Abstract
The fauna of quill mites of the family Syringophilidae (Acari: Prostigmata: Cheyletoidea) of the Palaearctic Region is comprehensively revised. The external morphology of adults and immature instars was studied in detail. All known 118 species grouped in 32 genera and 2 subfamilies occuring in the Palaearctic Region are examined and keyed. Critical reviews of host associations, occupying habitat, and distribution are given for all mite species. Six new genera are proposed, 4 in the subfamily Syringophilinae: Betasyringophiloidus gen. nov., Krantziaulonastus gen. nov., Paraniglarobia gen. nov., Pteroclidisyringophilus gen. nov.; and 2 in the subfamily Picobiinae: Neopicobia gen. nov., and Rafapicobia gen. nov. Twenty five new species are described in the subfamily Syringophilinae: Aulobia leucostictus sp. nov., Aulonastus lanius sp. nov., A. loxius sp. nov., A. anthus sp. nov., A. fringillus sp. nov., Betasyringophiloidus saxicolus sp. nov., B. phoenicurus sp. nov., Krantziaulonastus oryzivorus sp. nov., K. lonchurus sp. nov., Meitingsunes columbicus sp. nov., Neoaulonastus remizus sp. nov., N. caligatus sp. nov., N. aegithalos sp. nov., N. picidus sp. nov., N. riparius sp. nov., Selenonycha chradriiformicus sp. nov., Syringophiloidus philomelosus sp. nov., S. petronicus sp. nov., S. parapresentalis sp. nov., S. coccothraustes sp. nov., Syringophilopsis idunae sp. nov., S. muscicapus sp. nov., S. passericus sp. nov., S. nucifragus sp. nov., Torotrogla pycnonotus sp. nov. and five new species are described in the subfamily Picobiinae: Picobia cetti sp. nov., P. riparius sp. nov., P. eremophila sp. nov., P. galerida sp. nov., and Rafapicobia zirnitra sp. nov. The following new combinations are proposed: Aulobia stachyris (Bochkov et al., 2000) comb. nov., Betasyringophiloidus motacillae (Bochkov and Mironov, 1998) comb. nov., B. schoeniclus (Skoracki, 2002) comb. nov., Columbiphilus alectoris (Fain et al., 2000) comb. nov., Columbiphilus polonica (Skoracki et al., 2001) comb. nov., C. pteroclesi (Skoracki and OConnor, 2010) comb. nov., Cuculisyringophilus chirovi (Bochkov and Mironov, 1998) comb. nov., Krantziaulonastus buczekae (Skoracki, 2002) comb. nov., Krantziaulonastus galbulicus (Skoracki, 2008) comb. nov., Neoaulonastus bisetatus (Fritsch, 1958) comb. nov., Neopicobia anthi (Fritsch, 1958) comb. nov., Neopicobia cardinalis (Skoracki et al., 2010) comb. nov., N. carpodacus (Skoracki et al., 2010) comb. nov., N. epthianura (Skoracki et al., 2008) comb. nov., N. glossopsitta (Skoracki et al., 2008) comb. nov., N. locustella (Skoracki et al., 2004) comb. nov., N. modularis (Skoracki and Magowski, 2001) comb. nov., N. pyrrholaemus (Skoracki and Glowska, 2008) comb. nov., N. troglodytes (Skoracki et al., 2010) comb. nov., N. zumpti (Lawrence, 1959) comb. nov., Paraniglarobia calidridis (Bochkov and Mironov, 1998) comb. nov., Paraniglarobia skorackii (Bochkov and Galloway, 2004) comb. nov., Pteroclidisyringophilus re (Skoracki and OConnor, 2010) comb. nov. Three species are considered as incertae sedis: Picobia brotogeris Fain et al., 2000, P. leucophaeus Skoracki et al., 2010, P. poicephali Skoracki and Dabert, 2002. Keys to the syringophilid genera and all Palaearctic species are provided. Main aspects of host–parasite relationships are discussed.
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Skoracki, M., Hendricks, S. & Spicer, G.S. (2010c) New species of parasitic quill mites of the genus Picobia (Acari: Syringophilidae: Picobiinae) from North American birds. Journal of Medical Entomology, 47, 727–742.
Skoracki, M., Hromada, M. & Kuczynski, L. (2001a) Torotrogla lullulae, a new species of the family Syringophilidae Lavoipierre, 1953 (Acari: Prostigmata: Cheyletoidea). Genus, 12, 87–92.
Skoracki, M., Hromada, M. & Tryjanowski, P. (2001b) Description of a new species of quill mite Syringophiloidus weiszii sp. n. (Acari, Prostigmata, Syringophilidae) from Great Grey Shrike Lanius excubitor. Acta Parasitologica, 46, 30–34.
Skoracki, M. & Kiljan, G. (2002) Picobia paludicola sp. n. a new species of quill mite (Acari: Prostigmata: Syringophilidae) from the Aquatic Warbler Acrocephalus paludicola (Passeriformes: Sylviidae). Zootaxa, 91, 1–6.
Skoracki, M., Lontkowski, J. & Stawarczyk, T. (2010d) New taxa of the parasitic quill mites associated with accipitrid birds indicating close relationship of falconid birds to Psittaci-Columbi clade. Journal of Natural History, 44, 1203–1214.
Skoracki, M. & Magowski, W. (2001) Two new species of quill mites of the genus Picobia (Acari: Prostigmata: Syringophilidae) from passerine birds of Poland. Acarina, 9, 113–119.
Skoracki, M., Magowski, W. & Dabert, J. (2001c) Picobia polonica sp. n. (Acari: Prostigmata: Syringophilidae), a new species of quill mite from the domestic hen, Gallus gallus domesticus (Aves: Phasianidae). Folia Parasitologica, 48, 154–158.
Skoracki, M., Michalik, J., Skotarczak, B., Rymaszewska, A., Sikora, B., Hofman, T., Wodecka, B. & Sawczuk M. (2006) First detection of Anaplasma phagocytophilum in quill mites (Acari: Syringophilidae) parasitizing passerine birds in Poland. Microbes and Infection, 8, 303–307.
Skoracki, M., Michalik, J. & Sikora, B. (2005) Habitat selection among the quill mites (Acari: Syringophilidae. In Arthropods, a variety of forms and interactions; Buczek, A. & Blaszak, C. (eds.). Liber, Lublin, Poland, 99–105pp.
Skoracki, M., Michalik, J. & Sikora, B. (2010e) Prevalence and habitat preference of quill mites (Acari, Syringophilidae) parasitizing forest passerine birds in Poland. Acta Parasitologica, 55, 188–193.
Skoracki, M., Møller, A.P. & Tryjanowski, P. (2003) A new species of parasitic mites of the genus Syringophiloidus Kethley, 1970 (Acari: Syringophilidae) from the Barn Swallow Hirundo rustica Linnaeus, 1758. Parasite, 10, 17–20.
Skoracki, M. & OConnor, B. (2010) New taxa of quill mites (Acari: Cheyletoidea: Syringophilidae). Zootaxa, 2341, 1–32.
Skoracki, M. & Sikora, B. (2002) New ectoparasitic mites of the family Syringophilidae (Acari: Prostigmata: Cheyletoidea) associated with birds from Argentina. Zootaxa, 27, 1–8.
Skoracki, M. & Sikora, B. (2003) Quill mites (Acari: Prostigmata: Syringophilidae) from African passeriform birds. Zootaxa, 129, 1–10.
Skoracki, M. & Sikora, B. (2004a) Tinamiphilopsis elegans gen. nov. et sp. nov., a first record of the quill mites (Acari, Syringophilidae) from tinamou birds (Tinamiformes, Tinamidae). Acta Parasitologica, 49, 348–352.
Skoracki, M. & Sikora, B. (2004b) A new genus and four new species of quill mites (Acari: Prostigmata: Syringophilidae) from phasianid birds (Galliformes: Phasianidae). Parasite, 11, 379–386.
Skoracki, M. & Sikora, B. (2005) Neosyringophilopsis, a new genus of the subfamily Syringophilinae (Acari: Syringophilidae). Zootaxa, 1052, 21–28.
Skoracki, M. & Sikora, B. (2008) Blaszakia, a new genus of quill mites (Acari: Syringophilidae) parasitizing Turacos (Aves: Musophagiformes). Annales Zoologici, 58, 327–332.
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Skoracki, M., Tryjanowski, P. & Hromada, M. (2002) Two new species of the genus Syringophilopsis Kethley, 1970 (Acari: Syringophilidae) parasitizing quills of true shrikes (Aves: Laniidae). Parasite, 9, 11–16.
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