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Type: Article
Published: 2018-09-17
Page range: 1–51
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A revision of the Afrotropical species of Pachylophus Loew (Diptera: Chloropidae) and some related Ovoviviparous genera

Hon. Research Fellow, National Museum of Wales, Cathays Park, Cardiff CF10 3NP, U.K.
Diptera Chloropidae taxonomy Pachylophus Platycephala Phyladelphus Eurina Sabeurina ovoviviparity biology

Abstract

In Pachylophus all described Afrotropical species are treated and P. aristalis, ismayi, nigeriensis, tschirnhausi, mirabilis, lacteibasis and stuckenbergi are described as new species. Pachylophus tellinii Bezzi is placed in synonymy. In Phyladelphus the type species thalhammeri and the single described Afrotropical species, P. geminus, are treated and P. woodi and P. maraisi are described as new species. The genus Sabeurina is erected to accommodate Eurina minuta Loew and a new Afrotropical species S. occidentalis. Some taxonomic notes are given on the Afrotropical species currently included in Platycephala. The immature stages and biological notes concerning the treated genera are described and discussed in the contexts of ovoviviparity and classification. Keys are provided for the identification of all the Afrotropical species known in each of the genera treated.

 

References

  1. Adams, C.F. (1905) Diptera africana, 1. Kansas University Science Bulletin Lawrence, New Series, 3, 149–208.
    https://doi.org/10.5962/bhl.part.8100

    Andersson, H. (1977) Taxonomic and phylogenetic studies on Chloropidae (Diptera) with special reference to Old World genera. In: Entomologica scandinavica. 8 (Supplement). Scandinavian Science Press, Klampenborg, pp. 1–200.

    Andersson, H. (1979) Problem vid kladistisk analys av flugfamiljen Chloropidae. Entomologisk Tidskrift, 100 (3–4), 180–187.

    Becker, T. (1907) Die Ergebnisse meiner dipterologischen Frühjahrsreise nach Algier und Tunis, 1906. Zeitschrift für systematische Hymenopterologie und Dipterologie, 7, pp. 33–61 + 97–128 + 225–256 + 369–407 + 454–455.

    Becker, T. (1910) Chloropidae. Eine monographische Studie. 2. Teil. Aethiopische Region. Annales historico-naturales Musei nationalis hungarici Budapest, 8, 377–443, tab. XIII.

    Becker, T. (1913) Chloropiden aus Abessynien gesammelt von E. Kovács. Annales historico-nationalis Musei nationalis hungarici Budapest, 11, 147–167.

    Becker, T. (1914) Diptères nouveaux récoltés par MM. Ch. Alluaud et R. Jeannel en Afrique orientale 1911–1912. Annales de la Societé Entomologique de France, Paris, 83 (1913–1914), 120–130.

    Becker, T. (1916) Neue Chloropiden aus dem Ungarischen National Museum. Annales historico-naturales Musei nationalis Hungarici, 14 (42), 423–453.

    Beschovski, V. (1983) On the systematic status of Steleocerus lindbergi (Duda, 1933) (Diptera, Chloropidae). Acta zoologica bulgarica, 21, 78–82. [Zoological Record translation of title in Bulgarian]

    Bezzi, M. (1908) Ditteri eritrei raccolti dal Dott. Andreini e dal Prof. Tellini. Parte seconda. Diptera cyclorrhapha. Bollettino della Società Entomologica Italiana, Firenze, 39, (1907), 3–199.

    Collin, J.E. (1949) Results of the Armstrong College Expedition to Siwa Oasis (Libyan Desert), 1935 under the leadership of Prof. J. Omer-Cooper. Diptera Empididae, Dolichopodidae, Aschiza and Acalypterae. Bulletin de la Société Fouad 1er d’Entomologie, Le Caire, 33, 175–225.

    Cumming, J.M. & Wood, M. (2009) Chapter 2. Adult morphology and terminology. In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, M.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NRC Research Press, Ottawa, Ontario, pp. 9–50.

    Curran, C. H. (1928) Diptera of the American Museum Congo Expedition. Part II—Asilidae, Conopidae, Pyrgotidae, Micropezidae, Chloropidae, Drosophilidae, Lonchaeidae, Sapromyzidae, Muscidae, Calliphoridae and Tachinidae. Bulletin of the American Museum of Natural History, 57 (6), 327–399.

    Deeming, J.C. (1974) Notes on immature stages of Pachylophus beckeri Curran (Dipt. Chloropidae). Entomologist's Monthly Magazine, 109 (1973), 155–157.

    Deeming, J.C. (1977) The immature stages of Elachiptereicus abessynicus Becker and Pachylophus lugens Loew (Diptera: Chloropidae). Bulletin of Entomological Research, 67 (2), 325–328.
    https://doi.org/10.1017/S0007485300011147

    Deeming, J.C. (1981) A contribution to the study of afrotropical Chloropine Diptera. Journal of Natural History, 15 (5), 789–828.
    https://doi.org/10.1080/00222938100770581

    Deeming, J.C. (1985) A description of the adult and immature stages of Ectecephala capillata (Coquillett) (Diptera: Chloropidae). Proceedings of the Entomological Society of Washington, 1987 (2), 402–407.

    Deeming, J.C. (2011) Order Diptera, family Chloropidae. In: van Harten, A. (Ed.), Arthropod Fauna of the United Arab Emirates. Vol 4. Dar Al Ummah, Abu Dhabi, pp. 784–806.

    Duda, O. (1932–1933) 61. Chloropidae. In: Lindner, E. (Ed.), Die Fliegen der Paläarktischen Region. Band 6 (1). Schweitzebart, Stuttgart, pp. I–IV + 1–248, taf. I–III.

    Ebejer, M.J. (2009) Order Diptera, Family Scenopinidae. In: van Harten, A. (Ed.), Arthropod Fauna of the United Arab Emirates. Vol 2. Dar Al Ummah, Abu Dhabi, pp. 741–749

    Elzinga, R.J. & Broce, A.B. (1986) Labellar modifications of Muscomorpha flies (Diptera). Annals of the Entomological Society of America, 79 (1), 150–209.
    https://doi.org/10.1093/aesa/79.1.150

    Ferrar, P. (1987) A Guide to the Breeding Habits and Immature Stages of Diptera Cyclorrhapha. 18. Family Chloropidae. In: Entomonograph. Vol. 8. Parts 1 & 2. E.J. Brill/Scandinavian Science Press, Leiden, pp. 106–120.

    Gould, S.J. (1977) Ontogeny and Phylogeny. Belknap Press of Harvard University Press, Cambridge, Massachusetts, xiii + 501 pp.

    Johnson, C.W. (1898) Diptera collected by Dr A. Donaldson Smith in Somaliland, eastern Africa. Proceedings of the Academy of Natural Sciences of Philadelphia, 1898, 157–164.

    Kaplan, F. (1977) The Chloropidae of Israel. MSc Thesis, Tel-Aviv University, Department of Zoology, II +130 pp., 21 pls. [in Hebrew with English summary]

    Lamb, C.G. (1917) Notes on exotic Chloropidae. Annals and Magazine of Natural History, London, Series 8, 19, 33–58.
    https://doi.org/10.1080/00222931709486911

    Loew, H. (1858) Ueber einige neue Fliegengattungen. Berliner entomologische Zeitschrift, 2, 101–122, taf. 1.

    Loew, H. (1860) Bidrag till kännedomen om Afrikas Diptera (part). Ofversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, Stockholm, 17 (2), 81–97.

    Meier, R., Kotrba, M. & Ferrar, P. (1999) Ovoviviparity and viviparity in the Diptera. Biological Reviews of the Cambridge Philosophical Society, 74 (3), 199–258.
    https://doi.org/10.1111/j.1469-185X.1999.tb00186.x

    Moyal, P. (1982) Deux foreurs du riz irrigué noveaux pour la Côte d'Ivoire, Hydrellia prosternalis Deeming (Diptera, Ephydridae) et Pachylophus beckeri Curran (Diptera, Chloropidae). Agronomie tropicale, 37 (3), 295–305.

    Nartshuk, E.P. (1962) Results of the Chinese-Soviet Zoological-Botanical Expeditions to South-Westerh [sic] China. Chloropidae (Diptera) of Yunnan and Kwandun Provinces China. Revue d'Entomologie de l'URSS, 41 (3), 672–684. [in Russian, English translation in Entomological Review, 41 (3), 417–424 (1962)]

    Nartshuk, E.P. (1964) A new genus and new species of Chloropidae (Diptera) from Kazakhstan. [translation of Russian title as cited in Zoological Record]. Trudy Zoologicheskogo Instituta. Academiya Nauk SSSR, Leningrad, 34, 302–324.

    Nartshuk, E.P. (1973) New species of Chloropidae (Diptera) of the Palaearctic fauna. [translation of Russian title]. Entomologicheskoe Obozrenie, Moskva, 52 (1), 215–227.

    Nartshuk, E.P. (1983) A system of the superfamily Chloropoidea (Diptera, Cyclorrhapha). Entomologicheskoe Obozrenie, 62 (3), 638–648. [in Russian, English translation in Entomological Review, 62 (3), 180–193 (1984)]

    Nartshuk, E.P. (1993) “News [i.e. new data] on the systematics and faunistics of the insects of Vietnam. Part 2. Flies of the family Chloropidae (Diptera) from Vietnam and South China”. [title translation from Russian]. Proceedings of the Zoological Institute of the Academy of Sciences, USSR, 240 (1991), 77–120.

    Nartshuk, E.P. (1997) Biodiversity of Chloropidae (Diptera) in tropical ecosystems, pp. 169–178. In: Ulrich, H. (Ed.), Tropical Biodiversity and Systematics. Proceedings of the International Symposium on Biodiversity and Systematics in Tropical Ecosystems, 2–7 May 1994. Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn, 357 pp.

    Nartshuk, E.P. (2014) Grass-fly Larvae (Diptera, Chloropidae): Diversity, Habitats, and Feeding Specializations. In: Entomological Review. 94 (4). Buda, Texas, pp. 514–525. [originally published in Zoologicheskii Zhurnal, 93 (1), 81–91 (2014, in Russian)]

    Pollock, J.N. (1996) Viviparous adaptations in the acalyptrate genera Pachylophus (Chloropidae) and Cyrtona (Curtonotidae) (Diptera: Schizophora). Annals of the Natal Museum, 37, 183–189.

    Roth, L.M. & Willis, E.R (1954) The reproduction of cockroaches. Smithsonian miscellaneous collections, 122 (12), 1–49.

    Sabrosky, C.W. (1945) Three new African Chloropidae, with a discussion of the status of Pseudogaurax (Diptera). Proceedings of the Zoological Society of London, 114 (4), 456–461.
    https://doi.org/10.1111/j.1096-3642.1945.tb00237.x

    Sabrosky, C.W. (1959) Chapter XXII. Diptera (Brachycera): Chloropidae. In: Hanström, B., Brink, P. & Rudebeck, G. (Eds.), Results of the Lund University Expedition in 1950–1951. South African Animal Life. Vol. 6. Almqvist & Wiksell, Stockholm, pp. 413–424.

    Sabrosky, C.W. (1980) 80. Family Chloropidae. In: Crosskey, R.W., Cogan, B.H., Freeman, P., Pont, A.C., Smith, K.G.V. & Oldroyd, H. (Eds.), Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London, pp. 695–712.

    Skidmore, P. (1985) The biology of the Muscidae of the world. Series entomologica 29. Springer, Dordrecht, xiv + 550 pp.

    Speiser, P. (1910) [Abteilung] 10. Diptera. 5. Cyclorhapha [sic!], Aschiza. Cyclorhapha, Schizophora, Schizometopa. (Auswahl); Holometopa. (Auswahl). In: Königlich schwedische Akademie der Wissenschaften (Ed.), Wissenschaftliche Ergebnisse der schwedischen zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas 1905–1906 unter Leitung von Prof. Dr. Yngve Sjöstedt. 2. Band, Abteilung 8–14. P. Palmquists Aktiebolag, Stockholm, pp. 113–202 incl. index.

    Spencer, K.A. (1985) Viviparity in the Chloropidae (Diptera). Entomologist's Monthly Magazine, 121, 179–180.

    Spencer, K.A. (1986) The Australian Chloropinae (Diptera: Chloropidae). Journal of Natural History, 20 (3), 503–615.
    https://doi.org/10.1080/00222938600770401

    Thomasson, J.R. (1986) 15. Fossil Grasses: 1820–1986 and Beyond. In: Sonderstrom, T.R., Hilu, K.W., Campbell, C.S. & Barkworth, M.E. (Eds.), Grass Systematics and Evolution (an international symposium held at the Smithsonian Institution. 27–31 July 1986, Washington, D.C. Smithsonian Institution Press, Washington, pp. 1–474.

    von Tschirnhaus, M. (1992) Minier- und Halmfliegen (Agromyzidae, Chloropidae) und 52 weitere Familien (Diptera) aus Malaise-Fallen in Kiesgruben und einem Vorstadtgarten in Köln. Decheniana-Beihefte, 31, 445–497, taf. 13–15.

    von Tschirnhaus, M. & Hoffeins, C. (2009) Fossil flies in Baltic amber—insights in the diversity of Tertiary Acalyptratae (Diptera, Schizophora), with new morphological characters and a key based on 1,000 collected inclusions. Denisia, 26, 171–212.

    Venturi, F. (1940) Contributi alla conoscenza dell’Entomofauna delle Graminacee, VI. Redia, 26, 27–70.

    Yang, D. & Yang, C.-K. (1990) New and little-known species of Chloropidae from Guizhou (Ⅲ) (Diptera: Acalyptratae). Guizhou kexue (= Guizhou Science), 10 (2), 53–55.

    Yang, D. & Yang, C. (1994) New species of Chloropidae from Yunnan (Diptera: Acalyptratae). Entomotaxonomia, Wukung, 16 (1), 75–78. [in Chinese with English summary]