Abstract
The Neotropical genus Engelimyia Lopes, 1975 is revised and found to consist of four species: the already named E. bosqi (Blanchard, 1939) and E. inops (Walker, 1849), with the former nominal species herewith resurrected from synonymy with the latter; and two new species, E. qibos sp. nov. and E. psino sp. nov. A lectotype is designated for Sarcophaga cassidifera Engel, 1931 [= E. inops (Walker, 1849)]. Male terminalia of all species are illustrated, and a key to the species is provided. The four species appear to form two sister-group pairs: E. inops + E. psino and E. bosqi + E. qibos, both of which show a South American—Central American Isthmian disjunction, which here is considered to reflect the same vicariance event and a maximum age of the four species of about 3 My.
References
Aldrich, J.M. (1916) Sarcophaga and allies in North America. Thomas Say Publications, 1, 1–301 + [1 (postscript)] + [3 (index)], pls. 1–16.
Arnaud, P.H., Jr. (1958) The entomological publications of Charles Henry Tyler Townsend [1863–1944]; with lists of his new generic and specific names. Microentomology, 23, 1–63.
Blackith, R.M., Blackith, R.E. & Pape, T. (1998) Taxonomy and systematics of the taxon Helicophagella Enderlein, 1928 (Diptera: Sarcophagidae), with the description of a new species. Studia dipterologica, 4, 383–434. [1997]
Blanchard, E.E. (1939) Los sarcofágidos argentinos. Contribución a su conocimiento. Physis, 17, 791–856.
Coates, A.G. & Obando, J.A. (1996) The geologic evolution of the Central American Isthmus. In: Jackson, J.B.C., Budd, A.F. & Coates, A.C. (Eds), Evolution and environment in tropical America. University of Chicago Press, pp. 25–61.
Dodge, H.R. (1968) The Sarcophagidae of Barro Colorado Island, Panama (Diptera). Annals of the Entomological Society of America, 61, 421–450.
Enderlein, G. (1928) Klassifikation der Sarcophagiden. Sarcophagiden-Studien I. Archiv für Klassifikatorische und Phylogenetische Entomologie, 1, 1–56.
Engel, E.O. (1931) Die Ausbeute der deutschen Chaco-Expedition 1925/26.—Diptera. XXVI. Anthomyidae, XXVII. Muscidae und XXVIII. Sarcophagidae. Konowia, 10, 133–154.
Evenhuis, N.L. (1994). Catalogue of the fossil flies of the world (Insecta: Diptera). Backhuys Publishers, Leiden, 600 pp.
Hall, D.G. (1933) The Sarcophaginae of Panama (Diptera: Calliphoridae). Bulletin of the American Museum of Natural History, 66, 251–285.
Kurahashi, H. & Kano, R. (1984) Phylogeny and geographical distribution of the genus Boettcherisca Rohdendorf (Diptera: Sarcophagidae). Japanese Journal of Medical Science and Biology, 37, 27–34.
Lopes, H.S. (1935) Algumas espécies de Sarcophaga do Brasil. Revista de Entomologia, 5, 38–46.
Lopes, H.S. (1939) Sôbre alguns sarcophagídeos de Misiones, Argentina. Physis, 17, 117–123.
Lopes, H.S. (1941) Sôbre alguns sarcofagídeos neotrópicos da coleção do Museu Britânico (Diptera). Arquivos de Zoologia do Estado de São Paulo, 2, 357–387.
Lopes, H.S. (1969) Family Sarcophagidae. In: Papavero, N. (Ed.), A catalog of the Diptera of the Americas south of the United States, 103, 1–88. Departamento de Zoologia, Secretaria da Agricultura, São Paulo.
Lopes, H.S. (1973) Collecting and rearing sarcophagid flies (Diptera) in Brazil, during 40 years. Anais da Academia Brasileira de Ciências, 55, 279–291.
Lopes, H.S. (1975) On Engelimyia, new genus for Sarcophaga cassidifera Engel (Diptera, Sarcophagidae). Revista Brasileira de Biologia, 34, 425–430. [1974]
Lopes, H.S. (1976) On the holotypes, mostly females, of some Sarcophagidae described by Francis Walker. Revista Brasileira de Biologia, 36, 629–641.
Lopes, H.S. (1983) The importance of the mandible and clypeal arch of the first instar larvae in the classification of the Sarcophagidae (Diptera). Revista Brasileira de Entomologia, 26, 293–326. [1982]
Lopes, H.S. (1984) A tentative arrangement of the Notochaetina (Diptera, Sarcophagidae), a contribution to the phylogeny of the group. Anais da Academia Brasileira de Ciências, 56, 339–350.
Lopes, H.S. (1990) On the genera of Sarcophagidae (Diptera) showing proclinate frontorbital bristles in males. Revista brasileira de Biología, 50, 279–292.
Lopes, H.S. & Tibana, R. (1982) Sarcophagid flies (Diptera) from Sinop, State of Mato Grosso, Brazil. Memórias do Instituto Oswaldo Cruz, 77, 285–298.
Mello-Patiu, C.A. & Pape T. (2000) Definitions of Dexosarcophaga Townsend, 1917 and Sarcofahrtiopsis Hall, 1933, including two new species and a redescription of Sarcofahrtiopsis cuneata (Townsend, 1935) (Diptera: Sarcophagidae). Boletín de Entomología Venezolana, 15, 181–194.
Merz, B. & Haenni, J.-P. (2000) Morphology and terminology of adult Diptera (other than terminalia). In: Papp, L. & Darvas, B. (Eds), Contributions to a manual of Palaearctic Diptera (with special reference to flies of economic importance). Science Herald Press, Budapest, pp. 21–51.
Pape, T. (1994) The world Blaesoxipha Loew, 1861 (Diptera: Sarcophagidae). Entomologica scandinavica, Supplement, 45, 1–247.
Pape, T. (1995) A catalogue of the Sarcophagidae (Insecta: Diptera) described by G. Enderlein. Steenstrupia, 21, 1–30.
Pape, T. (1996) Catalogue of the Sarcophagidae of the world (Insecta: Diptera). Memoirs on Entomology, International, 8, 1–558.
Pape, T. (2006) A new species of Hoplacephala Macquart (Diptera: Sarcophagidae) from Namibia, with a discussion of generic monophyly. Zootaxa, 1183, 57–68.
Roubik, D.W., Segura, J.A.L. & Camaro, J.M.F. de (1997) New stingless bee genus endemic to Central American cloudforests: phylogenetic and biogeographic implications (Hymenoptera: Apidae: Meliponini). Systematic entomology, 22, 67–80.
Sinclair, B.J. (2000) Morphology and terminology of Diptera male terminalia. In: Papp, L. & Darvas, B. (Eds), Contributions to a manual of Palaearctic Diptera (with special reference to flies of economic importance). Science Herald Press, Budapest, pp. 53–74.
Shewell, G.E. 1987. Sarcophagidae. In: McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Eds), Manual of Nearctic Diptera. Volume 2. Agriculture Canada Monograph, 28, 1159–1186.
Thompson, F.C. (ed.) (2005) Biosystematic Database of World Diptera, Version 7.5. Available from http://www.sel.barc.usda.gov/diptera/biosys.htm (last update of nomenclator: 24 October 2005; accessed 20 March 2006).
Townsend, C.H.T. (1917) Genera of the dipterous tribe Sarcophagini. Proceedings of the Biological Society of Washington, 30, 189–197.
Walker, F. (1849) List of the specimens of dipterous insects in the collection of the British Museum. Part IV, 689–1172. British Museum (Natural History), London.
Wells, J.D., Pape, T. & Sperling, F.A.H. (2001) DNA-based identification and molecular systematics of forensically important Sarcophagidae (Diptera). Journal of Forensic Sciences, 46, 1098–1102.
Wood, D.M. (1987) Tachinidae. In: McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Eds), Manual of Nearctic Diptera. Volume 2. Agriculture Canada Monograph, 28, 1193–1269.
Zamudio, K.R. & Greene, H.W. (1997) Phylogeography of the bushmaster (Lachesis muta: Viperidae): implications for neotropical biogeography, systematics, and conservation. Biological Journal of the Linnean Society, 62, 421–442.
Zumpt, F. (1953) Notes on Enderlein’s types of Sarcophaginae from the Ethiopian and Madagascan regions with remarks on his system of classification. Anais do Instituto de medicina Tropical, 10, 15–23.