, 5 (Supplement), 25–44.

    Dell'Angelo, B., Lesport, J.-F., Cluzaud, A. & Sosso, M. (2018) The Oligocene to Miocene chitons (Mollusca: Polyplacophora) of the Aquitaine Basin, southwestern France, and Ligerian Basin, western France. Part 1: Leptochitonidae, Hanleyidae, Ischnochitonidae, Chitonidae, Spinochitonidae fam. nov. and Schizochitonidae. Bollettino Malacologico, 54, 1–47.

    Dell'Angelo, B. & Palazzi, S. (1989) Considerazioni sulla famiglia Leptochitonidae Dall, 1889 (Mollusca: Polyplacophora). III. Le specie terziarie e quaternarie europee, con note sistematiche e filogenetiche. Atti Prima Giornata di Studi Malacologici CISMA, 1989, 19–140.

    Dell'Angelo, B. & Palazzi, S. (1991) Considerazioni sulla famiglia “Leptochitonidae" Dall, 1889 (Mollusca: Polyplacophora). IV. Aggiunte e correzioni. Bollettino Malacologico, 27, 35–38.

    Dell'Angelo, B. & Palazzi, S. (1992) First record of a species of Polyplacophora in the Italian Oligocene. Journal of the Malacological Society of Australia, 13, 27–30.

    Dell’Angelo, B. & Palazzi, S. (1994) Callochiton calcatus n. sp. con note su Callochiton septemvalvis (Montagu, 1803). La Conchiglia, 26 (273), 15–23.

    Dell'Angelo, B., Palazzi, S. & Pavia, G. (1999) I molluschi del Messiniano Inferiore di Borelli (Torino). 4. Polyplacophora. Bollettino del Museo Regionale di Scienze Naturali, 16, 257–302.

    Dell'Angelo, B., Prelle, G., Sosso, M. & Bonfitto, A. (2011b) Intertidal chitons (Mollusca: Polyplacophora) from southern Madagascar. African Invertebrates, 52 (1), 21–37.
    https://doi.org/10.5733/afin.052.0103

    Dell'Angelo, B., Renda, W., Sosso, M., Sigwart, J.D. & Giacobbe, S. (2017) A new species of Callochiton (Mollusca: Polyplacophora) from the Strait of Messina (central Mediterranean). Archiv für Molluskenkunde, 146, 243–250.
    https://doi.org/10.1127/arch.moll/146/243-250

    Dell'Angelo, B. & Silva, C.M. da (2003) Polyplacophora from the Pliocene of Vale de Freixo: Central–West Portugal. Bollettino Malacologico, 39, 7–16.

    Dell'Angelo, B. & Smriglio, C. (1999) Chitoni Viventi del Mediterraneo. Edizioni Evolver, Roma, 256 pp. [English Ed. 2001: Living chitons of the Mediterranean]

    Dell’Angelo, B., Sosso, M., Prudenza, M. & Bonfitto, A. (2013) Notes on Fossil Chitons. 5. Polyplacophora from the Pliocene of Western Liguria, Northwest Italy. Rivista Italiana di Paleontologia e Stratigrafia, 119, 65–107.

    Dulai, A. (2001) Middle Miocene (Badenian) Polyplacophora (Mollusca) remains from borehole Szokolya–2 (Börzsöny Mts, Hungary, Central Paratethys). Fragmenta Palaeontologica Hungarica, 19, 39–49.

    Dulai, A. (2005) Badenian (Middle Miocene) Polyplacophora from the Central Paratethys (Bánd and Devecser, Bakony Mountains, Hungary). Fragmenta Palaeontologica Hungarica, 23, 29–50.

    Fabricius, O. (1780) Fauna Gronlandica. Impensis Ioannis Gottlob Rothe, Hafniae, 452 pp.

    Ferreira, A.J. (1979) The family Lepidopleuridae (Mollusca: Polyplacophora) in the Eastern Pacific. The Veliger, 22, 145–165.

    Ferreira, A.J. (1982) The Family Lepidochitonidae Iredale, 1914 (Mollusca: Polyplacophora) in the Eastern Pacific. The Veliger, 25 (2), 93–138a.

    Ferreira, A.J. (1983) Researches on the coast of Somalia. The chiton fauna (Mollusca Polyplacophora). Monitore Zoologico italiano, 9, 249–297.
    https://doi.org/10.1080/00269786.1983.11758574

    Ferrero Mortara, E.L., Montefameglio, L., Novelli, M., Opesso, G., Pavia, G. & Tampieri, R. (1984) Catalogo dei tipi e degli esemplari figurati della collezione Bellardi e Sacco. Parte II. Museo Regionale di Scienze Naturali, Cataloghi, 7, 1–484.

    Fischer, F.P. & Renner, M. (1979) SEM-Observations on the shell plates of three polyplacophorans (Mollusca, Amphineura). Spixiana, 2, 49–58.

    Garilli, V., Dell'Angelo, B. & Vardala-Theodorou, E. (2005) Polyplacophora from the Pleistocene of Kyllini (NW Peloponnese, Greece). Bollettino della Società Paleontologica Italiana, 44, 117–134.

    Gmelin, J.F. (1791) Caroli a Linnei systema natura per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentis, synonymis, locis etc. Editio decima tertia, aucta, reformata, cura J.F. Gmelin. 1 (6). Vermes testacea. G.E. Beer, Lipsiae, 1100 pp. [pp. 3021–4120]

    Gowlett-Holmes, K.L. (1998) Family Cryptoplacidae. In: Beesley, P.L., Ross, G.J.B. & Wells, A. (Eds.), Mollusca—The Southern Synthesis. Part A. Fauna of Australia. Vol. 5. CSIRO Publishing, Melbourne, pp. 189.

    Gray, J.E. (1821) A natural arrangement of Mollusca, according to their internal structure. London Medical Repository, 15, 229–239.

    Gray, J.E. (1828) Spicilegia Zoologica; or original figures and short systematic descriptions of new and unfigured animals. Vol. 1. Treüttel, Würtz and Co., London, 2 pp., 2 pls. [pp. 5–6, pls. 3 & 6]

    Gray J.E. (1847) A list of the genera of Recent Mollusca, their synonyms and types. Proceedings of the Zoological Society of London, 15, 129–206.

    Gray, J.E. (1857) Guide to the systematic distribution of Mollusca in the British Museum. Vol. 1. Taylor & Francis, London, xii + 230 pp.

    Gürs, K. (1992) Zwei neue Polyplacophorenarten (Mollusca) aus marinen Küstenablagerungen des Mainzer Beckens und der Kasseler Bucht (Oligozän, Rupelium und Chattium). Geologisches Jahrbuch Hessen, 120, 5–9.

    Herrmannsen, A.N. (1846–1847) Indicis Generum Malacozoorum Primordia. Vol. 1. T. Fisheri, Cassell, 637 pp.

    Issel, A. (1869) Malacologia del Mar Rosso. Ricerche zoologiche e paleontologiche. Biblioteca Malacologica, Pisa, xi + 388 pp., 5 pls.

    Itoigawa, J., Kuroda, M., Naruse, A. & Nishimoto, H. (1976) Polyplacophora assemblages from the Pleistocene formations of Bösö and Miura Peninsulas, environs of Tokyo, Japan. Bulletin of the Mizunami Fossil Museum, 3, 171–204.

    Itoigawa, J., Kuroda, M., Naruse, A., Nishimoto, H., Asada, T., Iwai, T. & Hayashi, K. (1978) Polyplacophora Assemblages from the Pleistocene Formations of Kisarazu, Ichihara and Their Environs, Boso Peninsula, Japan. Bulletin of the Mizunami Fossil Museum, 5, 143–155.

    Itoigawa, J. & Nishimoto, H. (1975) Miocene Polyplacophora assemblages from the Mizunami group, central Japan (First report). Bulletin of the Mizunami Fossil Museum, 2, 35–50, pls. 9–11.

    Itoigawa, J., Shibata, H., Nishimoto, H. & Okumura, Y. (1981–1982) Miocene fossils of the Mizunami group, central Japan. 2. Molluscs. Monograph of the Mizunami Fossil Museum, 3A & 3B, 1–55, pls. 1–52 & 1–330.

    Janjou, D., Lardeux, H., Chantraine, J., Callier, L. & Étienne, H. (1998) Carte Géologique de la France à 1/50000 (N° 422). Feuille Segré (explanatory note). Bureau de Recherches Géologiques et Minières, Orléans, 422, 1–68.

    Janssen, R. (1978) Revision der Polyplacophora des Oligozäns in Mitteleuropa. Archiv für Molluskenunde, 108, 215–235.

    Jeffreys, J.G. (1880) On a new species of chiton lately found on the British Coasts. Annals and Magazine of Natural History, Series 5, 6, 33–35.
    https://doi.org/10.1080/00222938009458892

    Kaas, P. (1979) On a collection of Polyplacophora (Mollusca, Amphineura) from the bay of Biscay. Bulletin du Muséum national d'Histoire Naturelle, Section A, (4) 1, 13–31.

    Kaas, P. (1981) Scandinavian species of Leptochiton Gray, 1847 (Mollusca: Polyplacophora). Sarsia, 66, 217–229.
    https://doi.org/10.1080/00364827.1981.10414541

    Kaas, P. (1985) The genus Acanthochitona Gray, 1821 (Mollusca, Polyplacophora) in the north–eastern Atlantic ocean and in the Mediterranean Sea, with designation of neotypes of A. fascicularis (L., 1767) and of A. crinita (Pennant, 1777). Bulletin du Muséum national d'Histoire Naturelle, Section A, Series 4, 7, 579–609.

    Kaas, P. & Van Belle, R.A. (1981) The genus Lepidochitona Gray, 1821 (Mollusca: Polyplacophora) in the Northeastern Atlantic Ocean, the Mediterranean Sea and the Black Sea. Zoologische Verhandelingen, 185, 3–43.

    Kaas, P. & Van Belle, R.A. (1985a) Monograph of Living Chitons (Mollusca: Polyplacophora). Vol. 1. Order Neoloricata: Lepidopleurina. E.J. Brill, W. Backuys, Leiden, 240 pp.

    Kaas, P. & Van Belle, R.A. (1985b) Monograph of Living Chitons (Mollusca: Polyplacophora). Volume 2. Suborder Ischnochitonina, Ischnochitonidae: Schizoplacinae, Callochitoninae & Lepidochitoninae. E.J. Brill, W. Backhuys, Leiden, 198 pp.

    Kaas, P. & Van Belle, R.A. (1990) Monograph of Living Chitons (Mollusca: Polyplacophora). Vol. 4. Suborder Ischnochitonina: Ischnochitonidae: Ischnochitoninae (continued). Additions to Vols. 1, 2 & 3. E.J. Brill, Leiden, 298 pp.

    Kaas, P. & Van Belle, R.A. (1994) Monograph of Living Chitons (Mollusca: Polyplacophora). Volume 5. Suborder Ischnochitonina: Ischnochitonidae: Ischnochitoninae (concluded), Callistoplacinae; Mopaliidae. Additions to Vols. 1–4. E.J. Brill, Leiden, 402 pp.

    Kaas, P., Van Belle, R.A. & Strack, H.L. (2006) Monograph of Living Chitons (Mollusca: Polyplacophora). Vol. 6. Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae. Additions to Vols. 1–5. Brill, Leiden-Boston, 463 pp.

    Kangas, M. & Shepherd, S.A. (1984) Distribution and feeding of chitons in a boulder habitat at West Island, South Australia. Journal of the Malacological Society of Australia, 6, 101–111.

    Koenen, A. von (1889-1894) Das Norddeutsche Unter-Oligocän und seine Mollusken-Fauna, 1–7. Abhandlungen zur geolo-gischen Specialkarte von Preussen und den Thüringischen Staaten 10, 1–1458. [issued in parts 1, 1–280, pls. 1–23 (1889); 2, 281–574 pls. 24–34 (1890); 3, 575–817, pls. 35–52 (1891); 4, 818–1004, pls. 53–62 (1892); 5, 1005–1248, pls. 63–86 (1983); 6, 1249–1392, pls. 87–99 (1894); 7, 1393–1458, pls. 100–101 (1894)]

    Koskeridou, E., Vardala–Theodorou, E. & Moissette, P. (2009) Pliocene and Pleistocene shallow–water chitons (Mollusca) from Rhodes Island, Greece. Neues Jahrbuch für Geologie und Paläontologie—Abhandlungen, 251, 303–330.
    https://doi.org/10.1127/0077-7749/2009/0251-0303

    Kroh, A. (2003) The Polyplacophora (Mollusca) of the Langhian (Lower Badenian) of the Molasse Zone and the northern Vienna Basin (Austria). Annalen des Naturhistorischen Museums in Wien, 104A, 129–143.

    Ladd, H.S. (1966) Chitons and Gastropods (Haliotidae through Adeorbidae) from the western Pacific islands. Geological Survey Professional Paper, 531, 1–98, 16 pls.

    Laghi, G.F. (1977) Polyplacophora (Mollusca) neogenici dell'Appennino settentrionale. Bollettino della Società Paleontologica Italiana, 16, 87–115, pls. 1–4.

    Laghi, G.F. (2005) Upper Triassic chitons from the Italian Dolomites. Lavori della Società Veneziana di Scienze Naturali, 30, 79–84.

    Landau, B. M., Ceulemans, L. & Van Dingenen, F. (2018) The upper Miocene gastropods of northwestern France, 2. Caenogastropoda. Cainozoic Research. [submitted]

    Landau, B. M., Van Dingenen, F. & Ceulemans, L. (2017) The upper Miocene gastropods of Northwest France, 1. Patellogastropoda and Vetigastropoda. Cainozoic Research, 17, 75–166.

    Lauriat-Rage, A. (1981) Les Bivalves du Redonien (Pliocène atlantique de France). Signification stratigraphique et paléobiogéographique.Mémoires du Muséum national d’Histoire naturelle, New Series, Série C, Sciences de la Terre, 45, 1–173.

    Leloup, E. (1941) A propos de quelques Acanthochitons peu connus ou nouveaux. II. Region Atlantique. Bulletin de Musee royal d’Histoire naturelle de Belgique, 17 (43), 1–15, pl. 1.

    Leloup, E. (1968) Acanthochitons de la cote atlantique africaine. Memórias da Junta de Investigações do Ultramar, 54, 55–84.

    Leloup, E. (1980) Chitons de la Mer Rouge, du golfe de Suez et de la Mediterranee. Bulletin Institut Royale des Sciences Naturelles de Belgique, 52 (5), 1–14, 2 pls.

    Leloup, E. (1981) Acanthochiton oblongus sp. nov. Bulletin Institut Royale des sciences Naturelles de Belgique, 53, 1–3, pl. 1.

    Linnaeus, C. (1767) Systema Naturae. Editio duodecima, reformata. “Vermes Testacea”. 1 (2). L. Salvii, Holmiae, 795 pp. [pp. 533–1327]

    Lovén, S.L. (1846) Index molluscorum litora Scandinaviae Occidentalia habitantium. Öfversigt Af K. Svenska Vetenskaps-Akademiens Förhandlingar, 3, 134–160.

    Macioszczyk, W. (1988) Polyplacophora from the Badenian deposits of Weglinek, Weglin and Łichów (Western Roztocze – Poland). Prace Museum Ziemi, 40, 47–58, pls. 1–4.

    Malatesta, A. (1962) Mediterranean Polyplacophora Cenozoic and Recent. Geologica Romana, 1, 145–171.

    Marquet, R. (1984) A remarkable molluscan fauna from the Kattendijk formation (Lower Pliocene) at Kallo (Oost-Vlaanderen, Belgium). Bulletin de la Société belge de Géologie, 93, 335–345.

    Marquet, R. (2002) The Neogene Amphineura and Bivalvia (Protobranchia and Pteriomorphia) from Kallo and Doel (Oost-Vlaanderen, Belgium). Palaeontos, 2, 1–100, pls. 1–34.

    Michelotti, G. (1847) Description des Fossiles des Terrains Miocènes de l'Italie Septentrionale. De Erven Loosjes, Haarlem, 408 pp., 17 pls.

    Millet de la Turtaudière, R.A. (1854) Paléontologie de Maineet-Loire. Cosnier et Lachèse, Angers, 144 pp.

    Montagu, G. (1803) Testacea Britannica or Natural History of British Shells, Marine, Land and Freshwater, Including the Most Minute, Systematically Arranged and Embellished with Figures. White, London, 1 & 2, 606 pp., 16 pls. [pp. i–xxxvii, 1–291 & pp. 292–606, pls. 1–16]

    Monterosato, M.T.A. (1879) Enumerazione e sinonimia delle Conchiglie mediterranee. Monografia dei Chitonidi del Mediterraneo. Giornale di Scienze Naturali ed Economiche, 14, 9–31.

    Néraudeau, D., Barbe, S., Mercier D. & Roman, J. (2003) Signatures paléoclimatiques des échinides marsupiaux du Messinien atlantique à faciès redonien. Annales de Paléontologie, 89, 153–170.
    https://doi.org/10.1016/S0753-3969(03)00023-5

    Nierstrasz, H.F. (1906) Remarks on the Chitonidae. II. Lepidopleurus africanus, nov. sp. Tijdschrift der Nederlandsche Dierkundige Vereeniging, Series 2, 10, 155–158, pl. 3.

    Payraudeau, B.C. (1826) Catalogue Descriptif et Méthodique des Annélides et des Mollusques de l'Ile de Corse. Béchet, Paris, 218 pp., 8 pls.

    Pennant, T. (1777) The British Zoology, 4. Crustacea, Mollusca, Testacea. Benjamin White, London, xviii + 156 pp., 193 pls.

    Pilsbry H.A. (1892–1894) Monograph of Polyplacophora. In: Tryon, G.W., Manual of Conchology. 14 & 15, pp. 1–128, pls. 1–30 (1892); pp. i–xxxiv + 129–350, pls. 31–68 (1893) & pp. 1–64, pls. 1–10 (1893); 65–133, pls. 11–17 (1894). [Academy of Natural Sciences, Philadelphia]

    Plate, L.H. (1898–1901) Die Anatomie und Phylogenie der Chitonen. In: Fauna Chilensis. Zoologische Jahrbücher, Supplement 4 (1), pp. 1–243, pls. 1–12 (1898); 5 (1), 5–216, pls. 2–11 (1899); 5 (2), 281–600, pls. 12–16 (1901).

    Poli, G.S. (1791–1795) Testacea Utriusque Siciliae Eorumque Historia et Anatome. Tomus primus & Tomus secundus. Bodoni ed., Parma, 257 pp., 18 pls. & 234 pp., 20 pls.

    Puchalski, S. (2005) Taphonomy of Chitons (Mollusca, Polyplacophora) and the Chiton Fossil Record. Thesis, Indiana University, Bloomington, Indiana, x + 129 pp.

    Puchalski, S.S., Eernisse, D.J. & Johnson, C.C. (2008) The effect of sampling bias on the fossil record of chitons (Mollusca, Polyplacophora). American Malacological Bulletin, 25, 87–96.
    https://doi.org/10.4003/0740-2783-25.1.87

    Rafinesque, C.S. (1815) Analyse de la nature ou tableau de l’univers et des corps organisés. Rafinesque, Palermo, 223 pp.

    Reuss, A.E. (1860) Die marinen Tertiärschichten Böhmens und ihre Versteinerungen. Sitzungberichte Akademie der Wissenschaften, Mathematisch–Naturwissenschaftliche Klasse, 39, 250–270.

    Risso, A. (1826) Histoire naturelle des principales productions de l’Europe méridionale et principalement de celles des environs de Nice et des Alpes-Maritimes. 4. Mollusques. Levrault, Paris, vii + 439 pp.

    Rochebrune, A.T. de (1881) Diagnoses specierum novarum familiae Chitonidarum. Journal de conchyliologie, 21, 42–46.

    Rochebrune, A.T. de (1882) Monographie des espèces fossiles appartenant à la classe des Polyplaxiphores. Annales des Sciences Géologiques, 14, 1–74, pls. 1–3.

    Rolle, F. (1862) Über einige neue oder wenig gekannte Mollusken-Arten aus Tertiär-Ablagerungen. Sitzungsberichte der K. Akademie der Wissenschaften Mathematisch Naturwissenschaftliche Klasse, Wien, 44, 205–224, pls. 1–2.

    Ruman, A. & Hudáčková N.H. (2015) Middle Miocene chitons (Polyplacophora) from the Slovak part of the Vienna Basin and the Danube Basin (Central Paratethys). Acta Geologica Slovaca, 7, 155–173.

    Sabelli, B. & Taviani, M. (1979) I Polyplacophora del Pleistocene inferiore del Torrente Stirone (Preappennino Parmense, Italia). Bollettino della Società Paleontologica Italiana, 18, 157–161, pl. 1.

    Sacco, F. (1897) I Molluschi dei Terreni Terziarii del Piemonte e della Liguria. Parte XXII. Gasteropoda (fine). Pulmonata. Amphineura (Chitonidae). Scaphopoda (Dentaliidae). Carlo Clausen, Torino, 148 pp., 10 pls.

    Sandberger, F. (1858–1863) Die Conchylien des Mainzer Tertiärbeckens. Kreidel's Verlag, Wiesbaden, (1), 1–40, pls. 1–5 (1858); (2), 41–72, pls. 6–10 (1858); (3), 73–112, pls. 11–15 (1859); (4), 113–152, pls. 16–20 (1860), (5–6), 153–232, pls. 21–30 (1861); (7), 233–270, pls. 31–35 (1862); (8), 271–458 (1863).

    Sars, G.O. (1878) Bidrag til Kundskaben om Norges Arktiske Fauna, 1. Mollusca regionis arcticae norvegiae. A.W. Brogger, Chris-tiania, xiii + [3] + 466 pp., 18 pls.

    Scacchi, A. (1836) Catalogus Conchyliorum Regni Neapolitani Quae Usque Adhuc Reperit A. Scacchi. Typis Filiatae Sebetii, Neapoli, 19 pp., 1 pl.

    Schmidt-Petersen, J., Schwabe, E. & Haszprunar, G. (2015) Acanthochitona pilosa spec. nov., a new species of Acanthochitona Gray, 1821 from the Mediterranean. Spixiana, 38, 11–20.

    Schumacher, C.F. (1817) Essai d’un nouveau système des habita-tions des vers testacés. Schultz, Copenhagen, 287 pp.

    Schwabe, E. (2010) Illustrated summary of chiton terminology. Spixiana, 33, 171–194.

    Segers, W., Swinnen, F. & de Prins, R. (2009) Marine molluscs of Madeira. The living marine molluscs of the Province of Madeira (Madeira and Selvagens Archipelago). Snoeck Publishers, Belgium, 612 pp.

    Seguenza, G. (1873–1877) Studii stratigrafici sulla formazione pliocenica dell'Italia meridionale. Bollettino del Regio Comitato Geologico d'Italia, 4 (1873), 29–30, 30–45, 84–103, 131–153, 213–230, 280–301, 345–357; 5 (1874), 4–15, 68–85, 146–152, 271–283, 332–347; 6 (1875), 18–31, 82–89, 145–153, 203–211, 275–283, 339–345; 7 (1876), 7–15, 91–103, 179–189, 259–271, 355–359; 8 (1877), 7–17, 91–99, 359–367.

    Sigwart, J.D., Andersen, S.B. & Schnetler, K.I. (2006) First record of a chiton from the Palaeocene of Denmark (Polyplacophora: Leptochitonidae) and ist phylogenetic affinities. Journal of Systematic Palaeontology, 5, 123–132.
    https://doi.org/10.1017/S1477201906001982

    Sigwart, J.D., Carey, N. & Orr, P.J. (2014) How subtle are the biases that shape the fidelity of the fossil record? A test using marine molluscs. Palaeogeography, Palaeoclimatology, Palaeoecology, 403, 119–127.
    https://doi.org/10.1016/j.palaeo.2014.02.025

    Sigwart, J.D., Schwabe, E., Saito, H., Samadi, S. & Giribet, G. (2011) Evolution in the deep sea: a combined analysis of the earliest diverging living chitons (Mollusca : Polyplacophora : Lepidopleurida). Invertebrate Systematics, 2010, 24, 560–572.

    Simroth, H. (1894) Die Hautanpassung eines Placophoren an die Luft. Berichte der Naturforschenden Gesellschaft zu Leipzig, 1893/1894, 104–106.

    Sirenko, B.I. (1973) Amphipacific distribution of chitons (Loricata) and their new species in the North‐West Pacific. Of Sea Shore, 1974 (Summer), 59–63. [English translation by Cowan, I.Mc.T. & Smith, A.G.; original printing in Russian, Zoologiceskij Zurnal, 52, 659–667]

    Sirenko, B.I. (2000) Genus Boreochiton G.O. Sars, 1878 (Mollusca, Polyplacophora)—old name, new composition. Ruthenica, 10, 71–72.

    Sirenko, B. (2006) New Outlook on the system of chitons (Mollusca: Polyplacophora). Venus, 65, 27–49.

    Sirenko, B.I. (2013) Four new species and one new genus of Jurassic chitons (Mollusca: Polyplacophora: Lepidopleurida) from the Middle Russian Sea. Proceedings of the Zoological Institute of the Russian Academy of Sciences, 317 (1), 30–44.

    Sirenko, B. (2014) Composition of the genus Hanleya (Mollusca: Polyplacophora: Lepidopleurida), with the description of two new species. Journal of Natural History, 48 (45–48), 2913–2945.
    https://doi.org/10.1080/00222933.2014.963722

    Sirenko, B.I. (2015) Shallow and deep-sea chitons of the genus Leptochiton Gray, 1847 (Mollusca: Polyplacophora: Lepidopleurida) from Peruvian and Chilean waters. Zootaxa, 4033 (2), 151–202.
    https://doi.org/10.11646/zootaxa.4033.2.1

    Sirenko, B., Sigwart, J. & Dell’Angelo, B. (2016) Hanleya hanleyi (Bean in Thorpe, 1844) (Mollusca, Polyplacophora) and the influence of the Gulf Stream System on its distribution. Ruthenica, 26, 57–70.

    Sosso M. & Dell'Angelo B. (2010) I Fossili del Rio Torsero. Cartotecnica Beusi, Prato, 96 pp.

    Sowerby, G.B.II (1832–1840) The Conchological Illustrations, or Coloured Figures of All the Hitherto Unfigured Recent Shells. Parts 38–45 (1833). Unkown publisher, London, 19 pp. [pp. 159–177 (1839–1840)]

    Speiser, D.I., Eernisse, D.J. & Johnsen, S. (2011) A Chiton Uses Aragonite Lenses to Form Images. Current Biology, 21, 665–670.
    https://doi.org/10.1016/j.cub.2011.03.033

    Spengler, L. (1797) Udförlig beskrivelse over det mangeskallede konkylie-slaegt, af Linnaeus kaldet Chiton; met endeel nye Arter og Varieteter. Skrivter af Naturhistorie Selskabet, 4, 62–103, pl. 6.

    Studencka, B. & Dulai, A. (2010) Chitons (Mollusca: Polyplacophora) from the Middle Miocene sandy facies of Ukraine, Central Paratethys. Acta Geologica Polonica, 60, 257–274.

    Studencka, B. & Studencki, W. (1988) Polyplacophora from the Badenian (Middle Miocene) marine sandy facies of the Holy Cross Mts. (Central Poland). Prace Muzeum Ziemi, 40, 37–46, pls. 1–4

    Šulc, J. (1934) Studien über die fossilen Chitonen. I. Die fossilen Chitonen im Neogen des Wiener Beckens und der angrenzenden Gebiete Annalen des Naturhistorischen Museums in Wien, 47, 1–31.

    Thiele, J. (1893) Das Gebiss der Schecken zur Begrundung einer naturlichen Classification. In: Troschel, F.H., Polyplacophora, Lepidoglossa, Schuppenzüngler, Berlin, 2, pp. 353–401, pls. 30–32.

    Thiele, J. (1909) Revision des Systems der Chitonen. Zoologica, 22, 1–132, pls. 1–10.

    Thorpe, C. (1844) British marine conchology. E. Lum-ley, London, lx + 267 pp.

    Urteaga, D., Griffin, M. & Pastorino, G. (2011) Callochiton monteleonensis n.sp., first record of Polyplacophora (Mollusca) from the Neogene of Argentina. Journal of Paleontology, 85 (6), 1181–1187.
    https://doi.org/10.1666/11-041.1

    Van Belle, R.A. & Dell'Angelo, B. (1998) Description of a new species Ischnochiton dolii sp. nov. (Polyplacophora: Ischnochitonidae) from Civitavecchia, Italy. Apex, 13, 77–79.

    Van Dingenen, F., Ceulemans, L. & Landau, B.M. (2014) Euroscaphella nov. gen. (Gastropoda: Volutidae) in the Neogene of Europe, with the description of a new species: Euroscaphella namnetensis nov. sp. from the Mio–Pliocene transition of northwestern France. Cainozoic Research, 14 (2), 101–111.

    Van Dingenen, F., Ceulemans, L., Landau, B.M. & Silva, C.M. da (2015) The family Nassariidae (Gastropoda: Buccinoidea) from the late Neogene of northwestern France. Cainozoic Research, 15 (1–2), 75–122.

    Van Dingenen, F., Ceulemans, L. & Landau, B.M. (2016) The lower Pliocene gastropods of Le Pigeon Blanc (Loire-Atlantique, north west France), 2. Caenogastropoda. Cainozoic Research, 16 (2), 109–219.

    Van Dingenen, F., Ceulemans, L. & Landau, B.M. (2017) The lower Pliocene gastropods of Le Pigeon Blanc (Loire-Atlantique, north west France), 4. Neogastropoda (in part). Cainozoic Research, 17 (1), 23–61.

    Vendrasco, M.J. (1999) Early evolution of the Polyplacophora (chitons). Thesis, University of California, Los Angeles, 232 pp.

    Vendrasco, M.J., Eernisse, D.J., Powell, C.L.II & Fernandez, C.Z. (2012) Polyplacophora (Mollusca) from the San Diego Formation: A Remarkable Assemblage of Fossil Chitons from the Pliocene of Southern California. Contributions in Science, 520, 15–72.

    Verbeek, R.D.M., Boettger, O. & Fritsch, K. von (1882) Die Conchylien der Mittelmiocaen–Schichten Sud–Sumatra's. In: Die Tertiärformation von Sumatra und ihre Thierreste. Palaeontographica, III (Supplement), II Theil, pp. 34–102, pls. 1–8.