Abstract
A cladistic analysis of the Empidoidea and basal lineages of the Cyclorrhapha, based on morphological characters, confirms the monophyly of both groups as well as that of the Eremoneura. The resulting final trees are used to revise the classification of the Empidoidea to include the following five families: Empididae, Hybotidae, Atelestidae (including Nemedininae n. subfam.), Brachystomatidae rev. stat. (comprising the subfamilies Brachystomatinae, Ceratomerinae and Trichopezinae), and Dolichopodidae s.lat. The family Microphoridae is not recognized, and the Microphorinae and Parathalassiinae are assigned to the Dolichopodidae s.lat. The Dolichopodidae s.str. includes 15 subfamilies that were previously recognized within the family. Within the Empidoidea we found support for Atelestidae as the sister group to the Hybotidae and for the monophyly of Parathalassiinae + Dolichopodidae s.str. The Empididae remains poorly defined and the genera Homalocnemis Philippi, Iteaphila Zetterstedt, Anthepiscopus Becker, and Oreogeton Schiner are classified as incertae sedis within the Empidoidea. In addition, the following higher taxa are proposed: Symballophthalmini n. tribe, Bicellariini n. tribe, Oedaleinae rev. stat., and Trichininae rev. stat., which are all assigned to the Hybotidae. The genus Sematopoda Collin is tentatively assigned to Trichopezinae, and Xanthodromia Saigusa is transferred from Hemerodromiinae to Brachystomatinae.
All morphological characters are extensively discussed and illustrated, including details of the antennae, mouthparts, internal thoracic structures, wings, and male and female terminalia. In addition, a key to families and unplaced genus groups of the Empidoidea is provided. Feeding habits are also discussed in terms of the empidoid ground plan condition.
References
Adisoemarto, S. & Wood, D.M. (1975) The Nearctic species of Dioctria and six related genera (Diptera: Asilidae). Quaestiones Entomologicae, 11, 505–576.
Agassiz, J.L.R. (1846) Nomina systematica generum Dipterorum (Pt. 4). In Agassiz, J.L.R., Nomenclator zoologicus continens nomina systematica generum animalium tam viventium quam fossilium, secundum ordinem alphabeticum disposita, adjectis auctoribis, libris, in quibus reperiuntur, anno editionis, etymologia et familias, ad quas pertinent, in singulis classibus. Fasc. 9/10, 1–42. Soloduri [= Solothurn].
Ale-Rocha, R. (2000) Description of Smithybos equatoriensis, new genus and species of Hybotinae from Ecuador (Diptera, Empididae). Revista Brasileira de Entomologica, 22(1/2), 39–42.
Ale-Rocha, R. & Carvalho, C.J.B. de (2003) Neohybos gen. nov. (Diptera, Empidoidea, Hybotinae) from the Neotropical Region. Zootaxa, 387, 1–16.
d’Assis Fonseca, E.C.M. (1978) Diptera Orthorrhapha Brachycera Dolichopodidae. Handbooks for the Identification of British Insects, 9(5), 1–90.
Bährmann, R. (1960) Vergleichend-morphologische Untersuchungen der männlichen Kopulationsorgane bei Empididen. Beiträge zur Entomologie, 10(5/6), 485–540.
Bährmann, R. (1966) Das Hypopygium von Dolichopus Latreille unter besonderer Berücksichtigung der Muskulatur und der Torsion (Diptera; Dolichopodidae). Beiträge zur Entomologie, 16(1/2), 61–72.
Barták, M. (2000) Atelestidae. In Barták, M. & Va¡hara, J. (Ed.), Diptera in an industrially affected region (North-Western Bohemia, Bílina and Duchcov Environs). Vol. 1. Folia Facultatis Scientiarium Naturalium Universitatis Masarykianae Brunensis, Biologia, 104, 201–202.
Becker, T. (1889) Berichtigung. Wiener Entomologische Zeitung, 8, 285.
Beutel, R.G. & Gorb, S.N. (2001) Ultrastructure of attachment specializations of hexapods (Arthropoda): evolutionary patterns inferred from a revised ordinal phylogeny. Journal of Zoological Systematics and Evolutionary Research, 39, 177–207.
Bickel, D.J. (1985) A revision of the Nearctic Medetera (Diptera: Dolichopodidae). U.S. Department of Agriculture, Technical Bulletin, 1692, 1–109.
Bickel, D.J. (1987) Babindellinae, a new subfamily of Dolichopodidae (Diptera) from Australia, with a description of symmetry in the dipteran male postabdomen. Entomologica Scandinavica, 18, 97–113.
Bickel, D.J. (1994) The Australian Sciapodinae (Diptera: Dolichopodidae), with a review of the Oriental and Australasian faunas, and a World Conspectus of the subfamily. Records of the Australian Museum, Supplement, 21, 1–394.
Bickel, D.J. (1996a) Thinempis, a new genus from Australia and New Zealand (Diptera: Empididae), with notes on the tribal classification of the Empidinae. Systematic Entomology, 21, 115–128.
Bickel, D.J. (1996b) Smoke flies (Diptera: Platypezidae) and the Sydney bushfires. Australian Entomologist, 23(2), 77–78.
Bickel, D.J. (1998) Cunomyia, a distinctive new hilarine fly genus from the Tasmanian World Heritage Area (Diptera: Empididae). Papers and Proceedings of the Royal Society of Tasmania, 132, 59–63.
Bickel, D.J. (2002) Bandella, a new hilarine fly genus from Australia (Diptera: Empididae). Records of the Australian Museum, 54(3), 313–324.
Bletchly, J.D. (1954) The mouth-parts of the Dance Fly, Empis livida L. (Diptera, Empididae). Proceedings of the Zoological Society of London, 124, 317–334.
Brauer, F. (1883) Die Zweiflügler des Kaiserlichen Museums zu Wien. III. Systematische Studien auf Grundlage der Dipteren-Larven nebst einer Zusammenstellung von Beispielen aus der Literatur über dieselben und Beschreibung neuer Formen. Denkschriften der Akademie der Wissenschaften, Wien, 47, 1–100, pls I–V.
Bremer, K. (1994) Branch support and tree stability. Cladistics, 10, 295–304.
Brooks, S.E. (2005a) Systematics and phylogeny of Dolichopodinae (Diptera: Dolichopodidae). Zootaxa, 857, 1–158.
Brooks, S.E. (2005b) Higher classification and phylogeny of Dolichopodidae. http://www.nadsdiptera.org/Doid/Doliclass/Doliclass.htm. [Accessed on 1 February 2006].
Brown, B.V. (1992) Generic revision of Phoridae of the Nearctic Region and phylogenetic classification of Phoridae, Sciadoceridae, and Ironomyiidae (Diptera: Phoridea). Memoirs of the Entomological Society of Canada, 164, 1–144.
Brown, B.V. (1995) Response to Disney. Journal of Natural History, 29, 259–264.
Carney, M. (1999) What was found in the burn. Seasons, 39(4), 26–31.
Chandler, P.J. (1981) Nemedina alamirabilis sp.n., a new genus and species of Diptera Eremoneura of uncertain affinities, from Hungary. Acta Zoologica Academiae Scientiarum Hungaricae, 27 (1/2), 103–113.
Chandler, P.J. (1994) The Oriental and Australasian species of Platypezidae (Diptera). Invertebrate Taxonomy, 8, 351–434.
Chandler, P.J. (1998) 3.2. Family Opetiidae. In Papp, L. & Darvas, B. (Ed.), Contributions to a Manual of Palaearctic Diptera, Vol. 3. Higher Brachycera. Science Herald, Budapest, pp. 17–25.
Chandler, P.J. (2001) The flat-footed flies (Diptera: Opetiidae and Platypezidae) of Europe. Fauna Entomologica Scandinavica, 36, 1–276.
Chandler, P.J. & Shatalkin, A.I. (1998) 3.3. Family Platypezidae. In Papp, L. & Darvas, B. (Eds.), Contributions to a Manual of Palaearctic Diptera, Vol. 3. Higher Brachycera. Science Herald, Budapest, pp. 27–49.
Chvála, M. (1975) The Tachydromiinae (Dipt. Empididae) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica, 3, 1–336.
Chvála, M. (1976) Swarming, mating and feeding habits in Empididae (Diptera), and their significance in evolution of the family. Acta Entomologica Bohemoslovaca, 73, 353–366.
Chvála, M. (1981) Classification and phylogeny of Empididae, with a presumed origin of Dolichopodidae (Diptera). Entomologica Scandinavica Supplement, 15, 225–236.
Chvála, M. (1983) The Empidoidea (Diptera) of Fennoscandia and Denmark. II. General Part. The families Hybotidae, Atelestidae and Microphoridae. Fauna Entomologica Scandinavica, 12, 1–279.
Chvála, M. (1986) Revision of Palearctic Microphoridae (Diptera) 1. Microphor Macq. Acta Entomologica Bohemoslovaca, 83, 432–454.
Chvála, M. (1987) Revision of Palearctic Microphoridae (Diptera) 2. Schistostoma Beck. Acta Entomologica Bohemoslovaca, 84, 133–155.
Chvála, M. (1988) Revision of Palearctic Microphoridae (Diptera) 3. Parathalassiinae (Parathalassius Mik and Microphorella Becker). Acta Entomologica Bohemoslovaca, 85, 352–372.
Chvála, M. (1989) Microphoridae. In Soós, Á. & Papp, L. (Eds.), Catalogue of Palaearctic Diptera, Volume 6, Therevidae — Empididae. Elsevier Science Publishing, pp. 171–174.
Chvála, M. (1991) First record of the relict Gondwanan genus Homalocnemis Philippi (Diptera: Empididae) from Namibia and the Afrotropical Region. Annals of the Natal Museum, 32, 13–18.
Chvála, M. (1994) The Empidoidea (Diptera) of Fennoscandia and Denmark. III. Genus Empis. Fauna Entomologica Scandinavica, 29, 1–187.
Chvála, M. (1999) Redescription, biology and distribution of Hilara splendida Straka (Diptera: Empididae), and notes on holoptism in the genus. Acta Universitatis Carolinae Biologica, 43, 211–220.
Chvála, M. (2005) The Empidoidea (Diptera) of Fennoscandia and Denmark. IV. Genus Hilara. Fauna Entomologica Scandinavica, 40, 1–234.
Chvála, M. & Kovalev, V.G. (1989) Hybotidae. In Soós, Á. & Papp, L. (Eds.), Catalogue of Palaearctic Diptera, Volume 6, Therevidae — Empididae. Elsevier Science Publishing, pp. 174–227.
Chvála, M. & Wagner, R. (1989) Empididae. In Soós, Á. & Papp, L. (Eds.), Catalogue of Palaearctic Diptera, Volume 6, Therevidae — Empididae. Elsevier Science Publishing, pp. 228–336.
Colless, D.H. (1963) An Australian species of Microphorella (Diptera: Empididae), with notes on the phylogenetic significance of the genus. Proceedings of the Linnean Society of New South Wales, 88, 320–323.
Colless, D.H. & McAlpine, D.K. (1991) Diptera (Flies) [Chapter] 39. In Naumann, I.D. (Ed.), The Insects of Australia, Volume 2. 2nd ed. Melbourne University Press, Carlton, pp. 171–786.
Collin, J.E. (1928) New Zealand Empididae based on material in the British Museum (Natural History). British Museum (Natural History), London, 110 pp.
Collin, J.E. (1933) Empididae. Diptera of Patagonia and South Chile, 4, 1–334.
Collin, J.E. (1960) Some Empididae from Palestine. Annals and Magazine of Natural History, (13) 2 (1959), 385–420.
Collin, J.E. (1961) Empididae. In British Flies, Volume 6. University Press, Cambridge, 782 pp.
Collins, K.P. & Wiegmann, B.M. (2002a) Phylogenetic relationships and placement of the Empidoidea (Diptera: Brachycera) based on 28S rDNA and EF–1a sequences. Insect Systematics and Evolution, 33, 421–444.
Collins, K.P. & Wiegmann, B.M. (2002b) Phylogenetic relationships of the Cyclorrhapha (Diptera: Brachycera) based on 28S rDNA. Insect Systematics and Evolution, 33, 445–456.
Courtney, G.W., Sinclair, B.J. & Meier, R. (2000) 1.4. Morphology and terminology of Diptera larvae. In Papp, L. & Darvas, B. (Ed.), Contributions to a Manual of Palaearctic Diptera, Vol. 1. General and Applied Dipterology. Science Herald, Budapest, pp. 85–161.
Crampton, G.C. (1942) The external morphology of the Diptera. In Guide to the Insects of Connecticut, Part VI. The Diptera or true flies of Connecticut, 1st fascicle. Connecticut State Geological and Natural History Survey, Bulletin , 64, 10–165.
Cregan, M.B. (1941) Generic relationships of the Dolichopodidae (Diptera) based on a study of mouth parts. Illinois Biological Monographs, 18(1), 1–68.
Cumming, J.M. (1989) Book review of Chvála, M. 1983. The Empidoidea (Diptera) of Fennoscandia and Denmark. II. General Part. The families Hybotidae, Atelestidae, and Microphoridae. Fauna Entomologica Scandinavica, Volume 12. The Canadian Field-Naturalist, 103, 620.
Cumming, J.M. (1990) Phylogenetic relationships within the Tachydromiinae (Diptera: Empidoidea). Second International Congress of Dipterology, Bratislava, Abstract Volume, p. 44.
Cumming, J.M. (1994) Sexual selection and the evolution of dance fly mating systems (Diptera: Empididae; Empidinae). The Canadian Entomologist, 126, 907–920.
Cumming, J.M. & Brooks, S.E. (2002) Electrophorella, a new genus of parathalassiine flies from Baltic amber, with a cladistic analysis of the Microphorinae + Dolichopodidae lineage (Diptera: Empidoidea). Studia Dipterologica, 9, 41–54.
Cumming, J.M. & Cooper, B.E. (1992) A revision of the Nearctic species of the tachydromiine fly genus Stilpon Loew (Diptera: Empidoidea). The Canadian Entomologist, 124, 951–998.
Cumming, J.M. & Cooper, B.E. (1993) Techniques for obtaining adult-associated immature stages of predacious tachydromiine flies (Diptera: Empidoidea), with implications for rearing and biocontrol. Entomological News, 104, 93–101.
Cumming, J.M. & Sinclair, B.J. (1990) Fusion and confusion: interpretation of male genitalic homologies in the Empidoidea (Diptera). Second International Congress of Dipterology, Bratislava, Abstract Volume, 3rd Supplement, p. 334.
Cumming, J.M. & Sinclair, B.J. (1996) The higher-level phylogeny of Eremoneura (Diptera: Brachycera). XX International Congress of Entomology, Proceedings. Florence, Italy, p. 24.
Cumming, J.M. & Sinclair, B.J. (2000) A new phylogenetic classification of the Empidoidea (Diptera: Eremoneura). Abstracts of XXI International Congress of Entomology (Brazil), Volume 2. p. 924.
Cumming, J.M. & Sinclair, B.J. (in press) Family Empididae. In Brown, B.V., Borkent, A., Wood, D.M. & Zumbado, M. (Eds.), Manual of Central American Diptera.
Cumming, J.M., Sinclair B.J. & Wood, D.M. (1995) Homology and phylogenetic implications of male genitalia in Diptera—Eremoneura. Entomologica Scandinavica, 26, 121–151.
Cumming, J.M. & Wood, D.M. (in press) Morphology and terminology. In Brown, B.V., Borkent, A., Wood, D.M. & Zumbado, M. (Eds.), Manual of Central American Diptera.
Daugeron, C. (1997) Evolution of feeding and mating behaviours in the Empidoidea (Diptera: Eremoneura). In Grandcolas, P. (Ed.), The Origin of Biodiversity in Insects: Phylogenetic Tests of Evolutionary Scenarios. Mémoires du Muséum National dHistoire Naturelle, Paris, 173, 163–182.
Disney, R.H.L. (1988) The form of articulation between the pedicel and first flagellar segment of the antenna in flies (Diptera). The Entomologist, 107, 99–103.
Disney, R.H.L. (2001) Sciadoceridae (Diptera) reconsidered. Fragmenta Faunistica, 44, 309–317.
Downes, J.A. (1969) The swarming and mating flight of Diptera. Annual Review of Entomology, 14, 271–298.
Downes, J.A. & Smith, S.M. (1969) New or little known feeding habits in Empididae (Diptera). The Canadian Entomologist, 101(4), 404–408.
Dufour, L. (1851) Recherches anatomiques et physiologiques sur les Diptères. Mémoires présentés par divers savants à l’Académie des Sciences de l’Institut de France et imprimés par son ordre: Sciences mathématiques et physiques. Paris, 11, 171–360.
Dyte, C.E. (1967) Some distinctions between the larvae and pupae of the Empididae and Dolichopodidae (Diptera). Proceedings of the Royal Entomological Society of London, (A) 42, 119–128.
Dyte, C.E. (1988) Mate guarding and sex ratio in Hydrophorus oceanus (Macquart) (Diptera: Dolichopodidae). The Entomologist, 107, 122–126.
Eltringham, H. (1928) On the production of silk by the genus Hilara Meig. (Diptera). Proceedings of the Royal Society, (B) 102, 327–338.
Funk, D.H. & Tallamy, D.W. (2000) Courtship role reversal and deceptive signals in the long-tailed dance fly, Rhamphomyia longicauda. Animal Behaviour, 59, 411–421.
Gatt, P. (2003) New species and records of Microphorella Becker (Diptera: Empidoidea, Dolichopodidae) from the Mediterranean region. Revue Suisse de Zoologie, 110(4), 669–684.
Giebel, C.G. (1856) Die Insecten und Spinnen der Vorwelt, mit steter Berücksichtigung der lebenden Insecten und Spinnen, monographisch dargestellt. F.A. Brockhaus, Leipzig. xvii + 511 pp.
Greathead, D.J. & Evenhuis, N.L. (2001) Annotated keys to the genera of African Bombylioidea (Diptera: Bombyliidae; Mythicomyiidae). African Invertebrates, 42, 105–224.
Griffiths, G.C.D. (1972) The phylogenetic classification of Diptera Cyclorrhapha, with special reference to the structure of the male postabdomen. Series Entomologica 8. Dr. W. Junk, The Hague, pp. 340.
Griffiths, G.C.D. (1983) Book review of Chvála, M. 1983. The Empidoidea (Diptera) of Fennoscandia and Denmark. II. The families Hybotidae, Atelestidae and Microphoridae. Fauna Entomologica Scandinavica, Volume 12. Quaestiones Entomologicae, 19(3/4), 483–485.
Griffiths, G.C.D. (1994) Relationships among the major subgroups of Brachycera (Diptera): a critical review. The Canadian Entomologist, 126, 861–880.
Griffiths, G.C.D. (1996) Review of the papers on the male genitalia of Diptera by D.M. Wood and associates. Studia Dipterologica, 3(1), 107–123.
Grimaldi, D. (1999) The co-radiations of pollinating insects and angiosperms in the Cretaceous. Annals of the Missouri Botanical Garden, 86(2), 373–406.
Grimaldi, D. & Cumming, J. (1999) Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History, 239, 1–124.
Grichanov, I.Y. (2006) All About Dolichopodidae (Diptera: Empidoidea). http://www.fortunecity.com/greenfield/porton/875/Genera3.htm. [Accessed on 21 February 2006].
Grootaert, P. (1994) Two new genera of empidoid flies (Diptera, Empidoidea, Hybotidae, Tachydromiinae) from Melanesia. Studia Dipterologica, 1(2), 141–156.
Hackman, W. & Väisänen, R. (1985) The evolution and phylogenetic significance of the costal chaetotaxy in the Diptera. Annales Zoologici Fennici, 22, 169–203.
Hardy, G.H. (1954) The phylogeny of Diptera 3.—Empididae. Entomologists Monthly Magazine, 90, 78–90.
Hardy, G.H. (1962) The anal field in the wing venation of Empididae and related families. Journal of the Entomological Society of Queensland, 1, 25–29.
Hendel, F. (1936–1937) Diptera. In Kükenthal, W. & Krumbach, T. (Ed.), Handbuch der Zoologie, 4 (2), 1729–1998.
Hennig, W. (1970) Insektfossilien aus der unteren Kreide. II. Empididae (Diptera, Brachycera). Stuttgarter Beiträge zur Naturkunde, 214, 1–12.
Hennig, W. (1971) Insektfossilien aus der unteren Kreide. III. Empidiformia (“Microphorinae”) aus der unteren Kreide und aus dem Baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beiträge zur Naturkunde, 232, 1–28.
Hennig, W. (1976) Das Hypopygium von Lonchoptera lutea Panzer und die phylogenetischen Verwandtschaftsbeziehungen der Cyclorrhapha (Diptera). Stuttgarter Beiträge zur Naturkunde, 283, 1–63.
Irwin, M.E. (1974) An account of the southern African genus Sciopolina with descriptions of new species (Diptera: Dolichopodidae: Sciapodinae). Annals of the Natal Museum, 22, 221–264.
Irwin, M.E. (1976) Morphology of the terminalia and known ovipositing behaviour of female Therevidae (Diptera: Asiloidea), with an account of correlated adaptations and comments on phylogenetic relationships. Annals of the Natal Museum, 22, 931–935.
Irwin, M.E & Lyneborg, L. (1981) Therevidae [Chapter] 37. In McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Coords.), Manual of Nearctic Diptera, Volume 1. Agriculture Canada Monograph, 27, 513–523.
Jürgens, G., Lehmann, R., Schardin, M. & Nüsslein-Volhard, C. (1986) Segmental organization of the head in the embryo of Drosophila melanogaster. Rouxs Archives of Developmental Biology, 195, 359–377.
Kessel, E.L. (1989) Autobiographical Anecdotes. Myia, 4, 1–186.
Krystoph, H. (1961) Vergleichend–morphologische Untersuchungen an den Mundteilen bei Empididen (Diptera). Beiträge zur Entomologie, 11(7/8), 824–872.
Latreille, P.A. (1797) Précis des caractères génériques des insectes, disposés dans un ordre naturel par le Citoyen Latreille. “An V”. Prévôt, Paris; F. Bordeaux, Brive, xiii + [i] + 201 + [7] pp.
Latreille, P.A. (1809) Genera crustaceorum et insectorum secundum ordinem naturalem in familias disposita, iconibus exemplisique plurimis explicata. Tomus quartus et ultimus. A. Koenig, Parisiis et Argentorati, 399 pp.
Laurence, B.R. (1953) On the feeding habits of Clinocera (Wiedemannia) bistigma Curtis (Diptera: Empididae). Proceedings of the Royal Entomological Society of London (A), 28, 139–144.
Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum caracteribus, differentiis, synonymis, locis. Tomus I. Edito decima, reformata. L. Salvii, Holmiae [= Stockholm], 824 pp.
MacDonald, J.F. (1988) New synonyms pertaining to Chelifera and generic key for North American Hemerodromiinae (Diptera: Empididae). Proceedings of the Entomological Society of Washington, 90, 98–100.
MacDonald, J.F. (1998) Review of the genus Hemerodromia Meigen of America north of Mexico (Diptera: Empididae; Hemerodromiinae). Proceedings of the Entomological Society of Washington, 100, 792–823.
Macquart, J. (1823) Monographie des insectes diptères de la famille des empides, observés dans le nord-ouest de la France. Recueil des Travaux de la Société d’Amateurs des Sciences, de l’Agriculture et des Arts à Lille, 1822, 137–165.
Macquart, J. (1827) Insectes diptères du nord de la France. Platypézines, dolichopodes, empides, hybotides. Lille, 159 pp.
Maddison, W.P. & Maddison, D.R. (2003) MacClade 4: Analysis of phylogeny and character evolution. Version 4.06. Sinauer Associates, Sunderland, Massachusetts.
Marina, T.A. (1985) Comparative morphology of prothoracic sclerites in Asilomorpha (Diptera). In Skarlato, O.A. (Ed.), Systematics of Diptera (Insecta) Ecological and Morphological Principles. Amerind Publ. Co., New Delhi, pp. 76–91 [Translated from Zoologicheskii Institut An SSSR Publishers, Leningrad (1979)].
Matsuda, R. (1965) Morphology and evolution of the insect head. Memoirs of the American Entomolgical Institute, 4, 1–334.
McAlpine, J.F. (1981) Morphology and terminology – Adults [Chapter] 2. In McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Coords.), Manual of Nearctic Diptera, Volume 1. Agriculture Canada Monograph, 27, 9–63.
McAlpine, J.F. (1989) Phylogeny and classification of the Muscomorpha. [Chapter] 116. In McAlpine, J.F. (Ed.), Manual of Nearctic Diptera, Volume 3. Agriculture Canada Monograph, 32, 1396–1518.
Meigen, J.W. (1803) Versuch einer neuen Gattungseintheilung der europäischen zweiflügeligen Insekten. Magazin für Insektenkunde, 2, 259–281.
Meigen, J.W. (1820) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Zweiter Theil. Forstmann, Aachen, pp. 363.
Meigen, J.W. (1822) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Dritter Theil. Schulz-Wundermann, Hamm, pp. 416.
Meigen, J.W. (1830) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Sechster Theil. Schulz, Hamm. xi + 401 pp.
Meijere, J.C.H., de (1900) Ueber der Larve von Lonchoptera. Ein Beitrag zur Kenntnis der cyclorrhaphen Dipterenlarven. Zoolgische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, 14, 87–132.
Meijere, J.C.H., de (1906) Die Lonchopteren des palaearktischen Gebietes. Tijdschrift voor Entomologie, 49, 44–98, pl. 4–5.
Melander, A.L. (1908) Family Empididae. In Williston, S.W. (Ed.), Manual of North American Diptera. 3rd Edition. J.J. Hathaway, New Haven, pp. 218–227.
Melander, A.L. (1928) Diptera, Fam. Empididae. In Wytsman, P. (Ed.), Genera Insectorum, 185 (1927), 1–434.
Melin, D. (1923) Contributions to the knowledge of the biology, metamorphosis and distribution of the Swedish asilids. Zoologiska Bidrag Från Uppsala, 8, 1–317.
Meyer, H (2005) Langbein-, Tanz- und Schwebfliegen (Diptera: Empidoidea: Dolichopodidae, Empididae, Hybotidae; Syrphidae) im Totholz von Laubwäldern Schleswig-Holsteins. Faunistisch-Ökologische Mitteilungen, 8, 363–382.
Mik, J. (1891) Vorläufige Notiz über Parathalassius blasigii, ein neues Dipteron aus Venedig. Wiener Entomologische Zeitung, 10, 216–217.
Mostovski, M.B. (1999) A brief review of brachycerous flies (Diptera, Brachycera) in the Mesozoic, with descriptions of some curious taxa. Proceedings of the First International Palaeoentomological Conference, Moscow 1998, pp. 103–110.
Moulton, J.K. & Wiegmann, B.M. (2004) Evolution and phylogenetic utility of CAD (rudimentary) among Mesozoic–aged eremoneuran Diptera (Insecta). Molecular Phlyogenetics and Evolution, 31, 363–378.
Naglis, S.M. (2001) Revision of the Neotropical Neurigoninae (Diptera: Dolichopodidae) I: Coeloglutus Aldrich, Neotonnoiria Robinson, and Paracoeloglutus gen. nov., with the definition of the tribe Coeloglutini stat. nov. Studia Dipterologica, 8, 189–206.
Naglis, S.M. (2002a) Revision of the Neotropical Neurigoninae (Diptera: Dolichopodidae) II: Argentinia Parent, Dactylomyia Aldrich, Macrodactylomyia gen. nov., and Systenoides gen. nov., with the definition of a new tribe Dactylomyiini. Studia Dipterologica, 8 (2001), 475–504.
Naglis, S.M. (2002b) Revision of the Neotropical Neurigoninae (Diptera: Dolichopodidae) III: Bickelomyia gen. nov., with the definition of a new tribe Neurigonini. Studia Dipterologica, 9, 225–241.
Naglis, S.M. (2003a) Revision of the Neotropical Neurigoninae (Diptera: Dolichopodidae) IV: Viridigona gen. nov. Studia Dipterologica, 9 (2002), 561–604.
Naglis, S.M. (2003a) Revision of the Neotropical Neurigoninae (Diptera: Dolichopodidae) V: Neurigona Rondani. Studia Dipterologica, 10, 267–314.
Negrobov, O.P. (1986) On the system and phylogeny of flies of the fam. Dolichopodidae (Diptera). Entomologicheskoe Obozrenie, 65, 182–186 [English translation in Entomological Review (1987), 65, 16–20].
Newkirk, M.R. (1970) Biology of the longtailed dance fly, Rhamphomyia longicauda (Diptera: Empididae); a new look at swarming. Annals of the Entomological Society of America, 63, 1407–1412.
Ovtshinnikova, O.G. (1994) On the homology of male genital sclerites of Brachycera Orthorrhapha and Cyclorrhapha (Diptera) based on musculature. Dipterological Research, 5, 263–269.
Papp, L. & Földvári, M. (2001) A new genus and three new species of Hybotidae with new records of the Hungarian Empidoidea (Diptera). Acta Zoologica Academiae Scientiarum Hungaricae, 47(4), 349–361.
Parent, O. (1938) Diptères Dolichopodidae. Faune de France, 35, 1–720.
Peterson, A. (1916) The head-capsule and mouthparts of Diptera. Illinois Biological Monographs, 3(2), 1–112.
Philippi, R.A. (1865) Aufzählung der chilenischen Dipteren. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 15, 595–782.
Plant, A.R. (1989) A revision of the Ocydromiinae (Diptera: Empidoidea: Hybotidae) of New Zealand with descriptions of new genera and species. New Zealand Journal of Zoology, 16, 231–241.
Plant, A.R. (1997) Atodrapetis, a new genus of empidid fly (Diptera, Empidoidea, Hybotidae, Tachydromiinae) from New Zealand. Studia Dipterologica, 4, 435–440.
Plant, A.R. (2003) Phenology of Empididae and Hybotidae (Diptera) in Great Britain. Dipterists Digest, 10, 13–30.
Plant, A.R. (2004) Hilara Meigen (Diptera: Empididae) in Britain: a provisional synopsis of distribution, habitat preferences and behaviour. Acta Universitatis Carolinae Biologica, 48, 165–196.
Plant, A.R. & Didham, R.K. (2006). A new genus of Drapetini (Diptera: Hybotidae: Tachydromiinae) from New Zealand. Entomologists Monthly Magazine, 142, 41–47.
Pollet, M., Brooks, S.E. & Cumming, J.M. (2004) Catalog of the Dolichopodidae (Diptera) of America north of Mexico. Bulletin of the American Museum of Natural History, 283, 1–114.
Pollet, M. & Cumming, J.M. (1998) Systematic revision of Nearctic species of Achalcus Loew (Diptera: Dolichopodidae) with comments on their phylogeny, ecology and zoogeography. Systematic Entomology, 23, 371–385.
Preston-Mafham, K.G. (1999) Courtship and mating in Empis (Xanthempis) trigramma Meig., E. tessellata F. and E. (Polyblepharis) opaca F. (Diptera: Empididae) and the possible implications of cheating behaviour. Journal of Zoology, London, 247, 239–246.
Rafael, J.A. & Cumming, J.M. (2004) The Neotropical genera Macrostomus Wiedemann and Porphyrochroa Melander (Diptera, Empididae, Empidinae). Revista Brasileira de Zoologica, 21(3), 439–448.
Robinson, H. (1969) A monographic study of the Mexican species of Enlinia (Diptera: Dolichopodidae). Smithsonian Contributions to Zoology, 25, 1–62.
Robinson, H. (1970) The subfamilies of the family Dolichopodidae in North and South America (Diptera). Papéis Avulsos Zoologia, S. Paulo, 23(6), 53–62.
Robinson, H. & Vockeroth, J.R. (1981) Dolichopodidae [Chapter] 48. In McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Coords.), Manual of Nearctic Diptera, Volume 1. Agriculture Canada Monograph, 27, 625–639.
Röder, G. (1984) Morphologische Untersuchungen an Praetarsen von Diptera und Mecoptera (Insecta). Dissertation, Naturwissenschaftliche Fakultäten der Friedrich-Alexander-Universität, Erlangen-Nürnberg, 393 pp.
Röder, G. (1986) Zur Morphologie des Praetarsus der Diptera und Mecoptera. Zoologische Jahrbücher (Anatomie), 114, 465–502.
Rondani, C. (1856) Dipterologiae Italicae prodromus. Vol. 1. Genera Italica ordinis Dipterorum ordinatim disposita et distincta et in familias et stirpes aggregata. A. Storschi, Parmae [= Parma], 228 pp.
Sabrosky, C.W. (1999) Family-group names in Diptera. An annotated catalog. Myia, 10, 1–360.
Saigusa, T. (1963) Systematic studies of Japanese Platypezidae I. Genus Opetia Meigen. Sieboldia, 3(1), 105–110.
Saigusa, T. (1986) New genera of Empididae (Diptera) from eastern Asia. Sieboldia, 5(1), 1–96.
Salzer, R. (1968) Konnstruktionsanatomische Untersuchung des männlichen Postabdomens von Calliphora erythrocephala Meigen (Insecta, Diptera). Zeitschrift für Morphologie der Tiere, 63, 155–238.
Satô, M. (1991) Comparative morphology of the mouthparts of the family Dolichopodidae (Diptera). Insecta Matsumurana, N.S., 45, 49–75.
Schiner, J.R. (1862) [Heft 8 (part)]. In Fauna Austriaca. Die Fliegen (Diptera). Theil 1. Carl Gerolds Sohn, Wien, pp. 657–674, i–lxxx, pl. 2.
Shamshev, I.V. & Grootaert, P. (2002) A new genus of Microphorinae (Diptera Empidoidea) from New Zealand. Belgian Journal of Entomology, 4, 129–144.
Shamshev, I.V. & Grootaert, P. (2004) Descriptions of four new species of the genus Microphorella Becker (Diptera: Empidoidea, Microphoridae, Parathalassiini) from southeast Asia and New Guinea, with notes on the relationships within the genus. The Raffles Bulletin of Zoology, 52(1), 45–58.
Shamshev, I.V. & Grootaert, P. (2005) Eothalassius, a new genus of parathalassiine flies (Diptera: Empidoidea: Dolichopodidae) from Southeast Asia and Papua New Guinea. European Journal of Entomology, 102, 107–118.
Shatalkin, A.I. (1995) Palaearctic species of Pseudopomyzidae (Diptera). Russian Entomological Journal, 3(3–4) (1994), 129–145.
Sinclair, B.J. (1992) A phylogenetic interpretation of the Brachycera (Diptera) based on the larval mandible and associated mouthpart structures. Systematic Entomology, 17, 233–252.
Sinclair, B.J. (1994) Revision of the Nearctic species of Trichoclinocera Collin (Diptera: Empididae; Clinocerinae). The Canadian Entomologist, 126, 1007–1059.
Sinclair, B.J. (1995a) Generic revision of the Clinocerinae (Empididae), and description and phylogenetic relationships of the Trichopezinae, new status (Diptera: Empidoidea). The Canadian Entomologist, 127, 665–752.
Sinclair, B.J. (1995b) New species of Hormopeza Zetterstedt from South Africa and Tasmania (Diptera: Empididae). Annals of the Natal Museum, 36, 203–208.
Sinclair, B.J. (1996) Revision of the southern Africa genus Acarterus Loew (Diptera: Empidoidea; Hybotinae), with description of seven new species. Annals of the Natal Museum, 37, 215–238.
Sinclair, B.J. (1997) Icasma Collin and an allied new genus, Glyphidopeza from New Zealand (Diptera: Empidoidea; Ceratomerinae). Records of the Australian Museum, 49(2), 195–211.
Sinclair, B.J. (1998) Biosystematic study of the Gondwanan subfamily Ceratomerinae (Empidoidea). In Ismay, J.W. (Ed.), Abstracts Volume. Fourth International Congress of Dipterology. Oxford, England, pp. 205–206.
Sinclair, B.J. (1999) Review of the genera Dipsomyia Bezzi, Zanclotus Wilder, and an allied new Gondwanan genus (Diptera: Empidoidea, Ragas–group). Entomological Science, 2, 131–145.
Sinclair, B.J. (2000a) 1.2. Morphology and terminology of Diptera male terminalia. In Papp, L. & Darvas, B. (Ed.), Contributions to a Manual of Palaearctic Diptera, Volume 1. General and Applied Dipterology. Science Herald, Budapest, pp. 53–74.
Sinclair, B.J. (2000b) Review of the genus Clinocera from Australia and New Zealand (Diptera: Empididae: Clinocerinae). Invertebrate Taxonomy, 14, 347–361.
Sinclair, B.J. (2001) Revision of the world species of Ragas Walker (Diptera: Empidoidea). Entomological Science, 4, 507–522.
Sinclair, B.J. (2002) Revision of the South African endemic genus Edenophorus Smith (Diptera: Empididae). African Invertebrates, 43, 109–122.
Sinclair, B.J. (2003a) Taxonomy, phylogeny and zoogeography of the subfamily Ceratomerinae of Australia (Diptera: Empidoidea). Records of the Australian Museum, 55, 1–44.
Sinclair, B.J. (2003b) Southern African Empidoidea (Diptera) – phylogenetic patterns and biogeographic implications. Cimbebasia, 29, 205–213.
Sinclair, B.J. & Arnaud, P.H., Jr. (2001) Nemedina eocenica new species (Diptera: Empidoidea) from Baltic amber. Myia, 6, 1–8.
Sinclair, B.J. & Cumming, J.M. (1994) Phylogenetic relationships within the Empidoidea (Diptera). In OHara, J.E. (Ed.), Abstract Volume. Third International Congress of Dipterology. Guelph, Canada, pp. 200–201.
Sinclair, B.J. & Cumming, J.M. (1998) The higher–level phylogeny of the Empidoidea (Diptera). In Ismay, J.W. (Ed.), Abstracts Volume. Fourth International Congress of Dipterology. Oxford, England, pp. 207–208.
Sinclair, B.J. & Cumming, J.M. (2000) Revision of the genus Apterodromia (Diptera: Empidoidea), with a redefinition of the tribe Ocydromiini. Records of the Australian Museum, 52, 161–186.
Sinclair, B.J., Cumming, J.M. & Wood, D.M. (1994) Homology and phylogenetic implications of male genitalia in Diptera—Lower Brachycera. Entomologica Scandinavica, 24(4) (1993), 407–432.
Sinclair, B.J. & Harkrider, J.R. (2004) The immature stages and rearing observations of the aquatic dance fly, Roederiodes wirthi Coquillett (Diptera: Empididae: Clinocerinae), with taxonomic notes on the genus. Studia dipterologica, 11, 51–61.
Sinclair, B.J. & Papp, L. (2004) The rediscovery of Nemedina alamirabilis Chandler from Hungary (Diptera: Empidoidea), and first description of the male. Bonner zoologische Beiträge, 52(1–2) (2003), 155–158.
Sinclair, B.J. & Saigusa, T. (2001) Revision of the world species of Ragas Walker (Diptera: Empidoidea). Entomological Science, 4(4), 507–522.
Sinclair, B.J. & Shamshev, I.V. (2003) A new species of Nemedina Chandler from mid–Asia (Diptera: Empidoidea) resolves the phylogenetic position of this enigmatic genus. Journal of Natural History, 37, 2949–2958.
Sivinski, J. (1997) Ornaments in the Diptera. Florida Entomologist, 80, 142–164.
Sivinski, J.M. & Petersson, E. (1997) Mate choice and species isolation in swarming insects. In Choe, J.C. & Crespi, B.J. (Ed.), The Evolution of Mating Systems in Insects and Arachnids. Cambridge University Press, pp. 294–309.
Skevington, J.H. & Yeates, D.K. (2000) Phylogeny of the Syrphoidea (Diptera) inferred from mtDNA sequences and morphology with particular reference to classification of the Pipunculidae (Diptera). Molecular Phylogenetics and Evolution, 16, 212–224.
Skevington, J.H. & Yeates, D.K. (2001) Phylogenetic classification of Eudorylini (Diptera: Pipunculidae). Systematic Entomology, 26, 421–452.
Smith, K.G.V. (1967) Family Empididae (Empidae, Hybotidae). In A Catalogue of the Diptera of the Americas south of the United States. Departamento de Zoologia, São Paulo, pp. 39.1–39.67.
Smith, K.G.V. (1969) The Empididae of southern Africa (Diptera). Annals of the Natal Museum, 19, 1–342.
Smith, K.G.V. (1975) Family Empididae (Empidae, Hybotidae). In Delfinado, M.D. & Hardy, D.E. (Ed.), A Catalog of the Diptera of the Oriental Region, Volume 2. The University of Hawaii Press, Honolulu, pp. 185–211.
Smith, K.G.V. (1980) 32. Family Empididae. In Crosskey, R.W. (Ed.), Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London, pp. 431–442.
Smith, K.G.V. (1989) 43. Family Empididae. In Evenhuis, N.L. (Ed.), Catalog of the Diptera of the Australasian and Oceanic Regions. Bishop Museum Special Publication 86, Honolulu, pp. 382–392.
Snodgrass, R.E. (1922) Mandible substitutes in the Dolichopodidae. Proceedings of the Entomological Society of Washington, 24(6), 148–152.
Snodgrass, R.E. (1943) The feeding apparatus of biting and disease-carrying flies: a wartime contribution to medical entomology. Smithsonian Miscellaneous Collections, 104(1), 1–51 (Publication 3732).
Solórzano Kraemer, M.M., Sinclair, B.J. & Cumming, J.M. (2005) Five new species of Tachydromiinae (Diptera: Empididae s.l.) from New World ambers. Zootaxa, 1010, 37–52.
Sommerman, K.M. (1962) Notes on two species of Oreogeton, predaceous on black fly larvae. Proceedings of the Entomological Society of Washington, 64, 123–129.
Sorenson, M.D. (1999) TreeRot, Version 2. Boston University, Boston, Massachusetts.
Ståhls, G., Hippa, H., Rotheray, G., Muona, J. & Gilbert, F. (2003) Phylogeny of Syrphidae (Diptera) inferred from combined analysis of molecular and morphological characters. Systematic Entomology, 28, 433–450.
Stark, A. (1994) Zum Beutespektrum und Jagdverhalten von Fliegen der Gattung Platypalpus (Empidoidea, Hybotidae). Studia Dipterologica, 1, 49–74.
Steyskal, G.C. & Knutson, L.V. (1981) Empididae [Chapter] 47. In McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Coords.), Manual of Nearctic Diptera, Volume 1. Agriculture Canada Monograph, 27, 607–624.
Stoffolano, J.G., Jr., Woodley, N.E., Borkent, A. & Yin, L.R.S. (1988) Ultrastructural studies of the abdominal plaques of some Diptera. Annals of the Entomological Society of America, 81(3), 503–510.
Stuckenberg, B.R. (1999) Antennal evolution in the Brachycera (Diptera), with a reassessment of terminology relating to the flagellum. Studia Dipterologica, 6, 33–48.
Stuckenberg, B.R. (2001) Pruning the tree: a critical review of classifications of the Homeodactyla (Diptera, Brachycera), with new perspectives and an alternative classification. Studia Dipterologica, 8, 3–41.
Sturtevant, A.H. (1925) The seminal receptacles and accessory glands of the Diptera, with special reference to the Acalypterae. Journal of the New York Entomological Society, 33(4), 195–215; 34(1), 1–21, pls. 1–3.
Svensson, B.G. & Petersson, E. (1987) Sex-role reversed courtship behaviour, sexual dimorphism and nuptial gifts in the dance fly, Empis borealis (L.). Annales Zoologici Fennici, 24, 323–334.
Swofford, D.L. (2002) PAUP*. Phylogenetic Analysis Using Parsimony (*and Other Methods), Version 4.0b10. Sinauer Associates, Sunderland, Massachusetts.
Thompson, F.C. (1969) A new genus of microdontine flies (Diptera: Syrphidae) with notes on the placement of the subfamily. Psyche, 76, 74–85.
Thompson, F.C. (Ed.) (2005) Biosystematic Database of World Diptera. http://www.diptera.org/Diptera/biosys.htm, accessed on December 1, 2005.
Trehen, P. (1961) Contribution à l’étude morphologique des segments génitaux du mâle d’Hilara maura Fab. (Diptères Empidides). Bulletin de la Société Scientifique de Bretagne, 35, 285–295.
Trehen, P. (1969) Description des stades préimaginaux et données sur la biologie de Phyllodromia melanocephala Fabricius, 1794 (Diptères-Empididae). Revue d’Écologie et de Biologie du Sol, 6, 41–52.
Tuomikoski, R. (1952) Über die Nahrung der Empididen–Imagines (Dipt.) in Finnland. Annales Entomologici Fennici, 18(4), 170–181.
Tuomikoski , R. (1960) Mitteilungen über die Empididen (Dipt.) Finnlands VII. Die Gattung Hormopeza Zett. Annales Entomologici Fennici, 26, 99–107.
Tuomikoski, R. (1966) The Ocydromiinae group of subfamilies (Diptera, Empididae). Annales Entomologici Fennici, 32, 282–294.
Ulrich, H. (1971) Zur Skelett- und Muskelanatomie des Thorax der Dolichopodiden und Empididen (Diptera). Veröffentlichungen der Zoologischen Staatsammlung München, 15, 1–44, 13 pls.
Ulrich, H. (1974) Das Hypopygium der Dolichopodiden (Diptera): Homologie und Grundplanmerkale. Bonner Zoologische Monographien, 5, 1–60.
Ulrich, H. (1975) Das Hypopygium von Chelifera precabunda Collin (Diptera, Empididae). Bonner Zoologische Beiträge, 26, 264–279.
Ulrich, H. (1976) Morphology of the dolichopodid hypopygium and its significance for a phylogenetic system of that family (Diptera). An unpublished paper read at the XV International Congress of Entomology, Washington, DC, USA.
Ulrich, H. (1981) Trichopeza longicornis (Meigen) (Diptera, Empididae) hat Raubbeine. Bonner Zoologische Beiträge, 32, 237–239.
Ulrich, H. (1984) Skelett und Muskulatur des Thorax von Microphor holosericeus (Meigen) (Diptera, Empidoidea). Bonner Zoologische Beiträge, 35, 351–398.
Ulrich, H. (1988) Das Hypopygium von Microphor holosericeus (Meigen) (Diptera, Empidoidea). Bonner Zoologische Beiträge, 39, 179–219.
Ulrich, H. (1990) Evidence for the phylogenetic position of Parathalassiinae (Empidoidea) based on thoracic morphology. An unpublished paper read at the 2nd International Congress of Dipterology. See abstract in: Abstract Volume, Second International Congress of Dipterology, Bratislava, Czechoslovakia, p. 245.
Ulrich, H. (1991) Two new genera of parathalassiine-like flies from South Africa (Diptera, Empidoidea). Bonner Zoologische Beiträge, 42, 187–216.
Ulrich, H. (1994) A likely autapomorphy of the Empididae, and evidence for the systematic position of Edenophorus Smith. An unpublished paper read to the 3rd International Congress of Dipterology. See abstract in: OHara, J.E. (Ed.), Abstract Volume, Third International Congress of Dipterology, Guelph, Canada, p. 230.
Ulrich, H (1996) Proposals for the phylogenetic classification of the Empidoidea, or Orthogenya (Diptera, Eremoneura). An unpublished paper read at the 20th International Congress of Entomology. See abstract in: XX International Congress of Entomology, Proceedings. Florence, Italy, p. 27.
Ulrich, H. (2003) How recent are the Empidoidea of Baltic amber? Studia Dipterologica, 10, 321–327.
Ulrich, H. (2004) Phylogenetic considerations about an early colonization of the sea coasts by Dolichopodidae (Diptera). Studia Dipterologica, 11, 233–243.
Ulrich, H. (2005) Predation by adult Dolichopodidae (Diptera): a review of literature with an annotated prey-predator list. Studia Dipterologica, 11 (2004), 269–403.
Vaillant, F. (1952) Un empidide destructeur de simulies. Bulletin de la Société Zoologique de France, (1951) 76, 371–379.
Vaillant, F. (1953) Hemerodromia seguyi, nouvel empidide d’Algérie destructeur de simulies. Hydrobiologia, 5, 180–188.
Vaillant, F. (1981) Diptères Empididae Hemerodromiinae nouveaux ou peu connus de la région paléarctique (Première partie). Bonner Zoologische Beiträge, 32, 351–408.
Wagner, R. & Gathmann, O. (1996) Long-term studies on aquatic dance flies (Diptera, Empididae) 1983–1993: Distribution and size patterns along the stream, abundance changes between years and the influence of environmental factors on the community. Archiv für Hydrobiologie, 137, 385–410.
Walker, F. (1837) Notes on Diptera. The Entomological Magazine, 4 (1836), 226–230.
Werner, D. & Pont, A.C. (2003) Dipteran predators of simuliid blackflies: a worldwide review. Medical and Veterinary Entomology, 17, 115–132.
Wheeler, W.M. & Melander, A.L. (1901) Empidae. Biologia Centrali-Americana. Insecta. Diptera, 1 (Supplement), 366–376.
Wiegmann, B.M. (1989) A phylogenetic revision of the family Atelestidae (Diptera: Empidoidea) and its implications for the origin of the cyclorrhaphous Diptera. M.Sc. thesis. University of Maryland, Washington, 205 pp.
Wiegmann, B.M., Mitter, C. & Thompson, F.C. (1993) Evolutionary origin of the Cyclorrhapha (Diptera): Tests of alternative morphological hypotheses. Cladistics, 9, 41–81.
Wiegmann, B.M., Yeates, D.K., Thorne, J.L. & Kishino, H. (2003) Time flies, a new molecular time-scale for Brachyceran fly evolution without a clock. Systematic Biology, 52, 745–756.
Wilder, D.D. (1981) A revision of the genus Niphogenia Melander (Diptera: Empididae). Pan-Pacific Entomologist, 57, 422–428.
Winterton, S.L., Yang, L., Wiegmann, B.M. & Yeates, D.K. (2001) Phylogenetic revision of the Agapophytinae subfam. n. (Diptera: Therevidae) based on molecular and morphological evidence. Systematic Entomology, 26, 173–211.
Woodley, N.E. (1989) Phylogeny and classification of the “Orthorrhaphous” Brachycera [Chapter] 115. In McAlpine, J.F. (Ed.), Manual of Nearctic Diptera, Volume 3. Agriculture Canada Monograph, 32, 1371–1395.
Yang, D. Gaimari, S.D. & Grootaert, P. (2004) A new genus and species of Tachydromiinae (Diptera: Empididae) from the Oriental realm. Transactions of the American Entomological Society, 130, 489–492.
Yang, D. & Yang, C. (2004) Diptera Empididae: Hemerodromiinae, Hybotinae. Fauna Sinica, Insecta, 34, i–x, 1–335, pl. 1.
Yang, L., Wiegmann, B.M., Yeates, D.K. & Irwin, M.E. (2000) Higher-level phylogeny of the Therevidae (Diptera: Insecta) based on 28S ribosomal and elongation factor-1a gene sequences. Molecular Phylogenetics and Evolution, 15(3), 440–451.
Yeates, D.K. (1992) Towards a monophyletic Bombyliidae (Diptera): the removal of the Proratinae (Diptera: Scenopinidae). American Museum Novitates, 3051, 1–30.
Yeates, D.K. (1994) The cladistics and classification of the Bombyliidae (Diptera: Asiloidea). Bulletin of the American Museum of Natural History, 219, 1–191.
Yeates, D.K. (1996) Groundplans and exemplars: paths to the tree of life. Cladistics, 11, 343–357.
Yeates, D.K. (2002) Relationships of extant lower Brachycera (Diptera): a quantitative synthesis of morphological characters. Zoologica Scripta, 31, 105–121.
Yeates, D.K. & Wiegmann, B.M. (1999) Congruence and controversy: toward a higher-level phylogeny of Diptera. Annual Review of Entomology, 44, 397–428.
Young, J.H. & Merritt, D.J. (2003) The ultrastructure and function of the silk-producing basitarsus in the Hilarini (Diptera: Empididae). Arthropod Structure and Development, 32, 157–165.
Zatwarnicki, T. (1996) A new reconstruction of the origin of eremoneuran hypopygium and its implications for classification (Insecta: Diptera). Genus, 7(1), 103–175.
Zimmer, M, Diestelhorst, O. & Lunau, K. (2003) Courtship in long-legged flies (Diptera: Dolichopodidae): function and evolution of signals. Behavioral Ecology, 14, 526–530.