Abstract
The World species of the genus Tachyusa, Erichson are revised and the genus is redefined. Tachyusilla and Tachyusota are placed in synonymy with Tachyusa. The revised Tachyusa includes 53 species, 16 of which are described as new: T. americanoides sp. n., T. bertiae sp. n., T. coarctatoides sp. n., T. hammondi sp. n., T. harbinica sp. n., T. krugeri sp. n., T. loebli sp. n., T. mindoroensis sp. n., T. nilensis sp. n., T. pseudosulciventris sp. n., T. riftensis sp. n., T. schawalleri sp. n., T. schillhammeri sp. n., T. smetanai sp. n., T. testacea sp. n. and T. turcica sp. n. The following new synonymies are proposed (each first name being valid): Tachyusa nitella Fauvel, 1895 = Tachyusa jonica Scheerpeltz, 1958, syn. nov., T. arida Casey, 1906 = T. ohioana Casey, 1911, syn. n.; T. elegans Cameron, 1939 = T. singalorum Pace, 1987, syn. n.; T. nitidula Mulsant & Rey, 1875 = T. iberica Fagel, 1957, syn. n.; T. obsoleta Casey, 1906 = T. pruinosa Casey, 1906, syn. n., = T. dakotana Casey, 1906, syn. n., = T. subaluatacea Casey, 1906, syn. n., = T. parviceps Casey, 1906, syn. n., = T. missouriana Casey, 1906, syn. n., = T. illini Casey, 1906, syn. n.; T. orientis Bernhauer, 1938 = T. reitteri Bernhauer, 1938, syn. n., = T. chinensis Pace, 1990, syn. n.; T. wei Pace, 1990 = T. hebeiensis Pace, 1998, syn. n. The following synonymies are confirmed: T. americana Casey, 1906 = T. meraca Casey, 1911, = T. silvatica Casey, 1911; T. balteata Erichson, 1839 = T. balteata elongata Kolenati, 1846; T. cavicollis LeConte, 1863 = T. carolinae Casey, 1906; T. coarctata Erichson, 1837 = T. coarctata cyanea Kraatz, 1856; T. faceta Casey, 1885 = T. vespertina Casey, 1906, = T. vaciva Casey, 1911; T. ferialis Erichson, 1839 = T. bicolor Mulsant & Rey, 1875; T. gracillima LeConte, 1863 = T. smithi Casey, 1906, = T. virginica Casey, 1911; T. nitidula Mulsant & Rey, 1875 = T. ventralis Fauvel, 1898; T. schuberti Jacobson, 1909 = T. bicolor Schubert, 1906, nom. preoc. A neotype is designated for Tachyusa nitella Fauvel, 1895. Both Tachyusa impressa Eppelsheim, 1877 and T. flavolimbata Eppelsheim, 1877, previously considered junior synonyms of T. agilis Baudi, 1869, and T. concinna Heer, 1838, previously considered a junior synonym of T. coarctata Erichson, 1937, are recognised as valid. All species are briefly described/redescribed and illustrated. An identification key to the World species of Tachyusa is provided. A phylogeny of fifty-three species belonging to the genus Tachyusa is proposed, based on fifty-six morphological characters. Cladistic analysis revealed the monophyly of the genus with two major clades.
References
Baudi, F. (1869) Coleopterorum messis in insula Cypro et Asiae minore ab Eugenio Truqui congregatae reconsitis de Europaeis notis quibusdam additis. Berliner entomologische Zeitschrift, 13, 369–418.
Bernhauer, M. (1902). Die Staphyliniden der paläarktischen Fauna. 2. Verhandlungen der k. k. Zoologisch-botanischen Gesellschaft in Wien, 52, 87–284.
Bernhauer, M. (1912) Zur Staphylinidenfauna von Südamerika. Wiener Entomologische Zeitung, 31, 68–75.
Bernhauer, M. (1915) Neue Staphyliniden der indo-malaiischen fauna, insbesondere der Sunda-Insel Borneo. Verhandlungen der k. k. Zoologisch-botanischen Gesellschaft in Wien, 65, 134–158.
Bernhauer, M. (1929) Neue Staphyliniden aus Mittelamerika. Wiener Entomologische Zeitung, 46, 186–208.
Bernhauer, M. (1938) Neuheiten der Staphylinidenfauna der Mandschurei. Koleopterologische Rundschau, 24, 20–29.
Bernhauer, M. (1938a) Zur Staphylinidenfauna von China u. Japan. Entomologisches Nachrichtenblatt, 12, 97–109.
Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Schenkling, S (ed.) Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, 499–988.
Cameron, M. (1920) New species of Staphylinidae from Ceylon. Part II. Entomologists Monthly Magazine, 56, 94–99.
Cameron, M. (1928) New species of Staphylinidae from Borneo. Sarawak Museum Journal, 3, 413–422.
Cameron, M. (1939) The Fauna of British India, including Ceylon and Burma. Coleoptera, Staphylinidae. Vol. 4, pt. 1. Taylor & Francis, London, 410 pp.
Cameron, M. (1940) New species of Staphylinidae (Col.) From Java. Entomologists Monthly Magazine, 76, 103–106.
Casey, T. L. (1885) New genera and species of Californian Coleoptera. Bulletin of the California Academy of Sciences, 1, 285–336.
Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transaction of the Academy of Sciences of St. Louis, 16(6), 125–356.
Casey, T. L. (1911) New American species of Aleocharinae and Myllaeninae. Memoirs on the Coleoptera, 2, 1–245.
Ciceroni, A., Puthz, V. & Zanetti A. (1995) Coleoptera Polyphaga III (Staphylinidae). In: Minelli, A., Ruffo, S. & La Posta, S. (Ed.), Checklist della specie della fauna italiana. 48. Calderini, Bologna, pp. 1–64.
Eppelsheim, E. (1877) Staphylinidae. In: Schneider, O. & Leder, H. (Ed.) Beiträge zur Kenntnis der Kaukasischen Käferfauna. Verhandlungen des naturforschenden Vereines in Brünn, 16, 90–131.
Eppelsheim, E. (1893) Beitrag zur Staphylinen–Fauna des südwestlichen Baikal–Gebietes. Deutsche entomologische Zeitschrift, 1893, 17–67.
Erichson, W. F. (1837) Die Käfer der Mark Brandenburg. Bd. 1, Abt. 1. F. H. Morin, Berlin, 384 pp.
Erichson, W. F. (1839) Genera et species Staphylinorum, insectorum coleopterorum familiae, F. H. Morin, Berlin, 954 pp.
Fagel, G. (1957) Contribution à la connaissance des Staphylinidae. XLII. Sur quelques espèces de Berberie. Bulletin et Annales de la Société Royale d’Entomologie de Belgique, 93, 276–282.
Fairmaire, L. (1849) Essai sur les Coléoptères de la Polynésie. Revue et Magasin de Zoologie, 1, 277–291.
Fairmaire, L. (1859) Diagnoses de quelques Coléoptères européens nouveaux. Annales de la
Societe entomologique de France, 10, 184–185.
Fauvel, A. (1879) Les Staphylinides des Moluques et de la Nouvelle-Guinée. Annali del Museo civico di storia naturale di Genova, 15, 63–120.
Fauvel, A. (1895) Notes synonymiques. Revue d’Entomologie, 14, 92–99.
Fauvel, A. (1898) Des Staphylinides de Barbarie, et des Iles Açores, Madères, Salvages et Canaries. Revue d’Entomologie, 17, 93–113.
Fauvel, A. (1907) Voyage de M. Ch. Alluaud dans l’Afrique orientale. Staphylinidae. Revue d’Entomologie, 26, 10–70.
Fenyes, A. (1920) Coleoptera: Fam. Staphylinidae, Subfam. Aleocharinae. Genera Insectorum, 173b, 111–414.
Ganglbauer, L. (1895) Die Käfer von Mitteleuropa. Vol. 2. Carl Gerolds Sohn, Wien, 881 pp.
Gravenhorst, J. L. C. (1806) Monographia Coleopterorum Micropterorum. Henricus Dieterich, Gottingae, 248 pp.
Heer, O. (1839) Fauna Coleopterorum Helvetica. Pars I, fasc. 2. Orelii, Fuesslini et Sociorum, Turici, pp. 145–360.
ICZN (1961) Opinion 600. Ischnopoda Stephens, 1835, and Tachyusa Erichson, 1837 (Insecta, Coleoptera); designations of type–species under the plenary powers. Bulletin of Zoological Nomenclature, 18(4), 241–243.
ICZN (1999) International Code of Zoological Nomenclature. Fourth edition. The International Union of Biological Sciences, London, 306 pp.
Jacobs, W. (1925) Ueber den Gattungsnamen Cardiola Muls. and Rey. Entomologische Zeitschrift, 38, 82.
Jacobson, G. (1909) Zhuki Rossii i Zapadnoi Evropy. Rukovodstvo k opredieleniiu Zhukov. Fasc. 7. A. F. Devrien, S. Petersbourg, pp. 481-560. [In Russian]
Jekel, H. (1873) Coleoptera Jekeliana, adiecta Eleutheratorum Bibliotheca, Énumération systématique et synonymique des coléoptères européens et exotiques composant la collection de Henri Jekel. Observations critiques. Descriptions despèces nouvelles. Reproduction et traduction de genres et espèces publiés dans des ouvrages français et etrangers de manière à former insensiblement la bibliothèque du Coléoptèriste. Livraison 1. E. Deyrolle fils, Paris, 96 pp.
Klimaszewski, J. & Peck, S.B. (1986) A review of the cavernicolous Staphylinidae (Coleoptera) of Eastern North America: Part I. Aleocharinae. Quaestiones Entomologicae, 22(2), 51–113.
Koch, C. (1936) Wissenschaftliche Ergebnisse der entomologischen expeditionen Seiner Durchlaucht des Fürsten Alessandro C. della Torre e Tasso nach Aegypten und auf die Halbinsel Sinai. XIII. Staphylinidae. Pubblicazioni del Museo Entomologico “Pietro Rossi” Udine, 1, 115–232.
Kolenati, F. A. (1846) Melethemata entomologica. Petropolis: Imperialis Academiae Scientiarum, 3, 1–44.
Kraatz, G. (1856) Naturgeschichte der Insecten Deutschlands. Erste Abteilung. Coleoptera. Bd. 2. Lief. 1–2. Verlag der Nicolaischen Buchhandlung, Berlin, pp. 1–376.
Last, H. R. (1966) Coleoptera from Southeast Asia (IV). Family Staphylinidae. Memoirs of the Faculty of Liberal Arts and Education Kagawa University, part II, No. 135, 1–10.
Leach, W. E. (1819) [New genera] In: G. Samouelle, The Entomologist's Useful Compendium; or an introduction to the knowledge of British insects, comprising the best means of obtaining and preserving them, and a description of the apparatus generally used; together with the genera of Linné, and the modern method of arranging the classes Crustacea, Myriapoda, spiders, mites, and insects from their affinities and structure, according to the views of Dr. Leach. Also an explanation of the terms used in entomology; a calendar of the times of appearance, and usual situations of near 3000 species of British insects; with instructions for collecting and fitting up objects for the microscope, Thomas Boys, London, pp. 1–496.
LeConte, J. L. (1863) New species of North American Coleoptera. Smithsonian Miscellaneous Collections, 6(167), 1–92.
Lohse, G. A. (1974) Staphylinidae II (Hypocyphtinae und Aleocharinae). In: Freude, H., Harde, K. W. & Lohse, G. A. (Ed.) Die Käfer Mitteleuropas, Band 5. Goecke & Evers, Krefeld, 381 pp.
Lohse, G. A. (1989) Familie: Staphylinidae. In: Lohse, G. A. & Lucht, W. H. (Ed.) Die Käfer Mitteleuropas, Band 12, 1 Suplementband mit Katalogteil. Goecke & Evers, Krefeld, 185–240 pp.
Lucht, W. & Klausnitzer, B. (1998) Die Käfer Mitteleuropas. 4. Supplementband. Goecke & Evers, Krefeld, pp. 119–197.
Mannerheim, C.G. (1830) Précis dun nouvel arrangement de la famille des Brachélytres, de lordre des Insectes Coléoptères. St. Pétersbourg, 87 pp.
Mulsant, M. E. & Rey, C. (1870) Quelques insectes nouveaux ou peu connus. Opuscules Entomologiques, 14, 153–193.
Mulsant, M.E. & Rey, C. (1873) Description de divers coléoptères brévipennes nouveaux ou peu connus. Opuscules Entomologiques, 15, 147–189.
Mulsant, M.E. & Rey, C. (1874) Tribu des Brévipennes. Famille des Aléochariens. Septième branche: Myrmédoniaires. Annales de la Société dAgriculture, Histoire Naturelle et Arts Utiles de Lyon, série 4. 6 [1873], 33–738.
Mulsant, M. E. & Rey, C. (1875) Histoire Naturelle des Coléoptères de France: Brévipennes: Aléochariens (suite). Myrmédoniaires, pt. 2. Deyrolle, Paris, 470 pp.
Muona, J. (1979) Staphylinidae. In: Silfverberg, H. (Ed.) Enumeratio Coleopterorum Fennoscandiae et Daniae. Helsingfors Entomologiska Bytesförening, Helsinki, 14–28.
Newton, A.F. & Thayer, M.K. (1992) Current Classification and Family-Group Names in Staphyliniformia (Coleoptera). Fieldiana (Zoology), 67, 1–92.
Newton, A.F., Thayer, M.K., Ashe, J.S. & Chandler, D.S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R.H. & Thomas, M.C. (Ed.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Baton, Florida, pp. 272–443.
Pace, R. (1984) Aleocharinae della Thailandia e della Birmania riportate da G. de Rougemont. Bollettino del Museo Civico di Storia Naturale di Verona, 11, 427–468.
Pace, R. (1985) Aleocharinae dell Himalaya raccolte da Guillaume de Rougemont (Coleoptera, Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona, 12, 165–191.
Pace, R. (1986) Aleocharinae dell Asia sudorientale raccolte da G. de Rougemont. Bollettino del Museo Civico di Storia Naturale di Verona, 13, 139–237.
Pace, R. (1987) Aleocharinae dello Sri Lanka raccolte du Marc Tronquet. Bollettino del Museo Civico di Storia Naturale di Verona, 14, 315–338.
Pace, R. (1990) Aleocharinae della Cina (Coleoptera, Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona, 17, 69––126.
Pace, R. (1998) Aleocharinae della Cina: Parte II (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 105(2), 395––463.
Pace, R. (1999) Faune de Madagascar. Insectes, Coléopteres, Staphylinidae, Aleocharinae. Paris, 261 pp.
Pace, R. (2000) Aleocharinae di Papua-Nuova Guinea (Coleoptera, Staphylinidae). Bulletin de lInstitut Royal des Sciences Naturelles de Belgique, Entomologie, 70, 109–163.
Paœnik, G. (2001) The North Korean Aleocharinae (Coleoptera, Staphylinidae): diversity and biogeography. Acta zoologica cracoviensia, 44(3): 185–234.
Redtenbacher, L. (1849) Fauna Austriaca. Die Käfer. Nach der Analytischen Methode bearbeitet. C. Gerold, Wien, xxvii + 883 pp.
Redtenbacher, L. (1856) Fauna Austriaca. Die Käfer. Nach der Analytischen Methode bearbeitet. C. Gerold, Wien, pp. 1–128.
Reitter, E. (1909) Fauna Germanica. Die Käfer des Deutschen Reiches. Nach der analytischen Methode bearbeitet. Bd. 2. K. G. Lutz, Stuttgart, 392 pp.
Saulcy, F. de (1864) Descriptions des espèces nouvelles de Coléoptères recueillies en Syrie, en Égypte et en Palestine, pendant les mois doctobre 1863 à janvier 1864, par M. de Saulcy, Senateur, Membre de lInstitut. Pt. 2. Annales de la Société Entomologique de France, (4)4, 629–660.
Scheerpeltz, O. (1958) Zoologischen Studien in West-Griecheland von Max Beir, Wien. VIII. Staphylinidae (Col.). Sitzungsberichte der Österreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abt. I, 167, 363–432.
Scheerpeltz, O. (1963) Ergebnisse der Zoologischen Nubien-Expedition 1962 (Coleoptera, Staphylinidae). Annalen des Naturhistorischen Museums in Wien, 66, 425–450.
Scheerpeltz, O. (1974) Coleoptera: Staphylinidae (exclus. Subfam. Paederinae, except. pars min.). In: Hanstrom B., Brinck P. & Rudebeck G. (eds) South African Animal Life, 15, 43–394.
Schubert, K. (1906) Einige neue paläarctische Staphyliniden aus Kashmir. Deutsche entomologische Zeitschrift, 1906, 378–383.
Seevers, C. H. (1978) A generic and tribial revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana (Zoology), 71, 1–275.
Stephens, J. F. (1832) Illustrations of British entomology; or, a synopsis of indigenous insects; containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable, Mandibulata, V. Baldwin and Cradock, London, 5, 1–240.
Swofford, D. L. (2001) PAUP*. Phylogenetic Analysis Using Parsimony (* and Other Methods). Version 4.0b10. Sinauer Associates, Sunderland, Massachusetts.
Thomson, C.G. (1858) Försök till uppställning af Sveriges Staphyliner. Öfversigt af Kongl. Vetenskaps-Akademiens Förhandlingar, 15, 27–40.
Thomson, C.G. (1859). Skandinaviens Coleoptera, synoptiskt bearbetade. Tom 1. Berlingska Boktryckeriet, Lund, v+290 pp.
Tottenham, C. E. (1945) Some notes on the nomenclature of the Staphylinidae (Coleoptera). The Proceedings of the Royal Entomological Society of London, (B), 14, 70–71.
Yosii, R. & Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II. Diagnostic characters of Genera and Subgenera with description of representative Species. Contributions from the Biological Laboratory, Kyoto University, 25(1), 11–140.