Abstract
Ilyocypris species are abundant and widespread in North East Asian rice fields, but for many species their taxonomy is poorly defined, which negatively impacts ecological studies on rice field ecosystems. Herein, two of these species, Ilyocypris dentifera Sars, 1903 and Ilyocypris japonica Okubo, 1990, are redescribed, and a third species, Ilyocypris incus sp. nov., is described and named. This latter species has previously been reported from Japan and possibly Korea under the names Ilyocypris angulata Sars, 1903 and I. dentifera, but study of the hemipenes demonstrates that it is a separate species. Ilyocypris nipponica Okubo, 2004 nomen nudum previously reported from Japan is herein considered to be a junior synonym of I. dentifera. Tuberculated and non-tuberculated forms of both I. dentifera and I. japonica were recovered during this study and the presence or absence of tubercles is postulated to be related to environmental conditions, such as water chemistry or predation pressures. Although tuberculated forms of I. dentifera do occur, we concur with some previous studies that Ilyocypris angulata Sars, 1903 should be treated as a separate species, based on features of the hemipenes and carapaces. Figures of the carapaces and hemipenes of I. angulata, and Ilyocypris mongolica Martens, 1991, which is morphologically very close to I. japonica, are also presented for comparisons. This study confirms that three species of Ilyocypris inhabit North East Asian rice fields: I. dentifera, I. japonica and Ilyocypris incus sp. nov. Extant Ilyocypris species in North East Asia, including those from habitats other than rice fields, can be divided into two groups based on the Zenker organ, hemipenis, and sixth limb morphologies. These groups are postulated to represent two distinct lineages of Ilyocypris in North East Asia.
References
Anadón, P., De Deckker, P. & Juliá, R. (1986) The Pleistocene lake deposits of the NE Baza Basin (Spain): salinity variations and ostracod succession. Hydrobiologia, 143, 199–208.
https://doi.org/10.1007/BF00026662
Baird, W. (1845) Arrangement of the British Entomostraca, with a list of species, particularly noticing those which have as yet been discovered within the bounds of the club. Transactions of the Berwickshire Naturalists’ Club, 2, 145–158.
Brady, G.S. & Norman, A.M. (1889) A monograph of the marine and freshwater Ostracoda of the North Atlantic and of northwestern Europe. Section I: Podocopa. Scientific Transactions of the Royal Dublin Society, Series 2, 4, 63–270.
Broodbakker, N.W. (1988) Ecology and distribution of groundwater Crustacea and freshwater Ostracoda in SW Honshu and the Nansei Islands, Japan. 1 Groundwater Crustacea and list of stations. Verslagen en Technische Gegevens, Instituut voor Taxonomische Zoölogie, 53, 1–21.
Broodbakker N.W. & Danielopol D.L. (1982) The chaetotaxy of Cypridacea (Crustacea, Ostracoda) limbs: proposals for a descriptive model. Bijdragen tot de Dierkunde, 52, 103–120.
https://doi.org/10.1163/26660644-05202003
Chandran, N.N., Fojtova, D., Blahova, L., Rozmankova, E. & Blaha, L. (2018) Acute and (sub)chronic toxicity of the neonicotinoid imidacloprid on Chironomus riparius. Chemosphere, 209, 568–577.
https://doi.org/10.1016/j.chemosphere.2018.06.102
Chang, C.Y., Lee, J. & Smith, R.J. (2012) Nonmarine ostracods (Crustacea) from South Korea, including a description of a new species of Tanycypris Triebel (Cyprididae, Cypricercinae). Zootaxa, 3161 (1), 1–19.
http://dx.doi.org/10.11646/zootaxa.3161.1.1
Chen, S.Z. (1982) Subclass Ostracoda. In: Dong, Y.M., Dai, A.Y., Jiang, X.Z., Chen, S.Z., Chen, Y.S. & Cai, R.X. (Eds.), Atlas of the Animals in China: Crustaceans. Volume 1, Version 2. Science Press, Beijing, pp. 48–63. [In Chinese]
Chen, S.Z. (1990) Description of male Ilyocypris angulata (Crustacea, Ostracoda). Acta Micropalaeontologica Sinica, 7, 253–256.
Churchel, M.A., Hanula, J.L., Berisford, C.W., Vose, J.M. & Dalusky, M.J. (2011) Impact of imidacloprid for control of hemlock woolly adelgid on nearby aquatic macroinvertebrate assemblages. Southern Journal of Applied Forestry, 35, 26–32.
D’ Ambrosio, D.S., Díaz, A.R., García, A. & Claps, M.C. (2015) First description of the soft part anatomy of Ilyocypris ramirezi Cusminsky & Whatley (Crustacea, Ostracoda) from Argentina, South America. Zootaxa, 3957 (1), 59–68.
https://doi.org/10.11646/zootaxa.3957.1.4
De Deckker, P. (1981) Taxonomy and ecological notes of some ostracods from Australian inland waters. Transactions of the Royal Society of South Australia, 105, 91–138.
Delorme, D.L. (1970) Freshwater ostracodes of Canada. Part IV. Families Ilyocyprididae, Notodromadidae, Darwinulidae, Cytherideidae, and Entocytheridae. Canadian Journal of Zoology, 48, 1251–1259.
https://doi.org/10.1139/z70-214
Dodson, S.I. (1974) Adaptive change in plankton morphology in response to size-selective predation: A new hypothesis of cyclomorphosis. Limnology and Oceanography, 19, 721–729.
https://doi.org/10.4319/lo.1974.19.5.0721
Forró, L., Meisch, C., Petersen, H. & Martens, K. (1987) Ostracoda taxa described by E. Daday, together with a catalogue of pertinent material in the Hungarian Natural History Museum. Miscellanea Zoologica Hungarica, 4, 33–63.
Gauthier, H. (1928) Recherches sur la faune des eaux continentales de l’Algérie et de la Tunisie. Thèses présentées à la Faculté des sciences de l’Université de Paris, Séries A, 1160, 1–423.
Gauthier, H. (1929) Cladocères et ostracodes du Sahara Central. Bulletin de la Société d’Histoire Naturelle de l’Afrique du Nord, 20, 143–162.
Ghaouaci, S., Yavuzatmaca, M., Külköylüoglu, O. & Amarouayache, M. (2017) An annotated checklist of the non-marine ostracods (Crustacea) of Algeria with some ecological notes. Zootaxa, 4290 (1), 140–154.
https://doi.org/10.11646/zootaxa.4290.1.8
Hayashi, N., Ohuchi, T. & Miyata, N. (2009) Planktonic biota occurring in rice fields in Japan. Journal of Natural History Museum and Institute, Chiba, 10, 71–79. [In Japanese, with English abstract]
Hayashi, N., Ohuchi, T. & Miyata, N. (2012) Rotifera and crustacean fauna of Japanese rice-paddies. Lake Inba-numa Watershed Research and Management, 1, 57–62. [In Japanese]
Herzog, Q., Rabus, M., Ribeiro, B.W. & Laforsch, C. (2016) Inducible defenses with a “twist”: Daphnia barbata abandons bilateral symmetry in response to an ancient predator. PLoS ONE 11 (2), e0148556.
https://doi.org/10.1371/journal.pone.0148556
Huang, B. (1982) Pleistocene Ostracoda from Lijiang Basin, Yunnan. Acta Palaeontologica Sinica, 21, 174–180. [In Chinese with English summary]
Huang B.R. (1979) Quaternary Ostracoda from Qinghai Lake region. Bulletin of Nanjing Institute of Geology and Palaeontology, Academia Sinica, 7, 187–236, with 11 plates.
Huang, B.R., Yang, H.R. & You, K.Y. (1982) Pliocene and Quaternary Ostracoda of southern and southwestern Tibet. In: Investigation Group of Qinghai-Tibet Plateau of Chinese Academy of Sciences (Eds), Paleontology of Tibet, Vol. 4. Science Press, Beijing, pp. 326–348. [in Chinese]
Huang, M.X. (1986) A general survey of ecological distribution of Ostracoda (Crustacea) in Sichuan Province. Sichuan Journal of Zoology, 5, 8–10. [In Chinese]
Huang, M.X., Du, J. & Long, Z.H. (2006) The crustacean of Sichuan. Sichuan Science and Technology Publishers, Chengdu, pp. 128–149. [In Chinese]
Kaiser, K., Kempf, E.K., Leroi-Gourhan, A. & Schütt, H. (1973) Quartärstratigraphische Untersuchungen aus dem Damaskus-Becken und seiner Umgebung. Zeitschrift für Geomorphologie, N.F., 17, 263–353.
Kaji, T. & Tsukagoshi, A. (2010) Heterochrony and modularity in the degeneration of maxillopodan nauplius eyes. Biological Journal of the Linnean Society, 99, 521–529.
https://doi.org/10.1111/j.1095-8312.2009.01382.x
Kamiya, T. (1988a) Contrasting population ecology of Loxoconcha (Ostracoda, Crustacea) in Recent Zostera (eelgrass) beds: adaptive differences between phytal and bottom-dwelling species. Micropaleontology, 34, 316–331.
https://doi.org/10.2307/1485600
Kamiya, T. (1988b) Different sex-ratios in two Recent species of Loxoconcha (Ostracoda). Senckenbergiana lethaea, 68, 337–345.
Karanovic, I. (2012) Recent freshwater ostracods of the world. Springer-Verlag, Berlin, Heidelberg, 608 pp.
https://doi.org/10.1007/978-3-642-21810-1
Karanovic, I. & Lee, W. (2013) On the ostracod genus Ilyocypris, with description of one new species from Korea and the first report of males of I. bradyi (Crustacea: Ostracoda: Podocopida). Proceedings of the Biological Society of Washington, 126, 39–71.
https://doi.org/10.2988/0006-324X-126.1.39
Karuthapandi, M. & Rao, D.V. (2017) Crustacea: Ostracoda (seed shrimps). In: Chandra, K., Gopi, K.C., Rao, D.V., Valarmathi, K. & Alfred, J.R.B. (Eds.), Current Status of Freshwater Faunal Diversity in India, Published by the Director, Zoological Survey of India, Kolkata, pp. 253–271.
Karuthapandi, M., Rao, D.V. & Innocent, B.X. (2014) Freshwater Ostracoda (Crustacea) of India - a checklist. Journal of Threatened Taxa, 6, 6576–6581.
https://doi.org/10.11609/JoTT.o3682.6576-81
Kaufmann, A. (1900a) Cypriden und Darwinuliden der Schweiz. Revue Suisse de Zoologie, 8, 209–423.
https://doi.org/10.5962/bhl.part.10584
Kaufmann, A. (1900b) Neue Ostracoden aus der Schweiz. Zoologischer Anzeiger, 23, 131–133.
Kempf, E.K. (1967) Ilyocypris schwarzbachi n. sp. (Crustacea, Ostracoda) und ein vorläufiges Ostrakoden-Diagramm aus dem pleistozänen Löß von Kärlich (Neuwieder Becken). Sonderveröffentlichungen des Geologischen Instituts der Universität Köln, 13, 65–79.
Keyser, D. (2005) Histological peculiarities of the noding process in Cyprideis torosa (Jones) (Crustacea, Ostracoda). Hydrobiologia, 538, 95–106.
https://doi.org/10.1007/s10750-004-4940-x
Keyser, D. & Aladin, N. (2004) Noding in Cyprideis torosa and its causes. Studia Quaternaria, 21, 19–24.
Kim, W. & Min, G.S. (1991a) Redescription of four Recent freshwater ostracod (Crustacea: Ostracoda) species from Korea. Korean Journal of Zoology, 34, 307–322.
Kim, W. & Min, G.S. (1991b) Redescription of Recent freshwater ostracods (Crustacea: Ostracoda) from Korea: A systematic study of four species of family Cyprididae. The Korean Journal of Systematic Zoology, 7, 81–100.
Klassen, R.W., Delorme, L.D. & Mott, R.J. (1967) Geology and paleontology of Pleistocene deposits in southwestern Manitoba. Canadian Journal of Earth Sciences, 4, 433–447.
https://doi.org/10.1139/e67-022
Klie, W. (1926) Zweiter Beitrag zur Kenntnis der Süsswasser-Ostracoden Russlands. Raboty Volzhskoy Biologicheskoy Stantsii, 9, 35–53.
Kulkarni, M.R., Padhye, S., Vanjare, A.I., Jakhalekar, S.S., Shinde, Y.S., Paripatyadar, S.V., Sheth, S.D., Kulkarni, S., Phuge, S.K., Bhakare, K., Klukarni, A.S., Pai, K. & Ghate, H.V. (2015) Documenting the fauna of a small temporary pond from Pune, Maharashtra, India. Journal of Threatened Taxa, 7, 7196–7210.
https://doi.org/10.11609/JoTT.o4190.7196-210
Lee, E., Huh, M. & Schornikov, E.I. (2000) Ostracod fauna from the East Sea coast of Korea and their distribution - preliminary study on Ostracoda as an indicator of water pollution. Journal of the Geological Society of Korea, 36, 435–472. [In Korean with English abstract]
Ma, S. & Yu, N. (2018) Freshwater ostracods (Crustacea) from Tiantong National Forest Park and Dongqian Lake, eastern China, with descriptions of two new species. Journal of Natural History, 52, 1825–1868.
https://doi.org/10.1080/00222933.2018.1489085
Martens, K. (1985) Effects of temperature and salinity on postembryonic growth in Mytilocypris henricae (Chapman) (Crustacea, Ostracoda). Journal of Crustacean Biology, 5, 258–272.
https://doi.org/10.2307/1547873
Martens, K. (1987) Homology and functional morphology of the sexual dimorphism in the antenna of Sclerocypris Sars, 1924 (Crustacea, Ostracoda, Megalocypridinae). Bijdragen tot de Dierkunde, 57, 183–190.
https://doi.org/10.1163/26660644-05702003
Martens, K. (1991) On a small collection of non-marine ostracods from Mongolia, with the description of a new species (Crustacea, Ostracoda). Miscellanea Zoologica Hungarica, 6, 53–60.
Martens, K., Ortal, R. & Meisch, C. (1992) The ostracod fauna of Mamilla Pool (Jerusalem, Israel) (Crustacea, Ostracoda). Zoology in the Middle East, 7, 95–114.
https://doi.org/10.1080/09397140.1992.10637628
Martens, K. & Savatenalinton. S. (2011) A subjective checklist of the Recent, free-living, non-marine Ostracoda (Crustacea). Zootaxa, 2855 (1), 1–79.
https://doi.org/10.11646/zootaxa.2855.1.1
Martens, K., Savatenalinton, S., Schön, I., Meisch, C. & Horne, D.J. (2013) World checklist of freshwater Ostracoda species. Freshwater Animal Diversity Assessment, Belgian Biodiversity Platform. Available from: http://fada.biodiversity.be/group/show/18 (accessed 7 December 2016)
Masi, L. (1905) Nota sugli Ostracodi viventi nei dintorni di Roma ed osservazioni sulla classificazione della Cypridae. Bollettino della Società Zoological Italiana, Seria 2, 6, 115–128.
Mazzini, I., Gliozzi, E., Rossetti, G. & Pieri, V. (2014) The Ilyocypris puzzle: A multidisciplinary approach to the study of phenotypic variability. International Review of Hydrobiology, 99, 1–14.
https://doi.org/10.1002/iroh.201301729
Meisch C. (2000) Freshwater Ostracoda of Western and Central Europe. Süßwasserfauna von Mitteleuropa 8/3. Spektrum Akademischer Verlag, Heidelberg, Berlin, 522 pp.
Meisch, C., Smith, R.J. & Martens, K. (2019) A subjective global checklist of the extant non-marine Ostracoda (Crustacea). European Journal of Taxonomy, 492, 1–135.
https://doi.org/10.5852/ejt.2019.492
Mischke, S., Herzschuh, U., Kürschner, H., Chen, F., Meng, F. & Sun, Z. (2003) Sub-Recent Ostracoda from Qilian Mountains, NW China. Journal of Micropalaeontology, 22, 137–138.
https://doi.org/10.1144/jm.22.2.137
Mischke, S., Herzschuh, U., Sun, Z., Qiao, Z., Sun, N. & Zander, A.M. (2006) Middle Pleistocene Ostracoda from a large freshwater lake in the presently dry Qaidam Basin (NW China). Journal of Micropalaeontology, 25, 57–64.
https://doi.org/10.1144/jm.25.1.57
Müller, G.W. (1912) Crustacea. Ostracoda. In: Schulze, F.E. (Ed.), Das Tierreich 31, Reprinted 1966 by J. Cramer, Weinheim, pp. 1–434.
Müller, O.F. (1776) Zoologiae Danicae Prodromus, seu Animalium Daniae et Norvegiae Indigenarum characters, nomina, et synonyma imprimis popularium. Havniae. XXXII, 1–282.
https://doi.org/10.5962/bhl.title.63795
Nakao Y. & Tsukagoshi, A. (2002) Brackish-water Ostracoda (Crustacea) from Obitsu River estuary, Central Japan. Species Diversity, 7, 67–115.
https://doi.org/10.12782/specdiv.7.67
Nakao, Y. & Tsukagoshi, A. (2008) Seasonal and decadal changes in distribution and abundance of Ostracoda (Crustacea) in the brackish-water environments. Proceedings of the Institute of Natural Sciences, Nihon University, 43, 249–275.
Okubo, I. (1974) Freshwater Ostracoda from Japan, XI. Ilyocypris angulata Sars, 1902. The Research Bulletin of Okayama Shujitsu Junior College, 4, 46–50.
Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39–50.
Okubo I. (2004) Nihon tansui san kaimijinko rui ni tsuite. Kabushikigaisha Sanmon Insatsusho, Okayama, 72 pp. [In Japanese, privately published]
Okubo, I. & Ida, K. (1989) Freshwater Ostracoda from Gunma Prefecture, Japan. Bulletin of the Biogeographical Society of Japan, 44, 105–107. [In Japanese]
Okubo, I. & Terauchi, M. (1992) Freshwater Ostracoda from the Southwestern Islands of Japan. Proceedings of the Japanese Society of Systematic Zoology, 46, 101–108.
Osterberg, J.S., Darnell, K.M., Blickley, T.M., Romano, J.A. & Rittschof, D. (2012) Acute toxicity and sub-lethal effects of common pesticides in post-larval and juvenile blue crabs, Callinectes sapidus. Journal of Experimental Marine Biology and Ecology, 424–425, 5–14.
https://doi.org/10.1016/j.jembe.2012.05.004
Peel, M.C., Finlayson, B.L. & McMahon, T.A. (2007) Updated world map of the Köppen-Geiger climate classification. Hydrology and Earth System Sciences Discussions, European Geosciences Union, 11, 1633–1644.
https://doi.org/10.5194/hess-11-1633-2007
Ramdohr, F.A. (1808) Über die Gattung Cypris Müll. und drei zu derselben gehörige neue Arten. Magazin der Gesellschaft Naturforschender Freunde zu Berlin für die Neuesten Entdeckungen in der Gesammten Naturkunde, 2, 85–93.
Sánchez-Bayo, F. & Goka, K. (2006a) Ecological effects of the insecticide imidacloprid and a pollutant from antidandruff shampoo in experimental rice fields. Environmental Toxicology and Chemistry, 25, 1677–1687.
https://doi.org/10.1897/05-404R.1
Sánchez-Bayo, F. & Goka, K. (2006b) Influence of light in acute bioassays of imidacloprid and zinc pyrithione to zooplankton crustaceans. Aquatic Toxicology, 78, 262–271.
https://doi.org/10.1016/j.aquatox.2006.03.009
Sari, N., Külköylüoðlu, O. & Akdemir, D. (2012) First record and detailed description of the male of Ilyocypris inermis Kaufmann, 1900 (Crustacea, Ostracoda). Turkish Journal of Zoology, 36, 484–495.
Sars, G.O. (1866) Oversigt af Norges marine ostracoder. Forhandlinger I Videnskabs-Selskabet I Christiania, 1865, 1–130.
Sars, G.O. (1890) Oversigt af Norges Crustaceer med forelobige Bemaerkninger over de nye eller mindre bekjendte arter: 2 (Branchiopoda, Ostracoda, Cirripedia). Forhandlinger I Christiania Videnskabs-Selskabet, 1890, 1–80.
Sars, G.O. (1903) Fresh-water Entomostraca from China and Sumatra. Archiv for Mathematik og Naturvidenskab, 25, 3–44.
Shinde, Y.S., Victor, R. & Pai, K. (2014) Freshwater ostracods (Crustacea: Ostracoda) of the plateaus of the northern Western Ghats, India. Journal of Threatened Taxa, 6, 5667–5670.
https://doi.org/10.11609/JoTT.o3610.5667-70
Shuhaimi-Othman, M., Yakub, N., Ramle, N-A. & Abas, A. (2011) Toxicity of metals to a freshwater ostracod: Stenocypris major. Journal of Toxicology, 2011, 1–8.
https://doi.org/10.1155/2011/136104
Sidorov, D.A. & Semenchenko, K.A. (2012) New records of freshwater ostracods (Crustacea) from the Far East of Russia, with a checklist of recent freshwater ostracods of the region. Arthropoda Selecta, 21, 227–234.
https://doi.org/10.15298/arthsel.21.3.04
Smith, R.J., Janz, H. & Okubo, I. 2011. Recent Cyprididae and Ilyocyprididae (Crustacea: Ostracoda) from Lake Biwa, Japan, including a summary of the lake's ostracod fauna. Zootaxa, 2874, 1–37.
https://doi.org/10.11646/zootaxa.2874.1.1
Smith, R.J., Lee, J. & Chang, C.Y. (2014) Nonmarine Ostracoda (Crustacea) from Jeju Island, South Korea, including descriptions of two new species. Journal of Natural History, 49, 37–76.
https://doi.org/10.1080/00222933.2014.946110
Smith, R.J., Matzke-Karasz, R. & Kamiya, T. (2016a) Sperm length variations in five species of cypridoidean non-marine ostracods (Crustacea). Cell and Tissue Research, 366, 483–497.
https://doi.org/10.1007/s00441-016-2459-x
Smith, R.J., Matzke-Karasz, R., Kamiya, T. & De Deckker, P. (2016b) Sperm lengths of non-marine cypridoidean ostracods (Crustacea). Acta Zoologica, 97, 1–17.
https://doi.org/10.1111/azo.12099
Smith, R.J., Zhai, D., Savatenalinton, S., Kamiya, T. & Yu, N. (2018) A review of rice field ostracods (Crustacea) with a checklist of species. Journal of Limnology, 77, 1–16.
https://doi.org/10.4081/jlimnol.2017.1648
Stepanaitys, N.E. (1960) New species of ostracods from the Neogene-Quaternary deposits of SW Turkmenia. Trudy Instituta Geologii Akademii Nauk Turkmenskoi SSR, 2 (for 1959), 298–315. [In Russian]
Tabuki, R. & Tsuhako, Y. (2015) Freshwater ostracode fauna from the biotope of the University Museum (Fuujukan) of the University of the Ryukyus, middle part of Okinawa Island, Japan. Bulletin of Faculty of Education University of the Ryukyus, 87, 207–218. [In Japanese with English abstract]
Tanaka, H., Kotorii, H., Yokozawa, K., Wakabayashi, F., Komoto, K. & Sano, K. (2015) Distribution and occurrence of freshwater Ostracoda (Crustacea) from southwest region of Mt. Fuji. Taxa, Proceedings of the Japanese Society of Systematic Zoology, 38, 26–41. [In Japanese with English abstract]
Tinn, O. & Oakley, T.H. (2008) Erratic rates of molecular evolution and incongruence of fossil and molecular divergence time estimates in Ostracoda (Crustacea). Molecular Phylogenetics and Evolution, 48, 157–167.
https://doi.org/10.1016/j.ympev.2008.03.001
Van der Meeren, T., Almendinger, J.E., Ito, E. & Martens, K. (2010) The ecology of ostracodes (Ostracoda, Crustacea) in western Mongolia. Hydrobiologia, 641, 253–273.
https://doi.org/10.1007/s10750-010-0089-y
van Harten, D. (2000) Variable noding in Cyprideis torosa (Ostracoda, Crustacea): an overview, experimental results and a model from Catastrophe Theory. Hydrobiologia, 419, 131–139.
https://doi.org/10.1023/A:1003935419364
Victor, R. (2004) Crustacea: Ostracoda. In: Yule, C.M. & Sen, Y.H. (Eds.), Freshwater Invertebrates of the Malaysian Region. Akademi Sains Malaysia, Kuala Lumpur, pp. 225–253.
Victor, R. & Fernando, C.H. (1980) Freshwater Ostracoda from the ricefields of South East Asia. Tropical Ecology & Development, 1980, 957–970.
Victor, R. & Fernando, C.H. (1981) Ilyocypris dentifera Sars, 1903 versus Ilyocypris angulata Sars, 1903. A taxonomic study of two species of Ilyocypris Brady and Norman, 1889 (Crustacea, Ostracoda) described from China. Canadian Journal of Zoology, 59, 1103–1110.
https://doi.org/10.1139/z81-153
Wrozyna, C., Frenzel, P., Steeb, P., Zhu, L. & Schwalb, A. (2009) Recent lacustrine Ostracoda and a first transfer function for palaeo-water depth estimation in Nam Co, southern Tibetan Plateau. Revista Española de Micropaleontologia, 41, 1–20.
Yamada, S. (2007) Structure and evolution of podocopan ostracod hinges. Biological Journal of the Linnean Society, 92, 41–62.
https://doi.org/10.1111/j.1095-8312.2007.00870.x
Yamaguchi, S. & Endo, K. (2003) Molecular phylogeny of Ostracoda (Crustacea) inferred from 18S ribosomal DNA sequences: implication for its origin and diversification. Marine Biology, 143, 23–38.
https://doi.org/10.1007/s00227-003-1062-3
Yang, F., Sun, Z., Qiao, Z. & Zhang, Y. (2004) Revision of the diagnosis of the genus Qinghaicypris Huang, 1979 (Ostracoda) and the environmental significance of its type species. Acta Micropalaeontologica Sinica, 21, 367–381. [Chinese with English Summary]
Yang, F., Sun, Z., Zhang, Y. & Qiao, Z. (2002) Taxonomic significance of nodal ornamentation of Quaternary genus “Ilyocypris” (Ostracoda) from Qaidam Basin, Qinghai Province. Acta Micropalaeontologica Sinica, 19, 15–32. [In Chinese with an English summary]
Yin, Y. & Martens, K. (1997) On a new species of Fabaeformiscandona Krstic, 1972 (Crustacea, Ostracoda) from China, with a preliminary checklist of Recent Chinese non-marine ostracods. Hydrobiologia, 357, 117–128.
https://doi.org/10.1023/A:1003182720121
Yu, N. (2014) Non-marine Ostracoda from China. Shanghai Educational Publishing House, Shanghai, 283 pp.
Yu, N., Zhao, Q., Li, E., Chen, S. & Chen, L. (2009) An updated and annotated checklist of recent nonmarine ostracods from China. Zootaxa, 2067 (1), 29–50.
https://doi.org/10.11646/zootaxa.2067.1.2
Yu, N., Zhao, Q., Schornikov, E.I. & Chen, L. (2005) Recent ostracods from the Taihu Lake. Acta Micropalaeontologica Sinica, 22, 143–151.
Zenina, M.A. & Schornikov, E.I. (2008) Ostracod assemblages of the freshened part of Amursky Bay and lower reaches of the Razdolnaya River (Sea of Japan). In: Lutaenko, K.A. & Vaschenko, M.A. (Eds.), Ecological studies and the state of the ecosystem of Amursky Bay and the estuarine zone of the Razdolnaya River (Sea of Japan). APN, Vladivostok, Dalnauka, pp. 156–185.
Zhai, D. & Xiao, J. (2013) On a new species of the genus Ilyocypris Brady & Norman 1889 from Hulun Lake, China. Il Naturalista Siciliano, 37, 465–469.
Zhai, D. & Zhao, W. (2014) On some Recent non-marine ostracods from northern China, with descriptions of one new species. Crustaceana, 87, 985–1026.