Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2019-08-20
Page range: 437–473
Abstract views: 174
PDF downloaded: 5

A review of the genus Ancylosis Zeller, 1839 (Lepidoptera: Pyralidae: Phycitinae) from Ukraine

Institute for Evolutionary Ecology of the National Academy of Sciences of Ukraine, 37 Academician Lebediev str., 03143, Kyiv, Ukraine.
T.I. Vyazemsky Karadag Scientific Station—Nature Reserve of RAS, Kurortnoye, Pheodosia, Crimea Republic, 298188.
Institute for Evolutionary Ecology of the National Academy of Sciences of Ukraine, 37 Academician Lebediev str., 03143, Kyiv, Ukraine.
Ancylosoma new species new synonymy new host plants Palaearctic region Phycitinae Lepidoptera

Abstract

Nine species of Ancylosis are recorded from Ukraine. Ancylosis larissae sp. nov. related to A. hellenica (Staudinger, 1870) and A. rhodochrella (Herrich-Schäffer, 1852), is described from Crimea. Ancylosis albicosta (Staudinger, 1870) is reported from Ukraine for the first time. Ancylosis albidella Ragonot, 1888, A. syrtella (Ragonot, 1887), A. rhodochrella, A. harmoniella (Ragonot, 1887) and A. monella Roesler, 1973 are removed from the list of Pyralidae of Ukraine due to misidentifications. The host plants for A. sareptalla (Herrich-Schäffer, 1861) and A. substratellum (Christoph, 1877) comb. nov. are recorded for the first time. New host plant is recorded for A. roscidella (Eversmann, 1844). The diagnoses for all species known from Ukraine are provided. The adults and genitalia of both sexes are illustrated for all Ukrainian species and some closely related species from adjacent territories. A key to the species is given based on external characters and the genitalia of both sexes. The female genitalia of A. deserticola (Staudinger, 1870), A. hellenica (Staudinger, 1870), A. sabulosella (Staudinger, 1879) as well as the male genitalia of A. pallida (Staudinger, 1870) are described and illustrated as they have been dealt incorrectly in literature. The status of A. albidella and the records of this species from Europe are discussed. The new synonymy is proposed: Ancylosoma Roesler, 1973 syn. nov. of Ancylosis Zeller, 1839 and A. sareptalla gulbaharensis Roesler, 1973 syn. nov. of A. sareptalla (Herrich-Schäffer, 1861). Ancylosis decolorella (Ragonot & Hampson, 1901) spec. rev., stat. nov. is taken out from synonymy with A. sabulosella. The lectotype of Myelois deserticola Staudinger, 1870 is designated.

References

  1. Alberti, B. & Soffner, J. (1962) Zur Kenntnis der Lepidopteren-Fauna Süd- und Südostrußland. Mitteilungen der Münchner Entomologische Gesellschaft E.V., 52, 147–198.

    Amsel, H.G. (1935) Zur Kenntnis der Microlepidopterenfauna des südlichen Toten-Meer-Gebietes, nebst Beschreibungen neuer palästinensischer Macro- und Microlepidoptera. Veröffentlichungen aus dem deutschen Kolonial- und Übersee-Museum in Bremen, 1 (2), 203–221., pls. 11–12.

    Amsel, H.G. (1942) Revision der Gattung Holcopogon Stgr. (Lepidoptera: Scythrididae). Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen, 3, 224–237., pl. 3.

    Amsel, H.G. (1951) Die Microlepidopteren der Brandt’schen Iran-Ausbeute. 3. Teil. Arkiv för Zoologi (New Series), 1 (36), 525–563.

    Amsel, H.G. (1954) Die Microlepidopteren der Brandt’schen Iran-Ausbeute. 4. Teil. Arkiv för Zoologi (New Series), 6 (16), 255–326., pls. 1–33.

    Amsel, H.G. (1955) Über mediterrane Microlepidopteren und einige transcaspische Arten. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, 31 (83), 1–64.

    Amsel, H.G. (1959) Microlepidoptera aus Iran. Stuttgarter Beiträge zur Naturkunde, (28), 1–47.

    Amsel, H.G. (1961) Die Microlepidopteren der Brandt’schen Iran-Ausbeute. 5. Teil. Arkiv för Zoologi (New Series), 13 (17), 323–445., pls. 1–9.

    Asselbergs, J.E.F. (1999) Arenipses sabella Hampson, 1901, from South Spain: new to the European fauna; Thylacoptila paurosema Meyrick, 1885, new to the fauna of the Canary Islands; Ancylosis (Ancylosis) albidella Raginot, 1888, new to the Spanish fauna (Lepidoptera: Pyralidae, Galleriinae, Phycitinae). SHILAP Revista de Lepidopterologia, 27 (105), 125–129.

    Asselbergs, J.E.F. (2008) Order Lepidoptera, superfamily Pyraloidea. In: van Harten, A. (Ed.), Arthropod fauna of the UAE Vol. I. Dar Al Ummah Printing, Publishing, Distribution & Advertising, Abu Dhabi, pp. 469–561.

    Asselbergs, J.E.F. (2010) Order Lepidoptera, superfamily Pyraloidea. In: van Harten, A. (Ed.), Arthropod fauna of the UAE (Part 2). Dar Al Ummah Printing, Publishing, Distribution & Advertising, Abu Dhabi, pp. 554–563.

    Balinsky, B.I. (1987) Ancylosis (Heterographis) subpyrethrella (Ragonot, 1888) and five related new species from southern Africa (Lepidoptera: Phycitinae). Annals of the Transvaal Museum, 34 (14), 303–317.

    Balinsky, B.I. (1989) The Ancylosis /Heterographis/ Staudingeria group of Phycitinae (Lepidoptera: Pyralidae) in southern Africa. Annals of the Transvaal Museum, 35 (5), 75–107.

    Bidzilya, O.V. & Budashkin, Yu.I. (2004) New records of Lepidoptera from Ukraine. Proceedings of Zoological Museum of Kiev Taras Shevchenko National University, 2, 59–67. [in Ukrainian]

    Bidzilya, O.V. &. Budashkin, Yu.I. (2017) New records of Lepidoptera from Ukraine and description of a new species of Caloptilia Hübner, 1825 (Lepidoptera, Gracillariidae) from the mountains of Crimea. Nota lepidopterologica, 40 (2), 145–161.

    https://doi.org/10.3897/nl.40.13085

    Bidzilya, O.V., Budashkin, Yu. I. & Gaedike, R. (2016) A revision of Eudarcia glaseri-species group (Lepidoptera, Meessiidae) with description of two new species from Greece and Crimea. Zootaxa, 4179 (3), 547–560.

    https://doi.org/10.11646/zootaxa.4179.3.9

    Bidzilya, O.V., Budashkin, Yu.I. & Zhakov, A.V. (2003) New records of Lepidoptera (Insecta) from Ukraine. The Kharkov Entomological Society Gazette, 10 (1–2), 59–73. [in Russian]

    Bidzilya, O.V., Budashkin, Yu.I. & Zhakov, A.V. (2016) Checklist of grass-mining moths of Ukraine with description of one new species (Lepidoptera: Elachistidae). SHILAP Revista de lepidopteroloigia, 44 (173), 17–38.

    Bidzilya, O.V., Budashkin, Yu.I. & Zhakov, A.V. (2017) Check-list of scythridid moths (Lepidoptera, Scythrididae) of Ukraine with description of two new species. Zootaxa, 4291 (3), 481–503.

    https://doi.org/10.11646/zootaxa.4291.3.3

    Bidzilya, O.V., Budashkin, Yu.I., Zhakov, A.V., Kljuchko, Z.F. & Kostjuk, I.Yu. (2001) The fauna of Lepidoptera of the «Stone Graves» Nature Reserve and its taxonomic structure. In: Morozova A.L. & Gnyubkin, V.F. (Eds.), Karadag. History, biology, archeology. The scientific works dedicated to 85-th anniversary of Karadag scientific station. SONAT, Simpheropol, pp. 72–107. [in Russian]

    Bidzilya, O.V., Budashkin, Yu.I., Zhakov, A.V. & Kostjuk, I.Yu. (2011) New and interesting records of Microlepidoptera (Lepidoptera) from Ukraine. Eversmannia, 25–26., 64–74., pl. 5. [in Russian]

    Bisset, G.A. (1938) Genus Heterographis. In: Pierce, F.N. & Metcalfe, J.W. (Eds.), The genitalia of the british Pyrales with the Deltoids and Plumes. Warmington, Oundle, Northants, p. 59.

    Blanchard, A. & Knudson, E.C. (1985) New species of Phycitinae (Lepidoptera, Pyralidae) from Texas, with description of a new genus. Proceedings of the Entomological Society of Washington, 87 (1), 231–238.

    Budashkin, Yu.I. (1992) Vognivky (Lepidoptera, Pyraloidea) Karadaz’kogo zapovidnyka (Krym) [Pyraloidea of Karadagh Nature Reserve (Crimea)]. Problemy zagal’noi ta molelekuljarnoi biologii, 10, 23–33. [In Ukrainian]

    Budashkin, Yu.I. (2004) Itogi dvadtsatiletnego statsionarnogo izuchenija fauny cheshuekrylykh (Lepidoptera) Karadagskogo prirodnogo zapovednika [Results of twenty-years study of Lepidoptera of Karadagh nature reserve]. In: Morozova A.L. & Gnyubkin V.F. (Eds.), Karadag. History, geology, botany, zoology. The scientific works dedicated to 90-th anniversary of T.I. Vyasemsky Karadag scientific station and 25-th anniversary of Karadag Nature Reserve. Book 1. SONAT, Simpheropol, pp. 323–366. [in Russian]

    Budashkin, Yu.I. (2006) The materials on Lepidopterofauna of the Kazantip nature reserve. Proceedings of the Nikitsky Botanical Garden—National Scientific Center, 126, 263–291. [in Russian]

    Budashkin, Yu.I. (2014) The fourth addition to fauna and bionomy of the Crimean Lepidoptera. Optimization and Protection of Ecosystems, 10 (29), 12–20. [in Russian]

    Budashkin, Yu.I. & Bidzilya, O. (2018) Four new species of the genus Eudarcia Clemens, 1880 (Lepidoptera: Meessiidae) from Crimea. Zootaxa, 4446 (1), 111–124.

    https://doi.org/10.11646/zootaxa.4446.1.8

    Budashkin, Yu.I. & Savchuk, V.V. (2010) New faunal and bionomic materials of the Crimean Lepidoptera. Optimization and Protection of Ecosystems, 2 (21), 42–57. [in Russian]

    Budashkin, Yu.I., Savchuk, V.V. & Pusanov, D.V. (2009) New information about fauna and bionomics of lepidopterans (Lepidoptera) of Crimea. Optimization and Protection of Ecosystems, 19, 33–45. [in Russian]

    Caradja, A. (1910) Beitrag zur Kenntnis über die geographische Verbreitung der Pyraliden des europäischen Faunengebietes nebst Beschreibung einiger neuer Formen. Deutsche entomologische Zeitschrift Iris, 24 (6–7), 105–147.

    Caradja, A. (1916) Beitrag zur Kenntnis der geographischen Verbreitung der Pyraliden und Tortriciden des europäischen Faunengebietes, nebst Beschreibung neuer Formen. Deutsche entomologische Zeitschrift Iris, 30 (1), 1–88, errata p. 151–152.

    Caradja, A. & Meyrick, E. (1937) Materialien zu einer Mikrolepidopterenfauna des Yülingshanmassivs (Provinz Yünnan). Deutsche entomologische Zeitschrift Iris, 51 (4), 137–182.

    Chrétien, P. (1911) Description de nouvelles espèces de Phycides de Mauritanie (Lep.). Bulletin de la Société entomologique de France, 2, 33–36.

    Chrétien, M.P. (1917) Contrubution á la connaisance des Lépidopterès du Nord de l’Afrique. Notes biologiques et critiques. Annales de la Société Entomologique de France, 85, 369–503.

    Christoph, H.T. (1876–1877) Sammelergebnisse aus Nordpersien, Krasnowodsk in Turkmenien und dem Daghestan. Horae Societatis entomologicae Rossicae, 12, 181–299., pls. 5–8.

    Clemens, B. (1860) Contributions to American lepidopterology. Proceedings of the Academy of Natural Sciences of Philadelphia, 12, 4–15., 156–174., 203–221., 345–362., 522–547.

    Curtis, J. (1833) Characters of some undescribed genera and species, indicated in the “Guide to an Arrangement of British Insects”. Entomological Magazine, 1 (2), 186–199.

    De Prins, J. & De Prins, W. (2019) Afromoths, online database of Afrotropical moth species (Lepidoptera). World Wide Web electronic publication. Available from: http:// www.afromoths.net (accessed 6 Marсh 2019)

    Disqué, H. (1901) Verzeichnis der in der Pfalz vorkommenden Kleinschmetterlinge. Mitteilungen der Pollichia, 63, 29–42.

    Duponchel, P.-A.-J. (1836–1837 [imprint “1836”]) Nocturnes 7.—Histoire naturelle des Lépidoptères ou Papillons de France, Paris, 10, 1–384, errata et addenda, 267–286 pp.

    Dyar, H.G. (1914) Report on the Lepidoptera of the Smithsonian Biological Survey of the Panama Canal Zone. Proceedings of the United States National Museum, 47 (2050), 139–350.

    https://doi.org/10.5479/si.00963801.47-2050.139

    Eversmann, E.F. v. (1844) Fauna Lepidopterologica Volgo-Uralensis. Typis Universitatis, Casani, i–xiv, 633 pp.

    Falkovitsh, M.I. (1969a) Tcheshuekrylye (Lepidoptera) gor Kokshetau i Zharkol’-Shoindykol’skogo plato [Lepidoptera of Kokshetau Mountains and Zharkol’-Shoindykol’ plateau]. In: Biokompleksnye issledovaniya v Kazakhstane. Ch. 1. Rastitel’nye soobshchestva i zhivotnoe naselenie stepey i pustyn’ Tsentral’nogo Kazakhstana [Biocomplex researches in Kazakhstan. Part 1. Plant communities and animals of steppes and deserts of Central Kazakhstan]. Nauka, Leningrad, pp. 444–468. [in Russian]

    Falkovitsh, M.I. (1969b) O pishchevyh sviazyah pustynnyh tcheshuekrylyh (Lepidoptera) v Srednei Azii [On host-plant relationships of desert Lepidoptera in Middle Asia]. Doklady na 21 ezhegodnom chtenii pamyati N.A. Kholodkovskogo (12.04.1968) [Reports on annual meeting in memory of A.N. Kholodkovskiy]. Nauka, Leningrad, 53–88. [in Russian]

    Ford, L.T. (1957) A note on the Ova and Larva of Heterographis oblitella Zeller (Lep.: Phycitidae). Entomologist’s Gazette, 8, 27.

    Glaser, W. (1960). Zur Frage der ersten Stände von Heterographis oblitella Z. (Lep. Anerastiinae). Zeitschrift der Wiener Entomologischen Gesellschaft, 45, 157–158.

    Goater, B., Nuss, M. & Speidel, W. (2005) Pyraloidea I (Crambidae: Acentropinae, Evergestinae, Heliothelinae, Schoenobiinae, Scopariinae). In: Huemer, P. & Karsholt, O. (Eds.), Microlepidoptera of Europe. 4. Apollo Books, Stenstrup, pp. 1–304.

    Govorun, O.V. (2008) Rezul’taty issledovanija fauny ognjovok (Lepidoptera, Pyralidae) Chernomorskogo biosfernogo zapovednika [Results of study of Pyralidae of Chernomorskiy nature reserve]. Zapovidna sprava v Ukraini, 14 (1), 51–52. [in Russian]

    Hampson, G.F. (1901) The Lepidoptera Phalaenae of the Bahamas. Annals and Magazine of Natural History, including Zoology, Botany and Geology (Series 7), 7, 246–261.

    https://doi.org/10.1080/00222930108678466

    Hampson, G.F. (1917) Descriptions of new Pyralidae of the subfamilies Epipaschiinae, Chrysauginae, Endotrichinae, and Pyralinae. Annals and Magazine of Natural History, including Zoology, Botany and Geology (Series 8), 19, 65–100.

    https://doi.org/10.1080/00222931709486913

    Hampson, G.F. (1930) New genera and species of Phycitinae (Lepidoptera, Pyralidae). Annals and Magazine of Natural History, including Zoology, Botany and Geology (Series 10), 5, 50–80.

    https://doi.org/10.1080/00222933008673106

    Hayden, J.E. & Dickel, T.S. (2014) New North American records of Pyraloidea (Lepidoptera: Crambidae, Pyralidae) from southern Florida. Insecta Mundi, 361, 1–16.

    Heinemann, H.V. (1864) Drei neue Phycideen. Wiener Entomologische Monatschrift, 8, 288–296.

    Heinrich, C. (1956) American moths of the subfamily Phycitinae. Bulletin of the United States National Museum, 207, i–viii, 1–581, figs 1–1138.

    Herrich-Schäffer, G.A.W. (1847–1855 [imprint “1849”]) Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu Jakob Hübner’s Sammlung europäischer Schmetterlinge. 4: Die Zünsler und Wickler. G.J. Manz, Regensburg, pp. [1]–2–288, (Index) [1]–2–48, pls 1–23 (Pyralidides) + 1–59 (Tortricides).

    Herrich-Schäffer, G.A.W. (1843–1856 [“1856”]) Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu Jakob Hübner’s Sammlung europäischer Schmetterlinge. 6. G.J. Manz, Regensburg, pp. [i]–[iv] + [i]–ii–xviii + [i]+ii+viii + 1–178 + [1]–2–72 + [1]–2–48, pls. [i]–ii--xxii + i–xiv.

    Herrich-Schäffer, G.A.W. (1860) Neue Schmetterlinge aus Europa und den angrenzenden Laendern. G.J. Manz, Regensburg, 9–20 pp., pls. [10]–[18].

    Herrich-Schäffer, G.A.W. (1861) Neue Schmetterlinge aus Europa und den angrenzenden Laendern, pp. 24–32., pls. 19–26.

    Hormuzaki, K., (1907) Die Schmetterlinge (Lepidoptera) der Bukowina. III. Tiel: Familien Pyralidae bis Micropterigidae. Verhandlungen der Zoologish-Botanischen Gesellschaft in Wien, 57, 34–104.

    Hulst, G.D. (1888) New genera and species of Epipaschiae [sic] and Phycitidae. Entomologica Americana, 4, 113–118.

    Karolinskiy, Ye.A., Demyanenko, S.A., Guglya, Yu.A., Zhakov, A.V., Kavurka V.V. & Mushinskiy, V.G. (2018) On the fauna of Lepidoptera (Insecta) of the national nature park ‘Dvorichanskyi’ (Kharkiv region, Ukraine) and its environs. Contribution 2. The Kharkov Entomological Society Gazette, 26 (1), 55–114.

    Korb, S. (2017) Les types primaires de Pyraloidea de la collection Staudinger, conservée au Museum für Naturkunde de Berlin (Leibniz-Institut für Evolutions- und Biodiversitätsforschung). Alexanor, 27 (8), 516–526.

    Lebediev, A.G. (1937) A Contribution to the Knowledge of the Biocoenosis of Foliate Forests (part III). Recherches sur L`ecologie des Animaux Terrestres, 3, 25–71. [in Ukrainian]

    Lederer, J. (1855) Beitrag zur Schmetterlings-Fauna von Cypern, Beirut und einem Theile Klein-Asiens. Verhandlungen des zoologisch-botanischen Vereins in Wien 5 (Abhandlungen), 177–254., pls. 1–5.

    https://doi.org/10.5962/bhl.title.9422

    Lederer, J. (1863) Verzeichniss der von Herrn Johann und Frau Ludmilla Haberhauer 1861 und 1862 bei Varna in Bulgarien und Sliwno in Rumelien gesammelten Lepidopteren. Wiener Entomologische Monatschrift, 7, 17–27., 40–47., pl. 1.

    Lepiforum (2019) Ancylosis Zeller, 1839. Available from: http: //www.lepiforum.de/lepiwiki.pl (accessed 6 Marсh 2019)

    Leraut, P.J.A. (2008) Ebauche d’une liste des pyrales de France [Lepidoptera, Pyraloidea]. Revue française d’Entomologie (New Series), 29 (4), 149–166.

    Leraut, P.J.A. (2014) Moths of Europe, volume 4. Pyralids 2. Verrières-le-Buisson, N.A.P Editions, Paris, 441 pp.

    Lucas, D. (1948) Contribution a la faune des Lepidopteres de l`Afrique du Nord. Bulletin de la Société entomologique de France, 53, 88–90.

    Lucas, D. (1950) Contribution a l’étude des Lépidoptères Nord-Africains. Bulletin de la Société entomologique de France, 55 (1), 141–144.

    Lucas, D. (1955) Nouveaux Lepidopteres nord-africains. Bulletin de la Société des Sciences Naturelles du Maroc, 35, 251–258.

    Meyrick, E. (1936–1937) Exotic Microlepidoptera. Taylor and Francis, London, 160 pp.

    Nuss, M., Landry, B., Mally, Vegliante, R.F., Tränkner, A., Bauer, F., Hayden, L., Segerer, A., Schouten, R., Li, H., Trofimova, T., Solis, M.A., De Prins, J. & Speidel, W. (2003–2019) Global Information System on Pyraloidea. Available from: http://www.pyraloidea.org (accessed 6 March 2019)

    Oberthür, C. (1888) Faunes entomologiques. Descriptions d’insectes nouveaux ou peu connus. Études d’Entomologie, 12, 1–44., pls. 1–7.

    Pak, O.V. (2001) Vidovoj sostav ognjovok (Lepidoptera: Pyraloidea) jugo-vostoka Ukrainy po literaturnym svedenijam [Species composition of Pyraloidea of SE Ukraine based on literature sources]. Visnyk Donetskogo universitetu. Serija A: pryrodnichi nauky, Donetsk, 2, 281–286 pp. [in Russian]

    Pak, O.V. (2005) Three lepidoptera species new to the fauna of south-eastern Ukraine (Lepidoptera: Phycitidae, Pyraustidae, Lycaenidae). In: N.N. Yaroshenko (Ed.), Ecology and fauna of the south-east of Ukraine (miscellanea scientific papers). DonNU, Donetsk, 5, pp. 46–47. [in Russian]

    Pak, O.V. & Yaroshenko, N.N. (2001) Dopolnitelnye materialy po faune ognevok (Lepidoptera: Pyraloidea) yugo-vostoka Ukrainy [Additional materials to pyraloid fauna (Lepidoptera: Pyraloidea) of the South-Eastern Ukraine]. Bioriznomanittja pryrodnyh i tehnogennyh biotopiv Ukrainy (Materialy Vseukrains’koi konferencii studentiv, aspirantiv ta molodyh vchenyh, Donetsk, 19–22 lystopada 2001 r.), DonNU, Donetsk, 2, pp. 146–152. [in Russian]

    Plant, C.W. (2016) An Annotated Systematic, Synonymic and Distributional Checklist of the Pyraloidea of Bulgaria (Lepidoptera, Crambidae & Pyralidae). Neue Entomologische Nachrichten, 72, 1–231.

    Pohl, G.R., Patterson, B. & Pelham, J.P. (2016) Annotated taxonomic checklist of the Lepidoptera of North America, North of Mexico. Working paper published online by the authors at ResearchGate.net, 1–766 pp.

    Poltavsky, A.N. & Artokhin, K.S. (2018) The Pyraloidea (Lepidoptera) of the Rostov-on-Don Province of Russia, 3. Family Pyralidae. Entomologist’s Gazette, 69, 186–209.

    https://doi.org/10.31184/G00138894.693.1637

    Ragonot, E.L. (1879) Euzophera lafauryella, Lep. Bulletin de la Société entomologique de France, 208–209.

    Ragonot, E.L. (1885) Revision of the British species of Phycitidae and Galleriidae. The Entomologist’s Monthly Magazine, 17–32, 52–60.

    Ragonot, E.L. (1887) Diagnoses d’espèces nouvelles de Phycitidae d’Europe et des Pays limitrophes. Annales de la Société Entomologique de France (Series 6), 7 (3), 225–260.

    Ragonot, E.L. (1888) Nouveaux genres et espèces de Phycitidae & Galleriidae. Publié par l’auteur, 1–52.

    https://doi.org/10.5962/bhl.title.117519

    Ragonot, E.L. (1890) Diagnoses de quelques espèces nouvelles de Phycites recoltées par. M. le Dr Staudinger en Algerie. Bulletin des séances de la Société entomologique de France (Series 6), 10 (2), cix–cxi.

    Ragonot, E.L. & Hampson, G.F. (1901) Monographie des Phycitinae et des Galleriinae. In: Romanoff, N.M. Mémoires sur les Lépidoptères, 8. St. Petersburg, pp. i–xli, 1–602., pls. 24–57.

    Ren, Y. (2012) Phycitinae. In: Li, H.H. (Ed.), Microlepidoptera of Qinling Mountains. Science Press, Beijing, pp. 288–417.

    Robinson, G.S., Ackery, P.R., Kitching, I.J., Beccaloni, G.W. & Hernández, L.M. (2010–2019). HOSTS - a database of the world’s Lepidopteran hostplants. The Natural History Museum, London. http://www.nhm.ac.uk/our-science/data/hostplants/ [accessed 15 May 2019].

    Roesler, R.-U. (1969a) Phycitinen-Studien (Lepidoptera) III. Bonner zoologische Beiträge, 20 (1–3), 257–265.

    Roesler, R.-U. (1969b) Phycitinen-Studien V (Lepidoptera, Pyralidae). Entomologische Zeitschrift, 79 (3), 13–24., figs. 1–7.

    Roesler, R.-U. (1970) Die trifinen Phycitinae. Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei (Lepidoptera, Pyralidae). Reichenbachia, 13 (7), 35–105., figs. 1–30.

    Roesler, R.-U. (1973) Phycitinae. Trifine Acrobasiina. In: Amsel, H.G., Gregor, F. & Reisser, H. (Eds.), Microlepidoptera Palaearctica, 4 (1–2), Georg Fromme & Co., Wien. Part 1, i–xvi, 1–752 pp., part 2, 1–137 pp., pls. 1–170.

    Rothschild, L.W. (1913) Expedition to the central western Sahara by Ernst Hartert. VII. Lepidoptera. Novitates Zoologicae, 20, 109–143.

    Savchuk, V. (2019) Lepidoptera of Crimea. Available from: http://lepidoptera.crimea.ua/families_frame/frame_2000_Pyralidae.htm (accessed 20 June 2019)

    Schille, F. (1930) Fauna motyli Polski. II. Prace Monograficzne Komisji Fizjograficznej, PAU, Kraków, 7, 1–358.

    Shaffer, M., Nielsen, E.S. & M. Horak, M. (1996) Pyraloidea. In: Nielsen, E.S., Edwards, E.D. & Rangsi, T.V. (Eds.), Checklist of the Lepidoptera of Australia. Monographs on Australian Lepidoptera. 4. CSIRO Division of Entomology, Canberra, 164–199 pp.

    Sinev, S.Yu. (1986) 57. Sem. Phycitidae—uzkokrylye ogniovki. [Fam. Phycitidae—narrow-winged pyralids] In: Medvedev, G.S. (Ed.), Opredelitel’ nasekomyh evropeiskoi chasti SSSR. Cheshuekrylye, 4 (3), Nauka (Leningradskoje otdelenie), Leningrad, pp. 251–340. [in Russian]

    Sinev, S.Yu. (1990a) Type specimens of the Phycitidae (Lepidoptera) kept in the collection of the Zoological Institute of the Academy of Sciences of the USSR. I. Entomologicheskoe obsozrenie, 1, 118–133. [in Russian]

    Sinev, S.Yu. (1990b) Type specimens of the Phycitidae (Lepidoptera) kept in the collection of the Zoological Institute of the Academy of Sciences of the USSR. II. Entomologicheskoe obsozrenie, 69 (2), 419–431. [in Russian]

    Sinev, S.Yu. (2008) Pyralidae. In: Sinev, S.Yu. (Ed.), Catalogue of the Lepidoptera of Russia, KMK Scientific Press Ltd., St. Petersburg—Moskow, pp. 156–170. [in Russian]

    Sinev, S.Yu., Anikin, V.V. & Zolotuhin, V.V. (2017) Volga–Ural Pyraloidea and Pterophoridae described by E. Eversmann. In: Anikin, V.V., Sachkov, S.A. & Zolotuhin, V.V. (Eds.), “Fauna Lepidopterologica Volgo-Uralensis”: from P. Pallas to present days. Proceedings of the Museum Witt Munich, Vol. 7, Munich–Vilnius, pp. 380–386., 548–549., pl. 4.

    Slamka, F. (1997) Die Zünslerartigen (Pyraloidea) Mitteleuropas. Bestimmen. Verbreitung. Flugstandort. Lebensweise der Raupen. F. Slamka, Bratislava, 112 pp.

    Slamka, F. (2010) Pyraloidea (Lepidoptera) of Central Europe. Identification. Distribution. Habitat. Biology. F. Slamka, Bratislava, 176 pp.

    Slamka, F. (2013) Pyraloidea of Europe (Lepidoptera). Pyraustinae & Spilomelinae. Identification. Distribution. Habitat. Biology. F. Slamka, Bratislava, 360 pp.

    Sovinskij, V.V. (1926) “Microlepidoptera” z okolyts Dniprjanskoi Biologichnoi Stantsii U.A.N. [Microlepidoptera from the vicinity of Dnipryanska Biological Station of Ukraine Academy of Sciences]. Zbirnyk prats’ Dniprjanskoi Biologichnoi Stantsii, 1 (1), 251–266 [in Ukrainian]

    Sovinskij, V.V. (1935) Über die Pyralididen der Kiewer Provinz. Travaux du Musée zoologique, Kyjv, 15, 47–139. [in Ukrainian]

    Speidel, W. (1996) Pyralidae. In: Karsholt, O. & Razowski, J. (Eds.), The Lepidoptera of Europe. A Distributional Checklist, Apollo Books, Stenstrup, pp. 166–183.

    Stainton, H.T. (1859) Notes on Lepidoptera collected in Madeira by T. V. Wollaston, Esq.; with descriptions of some new species. Annals and Magazine of Natural History, including Zoology, Botany and Geology, (Series 3), 3, 209–214.

    Staudinger, O. (1859) Diagnosen nebst kurzen Beschreibungen neuer andalusischer Lepidopteren. Entomologische Zeitung, 20 (7–9), 211–259.

    Staudinger, O. (1870–1871) Beitrag zur Lepidopterenfauna Griechenlands. Horae Societatis entomologicae Rossicae, 7, 3–304., pls. 1–3.

    Staudinger, O. (1879) Ueber Lepidopteren des südöstlichen europäischen Russlands. Stettiner Entomologische Zeitung, 40, 315–328.

    Stockl, A. (1922) Motyle (Lepidoptera) rzadsz i nowe, zebrine w latach 1911 do 1921 w okolicach Lwowa, Janowa, Mikuliczyna I Worochty.—Czesc III. Polskie pismo entomologiczne, 1 (2), 48–73.

    Strand, E. (1920) Beiträge zur Kenntnis der Verbreitung und Formen exotischer Heterocera, insbesondere Pyralididen, auf Grund von Material im Deutschen Entomologischen Museum. Archiv für Naturgeschichte, 85 (1919), Abt. A (12), 114–166.

    Strelzov, A.N. (2016) Nadsem. Pyraloidea—Ognyovkoobraznye. In: Lelej, A.S. (Ed.), Annotated catalogue of the insects of Russian Far East. Volume 2. Lepidoptera, Dalnauka, Vladivostok, pp. 265–307.

    Toll, S. (1939) Microlepidoptera zebrane v latach 1934–1937 w poviatacj zaleszczyckim i borszczowskim na Podolu. Sprawosdanie Komosji Fizjograficznej, PAU, Kraków, 72, 133–221.

    Treitschke, F. (1832) Die Schmetterlinge von Europa (Fortsetzung des Ochsenheimer’schen Werks). Ernst Fleischer, Leipzig, i–viii, 272 pp.

    Trofimova, T.A. (2017) Pyraloidea and Pterophoridae moths described from the Volga–Ural Region. In: Anikin, V.V., S.A. Sachkov & Zolotuhin, V.V. (Eds.), “Fauna Lepidopterologica Volgo-Uralensis”: from P. Pallas to present days. Proceedings of the Museum Witt Munich. Vol. 7. Munich–Vilnius, pp. 482–491., 620–625., pl. 40–42.

    Tsvetkov, E.V. (2018a) On two species of narrow-winged pyralids, Ancylosis bartelella (Caradja, 1910) and Ancylosis monostictella (Ragonot, 1887) (bona spp.) (Lepidoptera: Pyralidae: Phycitinae). Eversmannia, 55–56, 3–6., pl. 1–2. [in Russian]

    Tsvetkov, E.V. (2018b) Description of two new species of the genus Ancylosis Zeller, 1839 from Russia and Kazakhstan (Lepidoptera: Pyralidae, Phycitinae). SHILAP Revista de Lepidopterologia, 46 (182), 241–249.

    Turati, E. (1924) Spedizione Lepidopterologica in Cirenaica, 1921-1922. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale, 63 (1/2), 21–191., pls. V-VI.

    Turati, E. (1934) Novità di lepidopterologia in Cirenaica IV. Atti della Societa Italiana di Scienze Naturali, 73, 159–212., pl. iii.

    Turati, E. & Zanon, V. (1922) Materiali per una faunula lepidopterologica di Cirenaica (con 19 species e forme nuove). Atti della Societa Italiana di Scienze Naturali, 61, 138–178., pl. A.

    Warren, W. (1888) On Lepidoptera collected in Major Yerbury in Western India in 1886 and 1887. Proceedings of the General Meetings for Scientific Business of the Zoological Society of London, 292–339.

    https://doi.org/10.1111/j.1469-7998.1888.tb06711.x

    Whalley, P.E.S. (1970) A synonymic catalogue of the genera of Phycitinae (Lepidoptera: Pyralidae) of the world. Bulletin of the British Museum (Natural History) Entomology series, 25 (2), 33–72.

    Zeller, P.C. (1839) Versuch einer naturgemäßen Eintheilung der Schaben. Isis von Oken, Leipzig, [32] (3), 167–219 pp.

    Zeller, P.C. (1848) Die Gallerien und nackthornigen Phycideen. Isis von Oken, Leipzig, (8–10), pp. (8) 569–618., (9) 641–691., (10) 721–754.

    Zeller, P.C. (1867) Crambina, Pterophorina and Alucitina, collected in Palestine, by the Rev. O. P. Cambridge, March to May, 1865. Transactions of the Entomological Society of London, (Series 3), 5 (6), 453–460., pl. 23.

    Zerny, H. (1914) Über paläarktische Pyraliden des k. k. naturhistorischen Hofmuseums in Wien. Annalen des naturhistorischen Hofmuseums, 28 (3–4), 295–348., pls. 25–26.

    Zhikharev, I. (1928) Shkidlyvi ta inshi luskokryltsi (Lepidoptera) Darnytskoi lisovoi doslidnoi dachi [Pests and other Lepidoptera of Darnytska forest experimental station]. Trudy z Lisovoi Doslidnoi Spravy na Ukraini, 9, 231–330. [in Ukrainian]