Abstract
New specimens of trilobites from the Lower Permian limestone reef Shakhtau (Sterlitamak, Bashkortostan, Russia) are described as Brachymetopus (Conimetopus) alekseevi Mychko, sp. nov. Although represented by a limited number of specimens, the new species is significant in extending the stratigraphic distribution of the subgenus Brachymetopus (Conimetopus) Hahn and Hahn, 1985, whose members were previously known exclusively from Carboniferous deposits. Material described previously from the Early Permian (Asselian) of the Chelyabinsk Region is also assigned to the new species.
References
Adrain, J.M. (2011) Class Trilobita Walch, 1771. In: Zhang, Z.-Q. (Ed.), Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148, pp. 104–109.
Amler, M.R.W. & Herbig, H.-G. (2006) Ostrand der Kohlenkalk-Plattform und Übergang in das Kulm-Becken im westlichsten Deutschland zwischen Aachen und Wuppertal. In: Amler, M.R.W. & Stoppel, D. (Eds.), Deutsche Stratigraphische Kommission. Stratigraphie von Deutschland 6: Unterkarbon (Mississippium). Schriftenreihe der Deutschen Gesellschaft für Geowissenschaften, 41, pp. 441–477. [in German]
Angelin, N.P. (1854) Palaeontologia Scandinavica Pars I, Iconographia crustaceorum formationis transitionis, Fasciculus II. T.O. Weigel, Lund, 72 pp. [pp. 21–92]
Archbold, N.W. (1981) New Permian trilobite from Irian Jaya, Indonesia. Geological Research and Development Centre. Paleontology Series, 2, 35–41.
Chernov, A.A. & Chernov, G.A. (1940) Geologicheskoye stroyeniye basseyna r. Kos'yu v Pechorskom kraye (otchet geologicheskogo otryada Pechorskoy kompleksnoy ekspeditsii SOPS Akademii nauk 1935 g.). AS USSR, Moscow-Leningrad, 103 pp. [in Russian]
Chernykh, V.V. (2006) Nizhnepermskie konodonty Urala. Institut geologii i geokhimii UrO RAN, Ekaterinburg, 130 pp. [in Russian]
Chuvashov, B.I. (1983) Permian reefs of the Urals. Facies, 8, 191–212.
Chuvashov, B.I., Dyupina, G.V., Mizens, G.A. & Chernykh, V.V. (1991) Opornyye razrezy karbona i nizhney permi zapadnogo sklona Urala. UrO US USSR, Sverdlovsk, 369 pp. [in Russian]
Diener, C. (1897) Himalayan fossils. The Permocarboniferous fauna of Chitichun No.1. Palaeontologia Indica, 15, 1 (13), 1–105.
Eichwald, E. (1861) Lethaea Rossica ou Paléontologie de la Russie. Ancienne Période. 3. Librairie et imprimerie de E. Schweizerbart, Stuttgart, 1657 pp.
Gandl, J. (1987) Die Karbon-Trilobiten des Kantabrischen Gebirges (NW-Spanien) 4: Trilobiten aus dem hoheren Namur und tieferen Westfal. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 543, 1–179. [in German]
Gandl, J. (2011) The Carboniferous trilobites of the Cantabrian Mountains (NW Spain), 5: upper Westphalian trilobites. Abhandlungen der Senckenberg Gesellschaft fuer Naturforschung, 569, 1–143. [in German]
Gerasimov, N.P. (1934) Nekotoryye zamechaniya k stat'ye prof. M.E. Noinskogo Shvagerinovyy gorizont i artinskiye otlozheniya na Urale». Uchenye zapiski Kazanskogo osudarstvennogo universiteta imeni V.I. Ul'yanova-Lenina, 94 (1/3), 38–67. [in Russian]
Hahn, G. & Hahn, R. (1970) Trilobiten aus dem Kohlenkalk von Sondern (Rheinland). Decheniana, 122 (2), 217–250. [in German]
Hahn, G. & Hahn, R. (1982) Die Trilobiten des belgisehen Kohlenkalkes (Unter-Karbon). 2. Brachymetopus. Geologica et Palaeontologica, 15, 89–114. [in German]
Hahn, G. & Hahn, R. (1985) Trilobiten aus dem hohen Ober-Karbon oder Unter-Perm von Alaska. Senckenbergiana lethaea, 66 (6), 445–485. [in German]
Hahn, G. & Hahn, R. (1988) Seltene Trilobiten aus dem Unter-Aprathium (Unter-Karbon) von Erdbach und Herborn (Hessen). Senckenbergiana lethaea, 68 (5/6), 347–370. [in German]
Hahn, G. & Hahn, R. (1992) Trilobiten aus dem Karbon von SE-Alaska, Teil 2. Geologica et Palaeontologica, 26, 99–133. [in German]
Hahn, G. & Hahn, R. (1996) Die trilobiten-taxa des Karbons und Perms 2. Brachymetopidae. Courier Forschungsinstitut Senckenberg, 195, 1–242. [in German]
Hahn, G., Hahn, R. & Yuan, J.L. (1989) Trilobites from the Upper Carboniferous (Westphalian A) of S-China (N-Guangxi). Geologica et Palaeontologica, 23, 113–203.
Jell, P.A. & Adrain, J.M. (2003) Available generic names for trilobites. Memoirs of the Queensland Museum, 48 (2), 331–553.
Kobayashi, T. & Hamada, T. (1979) Permo-Carboniferous trilobites from Thailand and Malaysia. Geology and Palaeontology of Southeast Asia, 20, 1–21.
Korolyuk, I.K. (1985) Metody i rezultaty izucheniya permskogo rifogennogo massiva Shakhtau (Bashkirskoye Priuralye). Nauka, Moscow, 120 pp. [in Russian]
Kotlyar, G.V., Zakharov, Y.D., Kochirkevich, B.V., Kropacheva, G.S., Rostovcev, K.O., Chedia, I.O., Vuks, G.P. & Guseva, E.A. (1983) Pozdnepermskiy etap evolyutsii organicheskogo mira (Dzhul'finskiy i dorashamskiy yarusy SSSR). Nauka, Leningrad, 200 pp. [in Russian]
Krasnopolsky, A.A. (1887) Kratkiy ocherk geologicheskogo stroyeniya Kynovskoy dachi na Urale. Izvestia Geologicheskogo kommiteta, 6, 267–303. [in Russian]
Krasnopolsky, A.A. (1891) Obshchaya geologicheskaya karta Rossii. List 126-y. Perm–Solikamsk. Trudy Geologicheskogo kommiteta, 11 (2), 30. [in Russian]
Kulagina, E.I., Skuin, I.A. & Kossovaya, O.L. (2015) Permskiy rif Shakhtau. Belaya reka, Ufa, 72 pp. [in Russian]
Kulik, E.A. (1978) Izvestkovyye zelenyye (sifonovyye) vodorosli assel'skogo i sakmarskogo vozrasta massiva Shakhtau (Bashkiriya). Voprosy mikropaleontologii, 21, 182–214. [in Russian]
McCoy, F. (1847) On the fossil botany and zoology of the rocks associated with the coal of Australia. Annals and Magazine of Natural History, Series 1, 20, 226–236.
https://doi.org/10.1080/037454809496037
Miloradovich, B.V. & Ilina, N.S. (1951) Litolgo-paleontologicheskaya kharakteristika kamennougol'nykh i nizhnepermskikh otlozheniy Priural'ya. Gostoptekhizdat, Moscow-Leningrad, 106 pp. [in Russian]
Murchison, R.I., de Verneuil, E. & von Keyserling, A. (1845) The Geology of Russia in Europe and the Ural Mountains. Vol. 1. Geology. Murray, London, 700 pp.
https://doi.org/10.5962/bhl.title.121912
Mychko, E.V. (2012) A New Species of Trilobites from Sakmarian Reefs of Bashkortostan. Paleontological Journal, 46 (1), 44–48.
https://doi.org/10.1134/S0031030112010108
Mychko, E.V. (2016) Trilobity srednego-verkhnego karbona i permi Severnoy Yevrazii. Dissertation of the candidate of geologo-mineralogical sciences. Lomonosov Moscow State University, Moscow, 393 pp. [in Russian]
Mychko, E.V. & Alekseev, A.S. (2017) Localities of middle Carboniferous—Permian trilobites in Russia and surrounding countries. Bulletin of Moscow Society of Naturalists. Geological Series, 92 (3), 40–83. [in Russian]
Möller, V. von (1867) Über die Trilobiten der Steinkohlenformation des Urals, nebst einer Uebersicht und einigen Ergänzungen der bisherigen Beobachtungen uber Kohlen Trilobiten im Allgemeinen. Bulletin of the Imperial Society of Naturalists of Moscow, 1, 1–81. [in German]
Möller, V.I. von (1868) O trilobitakh kamennougol'noy formatsii Urala, s obzorom i nekotorymi dopolneniyami predshestvuyushchikh nablyudeniy nad kamennougol'nymi trilobitami voobshche. Zapiski Imperatorskogo S.-Peterburgskogo mineralogicheskogo obschestva, 2, 3 (7), 5–72. [in Russian]
Osmólska, H. (1968) Brachymetopus McCoy (Trilobita) in the Carboniferous of Poland and U.S.S.R. Acta Palaeontologica Polonica, 13 (3), 359–374.
Owens, R.M. (1983) A review of Permian trilobite genera. Special Papers in Palaeontology, 30, 15–41.
Owens, R.M. (1986) The Carboniferous trilobites of Britain. Part 1. Monograph of the Palaeontographical Society, 138, 1–26.
Owens, R.M. (2003) The stratigraphical distribution and extinctions of Permian trilobites. Special Papers in Palaeontology, 70, 377–397.
Özdikmen, H. (2009) Nomenclatural changes for twenty trilobites genera. Munis Entomology and Zoology, 4 (1), 155–171.
Puchkov, V.N. (2010) Geologiya Urala i Priuralya (aktualnye voprosy stratigrafii, tektoniki, geodinamiki i metallogenii). DizaynPoligrafServis, Ufa, 280 pp. [in Russian]
Prantl, F. & Pribyl, A. (1950) Revise čeledi Otarionidae R. a E. Richter z českého siluru a devonu (Trilobitae). Sborník Státního Geologického Ústavu Čeckoslovenské, 17, 1–83. [in Czech]
Rauzer-Chernousova, D.M., Beljaev, G. & Reitlinger, E. (1936) Upper Paleozoic foraminifera of the Pechora region. Proceedings of the Polar Commission AS SSSR, 28, 159–232. [in Russian]
Rauzer-Chernousova, D.M. (1950) Fatsii verkhnekamennougolnykh i artinskikh otlozheniy Sterlitamaksko-Ishimbayskogo Priuralya (na osnove izucheniya fuzulinid). Trudy Instituta geologii AN SSSR, 119 (43), 1–109. [in Russian]
Rauzer-Chernousova, D.M., Ivanova, E.A. & Korolyuk, I.K. (1977) K kharakteristike stratotipa sterlitamakskogo gorizonta (nizhnyaya perm, massiv Shakhtau, Bashkiriya). Byulluten MOIP. Otdelenie geologii, 52 (6), 24–37. [in Russian]
Rauzer-Chernousova, D.M. & Korolyuk, I.K. (1991) Sterlitamakskie shikhany—rannepermskie rify. In: Mezhdunarodnyy kongress «Permskaya sistema Zemnogo shara». Putevoditel geologicheskikh ekskursiy. Chast II. Yuzhnouralskaya ekskursiya. Sverdlovsk, pp. 47–71. [in Russian]
Savage, N.M. & Barkeley, S.J. (1985) Early to Middle Pennsylvanian conodonts from the Klawak Formation and the Ladrones Limestone, Southeastern Alaska. Journal of Paleontology, 59 (6), 1451–1475.
Shamov, D.F. (1957) Fatsii sakmaro-artinskikh otlozheniy Ishimbaevskogo Priuralya. Trudy Ufimskogo neftyanogo nauchno issledovatelskogo instituta, 2, 3–77. [in Russian]
Shamov, D.F. (1984) Vtoroy den. Razrez permi gor-odinochek Shakhtau i Tratau. In: Einor, O.L. (Ed.), 27 Mezhdunarodnyy geologicheskiy kongress. Yuzhnyy Ural. Putevoditel ekskursii 047. Nauka, Moscow, pp. 22–25. [in Russian]
Tolstikhina, M.M. (1941) Karbon i nizhnyaya perm' vostoka Russkoy platformy i zapadnogo sklona Urala. In: Trudy Bashkirskogo geologicheskogo upravleniya. Vol. 11. Gosgeolizdat Moscow-Leningrad, pp. 1–177. [in Russian]
Verneuil, E. de (1845) Terrain Paléozoique. Animaux. In: Murchison, R.I., de Verneuil E. & Keyserling, A. (Eds.), Géologie de la Russie d’Europe et des montagnes de l’Oural. Vol. II. Troisème partie. Paléontologie. J. Murray, London, Bertrand, Paris, pp. 15–396.
https://doi.org/10.5962/bhl.title.121912
Weber, V.N. (1932) Trilobity Turkestana. Trudy Vsesoyuznogo Geologo-Razvedochnogo Ob’edineniya N.K.T.P—S.S.S.R, 178, 1–157. [in Russian]
Weber, V.N. (1937) Trilobity kamennougolnych i permskich otlozenij SSSR. Vypusk 1. Kamennougolnye trilobity. Monografii popaleontologii SSSR, 71, 1–104. [in Russian]
Weber, V.N. (1944) Trilobity kamennougol'nykh i permskikh otlozheniy SSSR. Vypusk 2. Permskiye trilobity. Monografii popaleontologii SSSR, 1944, 1–30. [in Russian]
Woodward, H. (1884) The Carboniferous Trilobites, Part 2. Monograph of the Palaeontographical Society, London, 182, 39–86.
Zhou, Z.-Q. (1983) Trilobita. In: Shengyang institute of geology and mineral resources (Ed.), Paleontological Atlas of Northeast China. Geological Publishing House, Beijing, pp. 475–477. [in Chinese]