Abstract
As part of a project intended to review the taxa of Marginellidae worldwide, the phylogenetic relationships of its main representatives are provided based on comparative phenotypy. Characters from most structures and organs are investigated and used for a phylogenetic analysis, resulting in the following cladogram: ((((Marginellona gigas ((Eratoidea watsoni ((Prunum sp—Leptegouana guttata) (Volvarina brasiliana (Prunum prunum—P. rubens)))) (Austroginella muscaria (Marginella ealesae ((Marginella rosea (M. glabella—M. sebastiani)) (Dentimargo bruneolus (Eratoidea scalaris (Dentimargo sp—D. aureocincta)))))))) Pachycymbiola brasiliana*) Persicula sagitatta*) Buccinanops gradatus*). Those marked with * are outgroups functionally analyzed as part of the ingroup (respectively a volutid, a cystiscid and a nassariid); the root is based on Trophon geversianus (Muricidae). The genera Prunum, Marginella and Dentimargo are revealed as non-monophyletic. The monophyly of the family Marginellidae is supported by 17 synapomorphies. The volutids appear to be its sister taxon, and the possibility of Marginellidae being only a branch of Volutidae is discussed.
References
Amitrov, O.V. (2010) Gastropods of rare families from the Tshegan Formation of Kazakhstan. Paleontological Journal, 44 (4), 24–29.
https://doi.org/10.1134/S0031030110040039
Barnard, K.H. (1962) A new genus in the family Marginellidae. Proceedings of the Malacological Society of London, 35 (1), 14–15.
Baugh, T.M. (1990) Note on the feeding behavior of the common Atlantic marginella Prunum apicinum (Gastropoda, Marginellidae). Florida Scientist, 53, 50–51.
Boss, K.J. (1982) Mollusca. In: Parker, S.P. (Ed.), Synopsis and Classification of Living Organisms, vol. 1. McGraw-Hill, New York, pp. 945–1166.
Bouchet, P. (1989) A marginellid gastropod parasitizes sleeping fishes. Bulletin of Marine Science, 45, 76–84.
Bouchet, P., Rocroi, J.P., Hausdorf, B., Kaim, A., Kano, Y. Nützel, A., Parkhaev, P., Schrödl, M. & Strong, E.E. (2017) Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia, 61, 1–526.
https://doi.org/10.4002/040.061.0201
Bouvier, E.L. (1889) Sur le siphon oesophagien des marginelles. Bulletin de la Societé Philomatique de Paris, 1 (1), 13–14.
Boyer, F. (2000) The genus Volvarina (Volutacea: Marginellidae) in Brazil. Part 1: revision of the species described by A. Bavay, and closely related species. Novapex, 1 (2), 35–55
Boyer, F. (2004) Description of a new Cystiscus (Gastropoda: Cystiscidae) from the Mascarene Islands. Journal of Conchology, 38 (4), 355–361.
Boyer, F. (2015a) Revision des marginelles de Linne (mollusques prosobranches: Marginellidae et Cystiscidae). Xenophora, Supplement 8, 33–55.
Boyer, F. (2015b) Sur quelques Volvarina (Marginellidae) de l’ocean indien occidental. Xenophora, Supplement 6, 19–31.
Calzada, S. & Corbacho, J. (2013) Problema taxonómico entre los géneros Pterigia y Marginella (Gasteropoda). Scripta Musei Geologici Seminarii Barcinonensis Series Palaeontologica, 13, 23–26.
Carpenter, J.M. (1988) Choosing among equally parsimonious cladograms. Cladistics, 4, 291–296.
https://doi.org/10.1111/j.1096-0031.1988.tb00476.x
Carriker, M.R. (1943) On the structure and function of the proboscis in the common oyster drill, Urisalpinx cinerea Say. Journal of Morphology, 73, 441–506.
https://doi.org/10.1002/jmor.1050730304
Clarke Jr, A.H. (1962) Annotated list and bibliography of the abyssal marine molluscs of the world. Bulletin of the National Museum of Canada, 181, 1–114.
Clench, W.J. & Turner R.D. (1964) The subfamilies Volutinae, Zidoninae, Odontocymbiolinae, and Calliotectinae in the Western Atlantic. Johnsonia, 4 (43), 129–80.
Coan, E.V. (1965) A proposed reclassification of the family Marginellidae. Veliger, 7 (3), 184–194.
Coan, E.V. & Roth, B. (1976) Status of the genus Hyalina Schumacher, 1817 (Mollusca: Gastropoda). Journal of Molluscan Studies, 42, 217–222.
Coovert, G.A. (1986a) Type species list. Marginella Marginalia, 1 (1), 1–3.
Coovert, G.A. (1986b) Generic group names. Marginella Marginalia, 1 (2), 5–7.
Coovert, G.A. (1986c) Two previously unfigured species from Dall. Marginella Marginalia, 1 (3), 8–12.
Coovert, G.A. (1986d) A review of marginellid egg capsule. Marginella Marginalia, 1 (4), 13–19.
Coovert, G.A. (1986e) Notes on the genus Cystiscus and its type species. Marginella Marginalia, 1 (5), 20–24.
Coovert, G.A. (1986f) Notes on the genus Crithe and its type species. Marginella Marginalia, 1 (1), 1–3.
Coovert, G.A. (1987a) The genus Afrivoluta and a review of the type species Afrivoluta pringlei Tomlin, 1947. Marginella Marginalia, 2 (1), 1–8.
Coovert, G.A. (1987b) The Persicula cornea group. Marginella Marginalia, 2(2/3), 10–19.
Coovert, G.A. (1987c) Additional notes on the genus Crithe. Marginella Marginalia, 2 (5), 28–35.
Coovert, G.A. (1987d) Extra extra: read all about it. Marginella Marginalia, 2 (6), 36–41.
Coovert, G.A. (1987e) Proper usage of the generic names Haloginella, Hyalina and Volvarina with a diagnosis of the genus Haloginella. Marginella Marginalia, 3 (1), 1–7.
Coovert, G.A. (1987f) A literature review and summary of marginellid external anatomy. Marginella Marginalia, 3 (2/3), 8–23.
Coovert, G.A. (1987g) Notes on the genus Granulina and its type species. Marginella Marginalia, 3 (4), 26–32.
Coovert, G.A. (1987h) The external anatomy of two species of Caribbean Marginellidae. Marginella Marginalia, 3 (5), 33–37.
Coovert, G. A. (1987i) Taxonomy: how and why we do what we do. Marginella Marginalia, 3 (6), 38–42.
Coovert, G.A. (1988a) Marginellidae of Florida, Part I: Granulina hadria. Marginella Marginalia, 4 (1), 1–8.
Coovert, G.A. (1988b) Type species of the genus Austroginella and Mesoginella and their synonyms. Marginella Marginalia, 4 (2/3), 9–26.
Coovert, G.A. (1988c) Marginellas from sunwatch, a prehistoric indian village in SW Ohio. Marginella Marginalia, 4 (4), 27–34.
Coovert, G.A. (1988d) Marginellidae of Florida, Part II: Prunum succinea with a discussion of Prunum and Volvarina. Marginella Marginalia, 4 (5), 35–42.
Coovert, G.A. (1988e) Taxonomic characters in the family Marginellidae: conchological characters. Marginella Marginalia, 4 (68), 43–47.
Coovert, G.A. (1988f) A bibliography of the recent marginellidae. Marginella Marginalia, 5 (1–5), 1–43.
Coovert, G.A. (1989a) The Marginella musica—diacocus group. Marginella Marginalia, 6 (3–5), 16–29.
Coovert, G.A. (1989b) A literature review and summary of published marginellid radulae. Marginella Marginalia, 7 (1–6), 1–37.
Coovert, G.A. & Coovert, H.K. (1990) A Study of marginellid radulae part I: type 6 radula, “Prunum/Volvarina” type. Marginella Marginalia, 8–9 (1–6), 1–68.
Coovert, G.A. & Coovert, H.K. (1995) Revision of the supraspecific classification of marginelliform gastropods. Nautilus, 109(2–3), 43–110.
Cossignani, T. (2006a) Tre nuove Volvarine dal W. Atlantico (Gastropoda: Prosobranchia, Marginellidae). Malacologia Mostra Mondiale, 50, 15–17.
Cossignani, T. (2006b) Marginellidae & Cystiscidae of the World. L’Informatore Piceno, 408 pp.
Cossignani, T. (2007a) Tre nuove marginelle dal Brasile (Gastropoda: Prosobranchia, Marginellidae). Malacologia Mostra Mondiale, 53, 12–15.
Cossignani, T. (2007b) Una nuova Volvarina dal Brasile (Gastropoda: Prosobranchia, Marginellidae). Malacologia Mostra Mondiale, 54, 14–15.
Cossmann, A.E.M. (1899) Essais de Paléoconchologie Comparé. Vol. 3. Paris, 201 pp., 8 pl.
Cotton, B.C. (1949) Australian Recent and Tertiary Mollusca, Family Marginellidae. Records of the South Australian Museum, 9, 197–224.
D’Asaro, C.N. (1970) Egg capsules of prosobranch mollusks from South Florida and the Bahamas and notes on spawning in the laboratory. Bulletin of Marine Science, 20, 414–440.
Davis, G.M. (1979) The origin and evolution of the gastropod family Pomatiopsidae, with emphasis on the Mekong River Triculinae. Monographs of the Academy of Natural Sciences of Philadelphia, 20, 1–120.
Dell, R.K. (1990) Antarctic Mollusca, with special reference to the fauna of the Ross Sea. Bulletin of the Royal Society of New Zealand, 27, 1–311.
Djajasasmith, M. & Coomans, H.E. (1980) Notes on Rivomarginella electrum (Reeve, 1865) from Lampung, south Sumatra (Gastropoda, Marginellidae). Bulletin Zoologisch Museum Universiteit van Amsterdam, 711, 117–122.
Eames, F.E. (1952) A contribution to the study of the Eocene in Western Pakistan and Western India. C. The description of the Scaphopoda and Gastropoda from standard sections in the Rakhi Nala and Zinda Pir areas of the Western Punjab and in the Koahat District. Philosophical Transaction of the Royal Society of London, Series B, 236 (631), 1–168.
https://doi.org/10.1098/rstb.1952.0001
Espinosa, J. & Ortea, J. (2000a) Descripción de un genero y once especies nuevas de Cystiscidae y Marginellidae (Mollusca: Neogastropoda) del Caribe de Costa Rica. Avicennia, 12–13, 95–114.
Espinosa, J. & Ortea, J. (2000b) Datos anatomicos de Prunum holandae (Mollusca: Neogastropoda: Marginellidae). Avicennia, 12–13, 91–94.
Espinosa, J & Ortea, J. (2013a) Nuevas espécies de los gêneros Dentimargo Cossmann, 1899 y Eratoidea Weinkauff, 1879, y nuevo gênero de marginellido de la Isla de la Guadeloupe, Antillas Menores, Mar Caribe (Mollusca: Neogastropoda: Marginellidae). Revista de la Academia Canaria de Ciencias, 25, 111–127.
Espinosa, J & Ortea, J. (2013b) Nuevas especies de la família Marginellidae (Mollusca: Gastropoda: Prosobranchia) de cuatro islas del caribe: Cuba, Curazao, Guadalupe y Martinica. Revista de la Academia Canaria de Ciencias, 25, 195–218.
Farris, J.S. (1969) A successive approximations approach to character weighting. Systematic Zoology, 18, 374–385.
https://doi.org/10.2307/2412182
Farris, J.S. (1988) Hennig 86, computer program. Version 1.5.
Fehse, D. (2015) On the identity of Hespererato pallida Oleinik, Petuch & Aley, 2012 (Mollusca: Trivioidea: Eratoidae). Visaya, 4 (4), 19–20.
Fleming, J. (1828) A History of British Animals, Exhibiting the Descriptive Characters and Systematical Arrangement of the Genera and Species of Quadrupeds, Birds, Reptiles, Fishes, Mollusca and Radiata of the United Kingdom. Edinburg and London, 565 pp.
https://doi.org/10.5962/bhl.title.12859
Fretter, V. (1976) The anatomy and feeding of the volutacean prosobranch Volvarina taeniolata Mörch. Journal of Molluscan Studies, 42, 327–336.
https://doi.org/10.1093/oxfordjournals.mollus.a065341
Gofas, S. (1989) Le genre Volvarina (Marginellidae) dans la Méditerranée et l’Atlantique du Nord Est. Bolletino Malacologico, 25 (5–6), 1–25.
Gofas, S. & Fernandes, F. (1988) The Marginellids of São Tomé, West Africa. Journal of Conchology, 33 (1), 1–30.
Gofas, S. & Fernandes, F. (1992) The Marginellidae of Angola: the genus Volvarina. Journal of Conchology, 34 (4), 187–198.
Gofas, S. & Fernandes, F. (1994) The Marginellidae of Angola: the genus Marginella. Journal of Conchology, 35 (2), 103–119.
Golding, R.E., Ponder, W.F. & Byrne, M. (2009) The evolutionary and biomechanical implications of snout and proboscis morphology in Caenogastropoda (Mollusca: Gastropoda). Journal of Natural History, 43 (43–44), 2723–2763.
https://doi.org/10.1080/00222930903219954
Gracia, A. & Boyer, F. (2004) Distribution of Volvarina hennequini Boyer, 2001 in the Western Caribbean Sea and description of a new bathyal Volvarina from Northern Colombia. Novapex, 5, 79–84.
Graham, A. (1966) The foregut of some marginellid and cancellariid prosobranchs. Studies in Tropical Oceanography, 4, 134–151.
Gray, J.E. (1857) Guide to the systematic distribution of Mollusca in the British Museum. London, 12, 230 pp.
Griffiths, H.J., Linse, K. & Crame, J.A. (2003) SOMBASE—Southern Ocean mollusc database: a tool for biogeographic analysis in diversity and evolution. Organisms Diversity and Evolution, 3, 207–213.
https://doi.org/10.1078/1439-6092-00079
Harasewych, M.G. (1984) Comparative anatomy of four primitive muricacean gastropods: implication for trophonine phylogeny. American Malacological Bulletin, 3 (1), 11–26.
Harasewych, M.G. & Kantor, Y.I. (1991) Tenebrincola frigida, a new genus and species of abyssal volute from the Northern Pacific Ocean. Nemouria, 38, 1–11.
Harasewych, M.G., Adamkewicz, S.L., Blake, J.A., Saudek, D., Springgs, T. & Bult, C.J. (1997) Neogastropod phylogeny: a molecular perspective. Journal of Molluscan Studies, 63, 327–351.
https://doi.org/10.1093/mollus/63.3.327
Hinds, R.B. (1844) Descriptions of Marginellidae collected during the Voyage of H. M. S. Sulphur, and from the collection of Mr. Cuming. Proceedings of the Zoological Society of London, 12 (134), 72–77.
Jousseaume, J.P. (1875) Coquilles de la Famille des Marginelles. Monographie. Revue et Magazine de Zoologie, 3, 164–271.
https://doi.org/10.5962/bhl.title.10780
Kantor, Y.I. (1996) Phylogeny and relationships of Neogastropoda, pp. 221–230. In: Taylor, J.D. (Ed.), Origin and Evolutionary Radiation of the Mollusca. Oxford University Press, Oxford, 392 pp.
Kiener, L.C. (1834–1841) Le Genre Marginelle in Spécies Général et Iconographie des Coquilles Vivants. J. B. Baillière et fils. Paris, 3, 1–30 (1834), 31–44 (1841).
Lamarck, J.B.P.A.M. (1799) Prodome d’une Nouvelle Classification des Coquilles. Mémoire de la Societé Histoire Naturelle de Paris, 1, 63–91.
Landau, B., Frydman, F. & Silva, C.M. (2012) A new species of Harpa (Gastropoda: Volutoidea) from the Neogene of the Dominican Republic: Paleobiogeographical implications. Veliger, 51, 131–136.
Laseron, C.F. (1957) A new classification of the Australian Marginellidae (Mollusca), with a review of species from the Solanderian and Damperian Zoogeographical Provinces. Records of the Australian Museum, 8 (3), 274–311.
https://doi.org/10.1071/MF9570274
Magaña, J., Espinosa, J. & Ortea, J. (2003) Descripción de dos nuevas especies del género Prunum Herrmannsen, 1852 (Mollusca: Gastropoda: Marginellidae) del Caribe y el Pacífico de Costa Rica. Avicennia, 16, 121–128.
Marcus, E. & Marcus, E. (1968) On the prosobranchs Ancilla dimidiata and Marginella fraterculus. Proceedings of the Malacological Society of London, 38, 55–69.
Maury, C.J. (1925) Fósseis Terciários do Brasil, com descripção de novas formas Cretáceas. Monografia do Serviço Geológico e Mineralógico, 4, 1–665., 24 pls.
McCleery, T. (2011) A revision of the genus Eratoidea Weinkauff, 1879 (Gastropoda: Marginellidae). Novapex, 8, 1–111.
Modica, M.V. & Holford, M. (2010) The Neogastropoda: evolutionary innovations of predatory marine snails with remarkable pharmacological potential. In: Pontarotti, P. (Ed.), Evolutionary Biology—Concepts, molecular and morphological evolution. Spring-Verlag Berlin Heidelberg, Berlin, 249–270.
https://doi.org/10.1007/978-3-642-12340-5_15
Murray, F.V. (1970) The reproduction and life history of Microginella minutissima (Tenison-Woods, 1876) (Gastropoda: Marginellidae). Memoirs of the National Museum of Victoria, 31, 31–35.
https://doi.org/10.24199/j.mmv.1970.31.04
Nehm, R.H. (2001) Calibrating spatial and temporal species richness in tropical American marginellid gastropods. Journal of Paleontology, 75 (3), 680–696.
https://doi.org/10.1017/S002233600003972X
Nehm, R.H. (2005) Patterns and processes of evolutionary stasis and change in Eratoidea (Gastropoda: Marginellidae) from the Dominican Republic Neogene. Caribbean Journal of Science, 41 (2), 189–214.
Numanami, H. (1996) Taxonomic study on Antarctic gastropods collected by Japanese Antarctic research expeditions. Memoirs of National Institute of Polar Research, Series E (Biology and Medical Science), 39, 1–244.
Orlin, Z. (2007) More on blood-sucking mollusk. American Conchologist, 35 (2), 23–24.
Penchaszadeh, P.E. & Rincon, A. (1996) Egg capsules and development of Prunum prunum (Gmelin, 1791) (Prosobranchia: Marginellidae) from the Venezuela Caribbean. Veliger, 39 (1), 83–86.
Ponder, W.F. (1970) Some aspects of the morphology of four species of the neogastropod family Marginellidae with a discussion on the evolution of the toxoglossan poison gland. Journal of the Malacological Society of Australasia, 2 (1), 55–81.
https://doi.org/10.1080/00852988.1970.10673841
Ponder, W.F. (1973) The origin and evolution of the Neogastropoda. Malacologia, 12 (2), 295–338.
Ponder, W.F. (1998) Family Marginellidae. In: Beesley, P.L., Ross, G.J.B. & Wells, A. (Eds.), Mollusca: The Southern Synthesis. Fauna of Australia. CSIRO Publishing. Melbourne, 5 B, pp. 838–841.
Ponder, W.F. & Lindberg, D.R. (1997) Towards a phylogeny of gastropod molluscs—a preliminary analysis using morphological characters. Zoological Journal of the Linnean Society of London, 119, 83–265.
https://doi.org/10.1111/j.1096-3642.1997.tb00137.x
Ponder, W.F. & Taylor, J.D. (1992) Predatory shell drilling by two species of Austroginella (Gastropoda: Marginellidae). Journal of Zoology, 228, 317–328.
https://doi.org/10.1111/j.1469-7998.1992.tb04611.x
Price, R.M. (2003) Columellar muscle of neogastropods: muscle attachment and the function of columellar folds. Biological Bulletin, 205, 351–366.
https://doi.org/10.2307/1543298
Raeihle, D. (1965) Studies of captive Prunum apicinum Menke. Annual Reports of the American Malacological Union, 32, 21–22.
Riedel, F. (2000) Ursprung und Evolution der “höheren” Caenogastropoda. Berliner Geowissenschaftiche Abhandlungen E32, 1–240, 21 pls.
Rios, E.C. (2009) Compendium of Brazilian seashells. Universidade Federal do Rio Grande. Rio Grande, 668 pp.
Rios, E.C. & Leal, J.H. (1993) Volvarina pontesi, a new bathyal marginellid (Mollusca, Gastropoda) from off Brazil. Apex, 8, 27–30.
Robertson, R. (1996) Fargoa bartschi Winkley, 1909: a little-known Atlantic and Gulf coast American odostomian Pyramidellidae, and its generic relationships. American Malacological Bulletin, 13, 11–21.
Rolán, E. & Gori, S. (2014) New information on the marginellids of Sao Tome and Principe, with new records and the description of four new species. Iberus, 32 (1), 1–25.
Roy, A. (2014) Description of six new species of Marginellidae from the deeper Eastern seaboard of South Africa. Malacologia Mostra Mondiale, 85, 20–27.
Sacco, F. (1890) Il Molluschi dei Terreni Terziarii del Piemonte e della Liguria Descritti da Luigi Bellardii. Part 6 (Volutidae, Marginellidae, Columbellidae). Memoide della Reale Accademia delle Science di Torino (Serie 2), 40, 295–368.
Silva, C.M., Landau, B. & La Perna, R. (2011) Biogeography of Iberian Atlantic Neogene marginelliform gastropods (Marginellidae, Cystiscidae): global change and transatlantic colonization. Journal of Paleontology, 85, 1052–1066.
https://doi.org/10.1666/11-104.1
Simone, L.R.L. (1996) Anatomy and systematics of Buccinanops gradatus (Deshayes, 1844) and Buccinanops moniliferus (Kiener, 1834) (Neogastropoda, Muricoidea) from the southeastern coast of Brazil. Malacologia, 38, 87–102.
Simone, L.R.L. (2001) Phylogenetic analyses of Cerithioidea (Mollusca, Caenogastropoda) based on comparative morphology. Arquivos de Zoologia, 36 (2), 147–263.
https://doi.org/10.11606/issn.2176-7793.v36i2p147-263
Simone, L.R.L. (2002) Comparative morphological study and phylogeny of representatives of the Superfamily Calyptraeoidea (including Hipponicoidea) (Mollusca, Caenogastropoda). Biota Neotropica, 2 (2), 1–137.
https://doi.org/10.1590/S1676-06032002000200013
Simone, L.R.L. (2004a) Comparative morphology and phylogeny of representatives of the superfamilies of architaenioglossans and the Annulariidae (Mollusca, Caenogastropoda). Arquivos do Museu Nacional, 62 (4), 387–504.
Simone, L.R.L. (2004b) Morphology and phylogeny of the Cypraeoidea (Mollusca, Caenogastropoda). Papel Virtual, Rio de Janeiro, 185 pp.
Simone, L.R.L. (2005) Comparative morphological study of representatives of the three families of Stromboidea and the Xenophoroidea (Mollusca, Caenogastropoda), with an assessment of their phylogeny. Arquivos de Zoologia, 37 (2), 141–267.
https://doi.org/10.11606/issn.2176-7793.v37i2p141-267
Simone, L.R.L. (2011) Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia, 42 (4), 161–323.
https://doi.org/10.11606/issn.2176-7793.v42i4p161-323
Simone, L.R.L. (2018) Main processes of body modification in gastropods: the lateralization. Malacopedia 1, 36–43.
http://www.moluscos.org/trabalhos/Malacopedia/01-05%20Simone%202018%20Malacopedia-Lateralization.pdf
Simone, L.R.L. & Mezzalira, S. (1994) Fossil molluscs of Brazil. Botelim do Instituto Geológico, 10, 1–202.
Sohl, N.F. (1963) New gastropod genera from the late Upper Cretaceous of the east Gulf Coastal Plain. Journal of Paleontology, 37 (4), 747–757.
Souza Jr, P.J.S. (1992) A new species of marginellid (Gastropoda: Prosobranchia) from the north coast of Brazil. Journal of Conchogy, 34, 237–242.
Souza Jr, P.J.S. (1997) A Tribo Marginellini Fleming, 1828 na Costa do Brasil (Mollusca, Gartropoda, Marginellidae). M.Sc. Dissertation, Museu Nacional, UFRJ, Rio de Janeiro, xiii, 79 pp.
Souza Jr, P.J.S. (2007) Análise cladística da família Marginellidae (Mollusca, Gastropoda) baseada em morfologia comparada. Ph.D. Dissertation, Instituto de Biociências da Universidade de São Paulo. São Paulo. Available from: http://www.teses.usp.br/teses/disponiveis/41/41133/tde-04102007-141745/fr.php (accessed 20 July 2014)
Souza Jr, P.J.S. & Coovert, G.A. (2001) Revision of the Recent Bullata Jousseaume, 1875 (Gastropoda: Marginellidae) with the description of two new species. Nautilus, 115 (1), 1–14.
Souza Jr, P.J.S. & Simone, L.R.L. (2019) Checklist of generic and supra-generic classification of Marginellidae (Caenogastropoda). Malacopedia, 2, 16–19.
https://doi.org/10.5962/bhl.part.11263
Swainson, W. (1840) A Treatise on Malacology, or Shells and Shellfish. Longman, Orme, Brown, Green, and Longmans and John Taylor. London, vii, 419 pp., 130 figs.
Swofford, D.L. (2002) PAUP: Phylogenetic Analysis Using Parsimony (and other Methods). Version 4.0 Beta. CD-ROM. Florida State University, Tallahassee. ???page numbers???
Swofford, D.L. & Maddison, W.P. (1987) Reconstructing ancestral character states under Wagner parsimony. Mathematical Biosciences, 87, 199–229.
https://doi.org/10.1016/0025-5564(87)90074-5
Thiele, J. (1929–1931) Handbuch der Systematischen Weichtierkunde. Gustav Fischer. Jena, 778 pp.
Tomlin, J.R. (1917) A Systematic list of the Marginellidae. Proceedings of the Malacogical Society of London, 12 (5), 242–306.
Tomlin, J.R. (1918) A Systematic list of the fossil Marginellidae. Proceedings of the Malacogical Society of London, 13 (3), 41–65.
Tracey, S., Todd, J.A. & Erwin, D.H. (1993) Chapter 8: Mollusca: Gastropoda. In: Benton, M. J. (Ed.), The Fossil Record. Chapman & Hall. London, pp. 131–167.
Veldsman, S.G. & Veldsman, V. (2015) Description of two new Marginella species (Marginellidae) from the East Coast Molluscan Province, South Africa: Marginella olivarum and Marginella sulizeae. Visaya, 4 (4), 55–63.
Vermeij, G.J. & Signor, P.W. (1992) The geographic, taxonomic and temporal distribution of determinate growth in marine gastropods. Biological Journal of the Linnean Society, 47, 233–247.
https://doi.org/10.1111/j.1095-8312.1992.tb00668.x
Vokes, E.H. (1971) The geologic history of the Muricinae and the Ocenebrinae. The Echo, 4, 37–54.
Wagner, P.J. (2001) Gastropod phylogenetics: progress, problems and implications. Journal of Paleontology, 75, 1128–1140.
https://doi.org/10.1017/S0022336000017182
Wakefield, A. (2003) A study of a population of Mesoginella pygmaea (Sowerby, 1846) (Gastropoda: Marginellidae) from the Bay of Islands, New Zealand. Novapex, 4, 93–100.
Wakefield, A. (2010) A revision of the genus Cryptospira Hinds, 1844 (Caenogastropoda Marginellidae). Novapex, 11, 1–55.
Wakefield, A. (2012) A review of the Marginella bicatenata Sowerby, 1914 complex (Gastropoda: Marginellidae) with the description of a new southeast African Marginella species. Novapex, 13 (2), 69–74.
Wakefield, A. & McCleery, T. (2005) Two new Volvarina (Marginellidae) from offshore subtidal habitats in northern Panama. Novapex, 6 (3), 45-52.
Wenz, W. (1938–1943) Gastropoda, Allgemeiner Teil und Prosobranchia. In: Schiendewolf , O.H. (Ed.), 1938–1944, Handbuch der Palozoologie, 6, (1–3) (1938), 1–480; (4–5) (1939), 481–720; (6) (1940), 721–960; (7) (1941), 961–1200; (8) (1943), 1201–1506; (9) (1944), 1507–1639, i–xii.
Winner, B.E. (1983) The direct development of Marginella (Prunum) apicinum. Of Sea and Shore, 13 (1), 37–38.
Woodring, W.P. (1970) Geology and paleontology of Canal Zone and adjoining parts of Panama: Description of Tertiary Mollusks (Gastropods: Eulimidae, Marginellidae to Helminthoglyptidae). Professional Papers of the United States Geological Survey, 306D, 299–452.
https://doi.org/10.3133/pp306D