Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2019-08-29
Page range: 457–493
Abstract views: 95
PDF downloaded: 2

A taxonomic revision of the genus Xylota Meigen (Diptera: Syrphidae) in Korea

Division of Biological Science and Technology, Yonsei University, 1 Yonseidae-gil, Wonju-si, Gangwon-do 26493, Republic of Korea.
Division of Biological Science and Technology, Yonsei University, 1 Yonseidae-gil, Wonju-si, Gangwon-do 26493, Republic of Korea.
Diptera Insecta Xylotina taxonomy hoverfly genitalia new species

Abstract

As a result of the study of the hoverfly subtribe Xylotina in Korea, we have recognized the following 14 Xylota species, including eight new to the Korean syrphid fauna (marked with asteristks) and three new to science: X. abiens, X. amaculata*, X. atricoloris*, X. coquilletti, X. filipjevi*, X. fo*, X. ignava, X. pseudoignava*, X. spurivulgaris*, X. tarda*, X. umbrosa*, X. hauseri sp. n., X. orientiflorum sp. n., and X. xanthotarsis sp. n. Among these species, we clarified the identities of three previously confused species, X. coquilletti, X. spurivulgaris and X. fo, which had been collectively identified as a single species, X. coquilletti, in the Korean literature. In addtion, we provided unequivocal characters to separate two sibling species pairs based on their external characters including genitalic structure (X. hauseri sp. n. vs. X. umbrosa; X. pseudoignava vs. X. xanthotarsis sp. n.; X. filipjevi vs. X. tarda). In order to aid accurate identification of the Korean Xylota, diagnoses, descriptions, and color photographs of external structures including male genitalia are provided. For the taxonomic key, we included all the Korean genera of the subtribe Xylotina (Xylota, Brachypalpus and Chalcosyrphus).

 

References

  1. Bagatshanova, A. (1980) New species of Syrphidae (Diptera) from central Yakutia. Entomologitsheskoe Obrozenie, 59, 421–427. [in Russian]

    Bańkowska, R. (1998) Syrphid flies (Syrphidae, Diptera) from North Korea collected by Polish expeditions. Pt. 1,(Milesiinae, Microdontinae). Fragmenta Faunistica, 41, 167–182.

    https://doi.org/10.3161/00159301FF1998.41.12.167

    Bartsch, H., Binkiewicz, E., Klintbjer, A., Rådén, A. & Nasibov, E. (2009) Nationalnyckeln till Sveriges flora och fauna. Tvåvingar: Blomflugor: Eristalinae & Microdontinae. Diptera: Syrphidae: Eristalinae & Microdontinae. ArtDatabanken, SLU, Uppsala, 470 pp. [in Swedish]

    Becker, T. (1921) Neue Dipteren meiner Sammlung. Mitteilungen aus dem Zoologischen Museum in Berlin, 10 (1), 1–93.

    Bigot, J.M.F. (1861) Trois dipteres nouveaux de la Corse. Annales de la Société Entomologique de France, 1, 227–229. [in French]

    Bigot, J.M.F. (1885) Diptères nouveaux ou peu connus. 25e partie. XXXIII. Anthomyzides nouvelles. Annales de la Société Entomologique de France, 4, 263–304.

    Coquillett, D.W. (1898) Report on a Collection of Japanese Diptera, Presented to the U. S. National Museum by the Imperial University of Tokyo. Proceedings of the United States National Museum, 21, 301–340.

    https://doi.org/10.5479/si.00963801.21-1146.301

    Curran, C.H. (1925) Contribution to a Monograph of the American Syrphidae from North of Mexico. Bulletin of the University of Kansas Science Bulletin, 15, 1–283.

    Curran, C.H. (1941) New American Syrphidae. Bulletin of the American Museum of Natural History, 78, 243–304.

    Doi, H. (1938) A List of the Diptera of Korea. Bulletin of Science Museum, Keizyo, Korea, 72, 8–11. [in Japanese]

    Entomological Society of Korea and Korean Society of Applied Entomology (ESK and KSAE) (1994) Syrphide. In: Check list of insects from Korea. Kon-kuk University press, Seoul, pp. 288–292. [in Korean]

    Fabricius, J.C. (1794) Entomologia Systematica Emendata et aucta. Secundum Classes, Ordines, Genera, Species Adjectis Synonymis, Locis, Observationibus, Descriptionibus. Vol. 4. impensis Christ. Gottl. Proft, Hafniae [=Copenhagen], 472 pp.

    Fabricius, J.C. (1805) Systema antliatorum secundum ordines, genera, species. C. Reichard, Brunsvigae, 30 pp.

    Han, H.Y. & Choi, D.S. (2001) Family Syrphidae. Economic Insects of Korea 15. Insecta Koreana, 22 (Supplement), 1–224. [in Korean]

    Han, H.Y., Choi, D.S., Kim, J.I. & Byun, H.W. (1998) A catalog of the Syrphidae (Insecta: Diptera) of Korea. Insecta Koreana, 15, 95–166.

    Han, H.Y., Suk, S.W., Lee, Y.B. & Lee, H.S. (2014) National List of Species of Korea, Insect (Diptera II). National Institute of Biological Resources, Incheon, 268 pp. [in Korean]

    Harris, M. (1776–1780) An exposition of English Insects, with curious observations and remarks, wherein each insect is particularly described; its parts and properties considered; the different Sexes distinguished, and the natural history faithfully related. Robson, London, 8 pp.

    https://doi.org/10.5962/bhl.title.122956

    He, J. & Chu, X. (1992) Studies on three genera of Xylotini (Sensu Hippa) from China (Diptera: Syrphidae). Journal of Shanghai Agricultural College, 10, 1–12. [in Chinese]

    Hendel, F. (1908) Diptera. Fam. Muscaridae, Subfam. Pyrgotinae. In: Wytsman, P. (Ed.), Genera insectorum, 79, pp. 1–33.

    Hervé-Bazin, J. (1914) Syrphides recueillis au Japon par M. Edme Gallois (Diptera). Annales de la Société Entomologique de France, 83, 398–416.

    https://doi.org/10.5962/bhl.part.26806

    Hippa, H. (1968) Classification of the Palaearctic species of the genera Xylota Meigen and Xylotomima Shannon (Dipt., Syrphidae). Annales Entomologicae Fennicae, 34, 179–197.

    Hippa, H. (1978) Classification of Xylotini (Diptera, Syrphidae). Acta zoological fennica, 156, 1–153.

    Hokuryukan, Co. (1956) Iconographia Insectorum Japonicorum. Hokuryukan, Tokyo, 228 pp. [pp. 1494–1721, in Japanese]

    Hokuryukan, Co. (1965) Iconographia Insectorum Japonicorum, Colore Naturali Edita. Vol. III. Hokuryukan, Tokyo, 358 pp., 156 colour pls. [in Japanese]

    Huang, C. & Cheng, X. (2012) Fauna Sinica: Diptera Syrphidae. Vol. 50. Insecta. Science press, Beijing, 852 pp. [in Chinese]

    Huang, C., Cheng, X. & Yang, C. (1996) Syrphidae. In: Xue, W. & Chao, C. (Eds.), Flies of China, I, pp. 118–233. [in Chinese]

    Hull, F.M. (1936) A Check list of the described species of Syrphidae from Australia and the Regional Islands. Journal of the Federated Malay States Museum, 18, 190–212.

    Hull, F.M. (1944) Two species of Xylota from southern Asia (Diptera: Syrphidae). Proceedings of the Entomological Society of Washington, 46, 45–47.

    Huo, K.-K., Ren, G.-D. & Zheng, Z.-M. (2007) Fauna of Syrphidae from Mt. Qinling-Bashan in China (insecta: Diptera). Science press, Beijing, 512 pp. [in Chinese]

    International Commission on Zoological Nomenclature (1963) Opinion 678. The suppression under the plenary powers of the pamphlet published by Meigen, 1800. Bulletin of zoological nomenculature, 20 (5), 339–342.

    Ito, S., Ikutani, T. & Hiura, I. (1993) Colored Illustrations of the Insects of Japan II. Hoikusha Publishing Co. Ltd., Osaka, pp. 252–256. [in Japanese]

    Jeong, S.H., Jung, J.M. & Han, H.Y. (2017) A taxonomic review of Brachypalpus Macquart and Chalcosyrphus Curran (Insecta: Diptera: Syrphidae) in Korea. Journal of Asia-Pacific Entomology, 20, 1043–1061.

    https://doi.org/10.1016/j.aspen.2017.07.001

    Kim, C.W. & Kim, J.I. (1972a) Insect fauna of Gucheondong, Muju-gun. Report of Korean Association for Conservation of Nature, 5, 91–93. [in Korean]

    Kim, C.W., Park, S.H., Lee, K.W., Lee, C.U., Lee, T.J. & Cho, B.S. (1971a) Family 26. Syrphidae. In: Kim, C.W., Park, S.H., Lee, K.W., Lee, C.U., Lee, T.J. & Cho, B.S. (Eds.), Illustrated Encyclopedia of Fauna and Flora of Korea. Vol. 12. Samhwa Publishing Co., Seoul, pp. 862–865. [In Korean]

    Kim, C.W. & Nam, S.H. (1982a) Changes in insect fauna due to urbanization of Seoul city. Bulletin of Korean Association for Conservation of Nature, 4, 249–296. [in Korean]

    Kim, C.W. & Nam, S.H. (1982b) Insect fauna in the areas of Mts. Gyebang, Sogyebang and Gachilbong in summer season. Report of Korean Association for Conservation of Nature, 20, 119–137. [in Korean]

    Kim, C.W. & Nam, S.H. (1982c) Insect Fauna of Seoul City Area. The Korean National Council for Conservation of Nature, 23, 125–176. [in Korean]

    Kim, C.W., Shin, Y.H. & Kim, J.I. (1971b) Report on the scientific survey of island Woolung-do (Insecta). Report of Korean Association for Conservation of Nature, 3, 47–62. [in Korean]

    Kim, J.I. (1975) A list of Syrphidae (Diptera) from Korea. Korean Journal of Entomology, 5, 38–42. [in Korean]

    Kim, J.I. (1980) The historical review and the tentative list of the Korean Syrphidae (Diptera). Sungshin University, 13, 365–389. [in Korean]

    Kim, J.I. (1995a) Coleoptera and Diptera (Insecta) from Mt. Sobaek. Report of Korean Association for Conservation of Nature, 33, 157–179. [in Korean]

    Kim, J.I. (1995b) Fauna of Coleoptera and Diptera (Insecta) from Pyonsan peninsula National Park. Report of Korean Association for Conservation of Nature, 34, 129–145. [in Korean]

    Kim, J.I. (1996) Insect fauna of Coleoptera and Diptera from Mt. Pangtae in summer season. Report of Korean Association for Conservation of Nature, 35 (1995), 163–179. [in Korean]

    Kim, J.I., Kim, B.J., Lee, O.K. & Park, H.C. (1991) Faunastic study on the insect from Mt. Songni. Report of Korean Association for Conservation of Nature, 29, 163–193. [in Korean]

    Kim, J.I. & Park, H.C. (1991) A Faunistic study of terrestrial insects in the Daechung lake. Report of Survey for Daechung lake Ecosystem, Chungcheongbuk-do. Korean Jornal of Entomology, 29, 215–236. [in Korean]

    Kim, J.I., Park, J.M., Kim, S.Y. & Choi, H.S. (1994a) fauna for Coleoptera, Hymenoptera and Diptera, in Mt.Kwangdok. Report of Detailed Area-wide Ecosystem Survey, Korean Ministry of Environment, 111, 101–116. [in Korean]

    Kim, J.I., Lee, E.J. & Kim, S.Y. (1994b) Insects fauna from the Mt. Gyebang, Kangwon-do. Korea. Report of Detailed Area-wide Ecosystem Survey, Korean Ministry of Environment, 147, 135–159. [in Korean]

    Kim, J.I. & Park, H.K. (1984) Taxonomical study on the Syrphidae (Diptera) from the anatomical characters of the digestive organ. Journal of Institute for Basic Science, 1, 41–49. [in Korean]

    Kim, J.I. & Yoo, H.J. (1987) Study on the insects fauna and its change (succession) from near DMZ of the province Kyonggi-do, Korea. Report on the Environmental Study of near DMZ, Korea. The Korean National Council for Conservation of Nature, Seoul, 40 pp. [pp. 489–528 pp., in Korean]

    Kim, J.I., Kim, T.W., Kim, A.Y., Jung, B.H. & Yoo, I.S. (2006) A Faunistic Study on the Insects from Mt. Achasan, Korea. Korean Journal of Nature Conservation, 4, 27–49.

    https://doi.org/10.30960/kjnc.2006.4.1.27

    Kwon, Y.J., Suh, S.J., An, S.L. & Huh, E.Y. (1996) Insect Diversity of Ullungdo and Tokto Islands in Korea. Report on the Survey of Natural Environment in Korea, 10, 441–532. [in Korean]

    Lee, C.E. & Kwon, Y.J. (1981) On the insect fauna of Island Ulreung and Dogdo in Korea. Report of Korean Association for Conservation of Nature, 19, 139–179. [in Korean]

    Lepeletier, A.L.M. & Serville, J.G.A. (1828) [Various articles]. In: Latreille, P.A., Lepeletier, A.L.M., Serville, J.G.A. & Guerin-Meneville, F.E. (Eds.), Entomologie, ou histoire naturelle des Crustaces, des Arachnides et des insectes. Encyclopédie Méthodique, Histoire Naturelle. Insectes. Vol. 10. Agasse, Paris, pp. 345–833.

    Linnaeus, C. (1758) Systema Naturae per regna trio naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus 1. Editio Decima, Reformata, Holmiae, 824 pp.

    https://doi.org/10.5962/bhl.title.542

    Lim, J.S., Lee, B.W., Park, S.Y. & Jo, D.G. (2011) Insect fauna of Maebongsan mountain, Hongcheon-gun, Gangwon-do. Journal of Korean Nature, 4, 293–307. [in English]

    https://doi.org/10.7229/jkn.2011.4.4.293

    Loew, H. (1871) Beschreibungen europischer Dipteren. Systematische Beschreibung der bekannten europäischen zweiflügligen Insekten. von Johann Wilhelm Meigen. II. Halle, 319 pp.Matsumura, S. (1911) Erster Beitrag zur Insekten-Fauna von Sachalin. Journal of the College of Agriculture, Imperial University of Tokyo, 4, 72–80.

    Macquart, J. (1834) Histoire naturelle des Insectes—Dipterès. In: Roret, N.E. (Ed.), Collection des suites à Buffon, formant avec les oeuvres de cet auteur un cours complet d’histoire naturelle. Librairie Encyclopédique de Roret, Paris, pp. 1–578.

    https://doi.org/10.5962/bhl.title.14274

    Matsumura, S. (1931) 6000 Illustrated Insects of Japan-Empire. Tokoshoin, Tokyo.

    Matsumura, S. & Adachi, J. (1916) Synopsis of the Economic Syrphidae of Japan (I). The Entomological Magzine, Kyoto, Japan, 2, 1–36.

    Meigen, J.W. (1800) Nouvelle classification des mouches a deux ailes (Diptera L.) d’apres un plan tout nouveau. Perronneau, Paris, 40 pp.

    https://doi.org/10.5962/bhl.title.119764

    Meigen, J.W. (1803) Versuch Einer Neuen Gattung Einteilung der Europäischen Zweiflügeligen Insekten. Magazin fur Insektenkunde, Herausgegeben von Karl Illiger, 2, 259–281.

    Meigen, J.W. (1822) Systematische Beschreibung der bekannten Europäischen zweiflügeligen Insekten. Dritter Theil. Schultz-Wundermann, Hamm, x + 416 pp., pls. 22–32.

    Mengual, X., Ståhls, G. & Rojo, S. (2015) Phylogenetic relationships and taxonomic ranking of pipizine flower flies (Diptera: Syrphidae) with implications for the evolution of aphidophagy. Cladistics, 31, 491–508.

    https://doi.org/10.1111/cla.12105

    Mutin, V.A. (1984) Two new species of syrphid flies (Diptera) from the south of the Soviet Far East. Zoologicheskii Zhurnal, 63, 783–785.

    Mutin, V.A. (1988a [1987a]) New data on hover-flies of genera Xylota Mg. and Chalcosyrphus Curr. (Diptera, Syrphidae). In: Lehr, P.A. & Kanyukova, E.V. (Eds.), Taksonomiya nasekomykh Sibiri i Dal’nego Vostoka SSSR. DV NTS AN SSSR, Vladivostok, pp. 102–106. [in Russian]

    Mutin, V.A. (1988b [1987b]) New species of hover-flies of genus Xylota Mg. (Diptera, Syrphidae) from the south of Russian Far East. In: Lehr, P.A. & Storozheva, N.A. (Eds.), Novye dannye o sistematike nasekomykh Dal’nego Vostoka. DVO AN SSSR, Vladivostok, pp. 119–121. [in Russian]

    Mutin, V.A. & Barkalov, A.V. (1999) 62. Family Syrphidae. In: Lehr, P.A. (Ed.), Key to the insects of Russian Far East. Vol. 6. Diptera and Siphonaptera. Part 1. Dal’nauka, Vladivostok, pp. 342–500. [in Russian]

    Mutin, V.A. & Gilbert, F. (1999) Phylogeny of the genus Xylota Meigen, 1822 (Diptera, Syrphidae) with descriptions of new taxa. Dipteron, 2, 45–68.

    Mutin, V.A. & Ichige, K. (2014) A new species of Xylota Meigen (Diptera: Syrphidae) from the Far East. Zootaxa, 3878 (2), 196–200.

    https://doi.org/10.11646/zootaxa.3878.2.6

    Ohara, K., Ohishi, H. & Ichige, K. (2014) Catalog of the insects of Japan. Vol. 8. Part 1. Diptera (Nematocera—Brachycera Aschiza). Entomological society of Japan, Touka Shobo, Fukuoka, xxxii + 539 pp. [in Japanese]

    Paek, M.K., Hwang, J.M., Jung, K.S., Kim, T.W., Kim, M.C., Lee, Y.J., Cho, Y.B., Park, S.W., Lee, H.S., Ku, D.S., Jeong, J.C., Kim, K.G., Choi, D.S., Shin, E.H., Hwang, J.H, Lee, J.S., Kim, S.S. & Bae, Y.S. (2010) Checklist of Korean insects. In: Peak, M.K. &Cho, Y.K. (Eds.), Nature & Ecology Academics. Series 2, Nature & Ecology, Seoul, pp. 1–598.

    Panzer, G.W.F. (1798) Faunae insectorum germanicae initia oder Deutschlands Insecten. 54, 59 & 60. Felsecker, Nurnberg [= Nuremberg], pp. 1–24.

    Panzer, G.W.F. (1806) Kritische Revision der Insektenfaune Deutschlands. Vol. 2. Nürnberg, In der Felfseckerschen Buchhandlung, 271 pp.

    Pape, T. & Thompson, F.C. (Eds.) (2013) Systema Dipterorum. Version [1.5]. Available from: http://www.diptera.org/ (accessed 31 October 2016)

    Park, J.S., Ku, D.S. & Han, K.D. (1993) Faunistic study on the insect from Hamyang-gun and Paemsagol area of Mt. Chiri. Report of Korean Association for Conservation of Nature, 31, 153–218.

    Park, J.S. (1996) Environment impact statement of Mt. Chiaksan. Korea national park service, Seoul, 64 pp. [pp. 130–193, in Korean]

    Park, K.T. (1998) Insects in Kangwon-do. Center for Insect Systematics and Institute for Biodiversity Research, Kangwon University, Chuncheon, 155. [in Korean]

    Peck, L.V. (1988) Family Syrphidae. In: Soos, A. (Ed.), Catalogue of Palaearctic Diptera. Vol. 8. Akademiai Kiado, Budapest, pp. 11–230.

    Philippi, R.A. (1865) Aufzahlung der chilenischen Dipteren. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 15, 595–782.

    https://doi.org/10.5962/bhl.title.9295

    Rotheray, G.E. & Gilbert, F. (2011) The natural history of hoverflies. Forrest text, Tresaith, Wales, 333 pp.

    Sack, P. (1932) 31. Syrphidae. In: Lindner, E. (Ed.), Die Fliegen der Palaearktischen Region. Band IV/6. E. Schweizerbart’sche Verlagsbuchhandlung, Stuttgart, pp. 1–451. [in German]

    Shannon, R.C. (1926) Review of the American xylotine syrphid-flies. Proceedings of the United States National Museum, 69, 1–52.

    Schrank, F.V.P. (1803) Fauna Boica. Durchgedachte Geschichte der in Baiern einheimischen und zahmen Thiere. Landshut, 3, 1–272.

    Shiraki, T. (1930) Die Syrphiden des Japanischen Kaiserreichs, mit Berucksichtigung benachbarter Gebiete. Memoirs of the Faculty of Science and Agriculture, Taihoku Imperial University, 1, 1–446.

    Shiraki, T. (1968) Fauna Japonica, Syrphidae (Insecta). Vol. 3. Biogeographical Society Japan, Tokyo, 272 pp.

    Speight, M.C.D. (1999) A key to European Xylotini (Dip.: Syrphidae). Entomologist’s Record, 111, 211–218.

    Stackelberg, A.A. (1952) Brief review of palaearctic species of the genus Zelima Mg. (Diptea, Syrphidae). Entomologiceskoe obozrenie, 32, 316–328. [in Russian]

    Stackelberg, A.A. (1964) Notes on palearctic Syrphidae (Diptera). Zoologicheskii Zhurnal, 43, 467–473.

    Ståhls, G. (2006) Placement of Cacoceria and phylogenetic relationships of the xylotine genera of the tribe Milesiini (Diptera, Syrphidae: Eristalinae) based on molecular characters. Zootaxa, 1171 (1), 17–29.

    https://doi.org/10.11646/zootaxa.1171.1.2

    Ståhls, G., Hippa, H., Rotheray, G., Muona, J. & Gilbert, F. (2003) Phylogeny of Syrphidae (Diptera) inferred from combined analysis of molecular and morphological characters. Systematic Entomology, 28, 433–450.

    https://doi.org/10.1046/j.1365-3113.2003.00225.x

    Stireman, J.O., Cerretti, P., O’Hara, J.E., Blaschke, J.D. & Moulton, J.K. (2019) Molecular phylogeny and evolution of world tachinidae (Diptera). Molecular Phylogenetics and Evolution. [in press]

    https://doi.org/10.1016/j.ympev.2018.12.002

    Stubbs, A.E. & Falk, S. (2002) British hoverflies, an illustrated identification guide. Second edition. British Entomological and Natural History Society, Penzance, Cornwall, 253 pp., 13 pls., supplement I–XV.

    Thompson, F.C. (1975) Notes on the status and relationships of some genera in the tribe Milesiini (Diptera: Syrphidae). Proceedings of Entomological Socieyt of Washington, 77, 291–305.

    Thompson, F.C. (1999) A key to the genera of the flower flies (Diptera: Syrphidae) of the Neotropical Region including descriptions of new genera and species and a glossary of taxonomic terms. Contributions on Entomology International, 3, 321–378.

    Thompson, F.C. & Vockeroth, J.R. (1989) Family Syrphidae, In: Evenhuis, N.L. (Ed.), Catalogue of the Diptera of Australasia and Oceania. Bishop Museum Special Publication 86. Bishop Museum Press, Honolulu, pp. 437–456.

    van Veen, M.P. (2004) Hoverflies of northwest Europe: identification keys to the Syrphidae. KNNV, Utrecht, 254 pp.

    https://doi.org/10.1163/9789004274495_004

    Violovitsh, N.A. (1975) Some new species of hover-flies (Diptera. Syrphidae) from the fauna of the USSR. Novye i meloizvestnye vidy fauny Sibiri, Vyp. 9, 73–89. [in Russian]

    Violovitsh, N.A. (1983) Syrphids of Siberia (Diptera, Syrphidae). Nauka, Novosibirsk, 242 pp. [in Russian]

    Violovitsh, N.A. (1986) Siberian Syrphidae (Diptera) (Sirfidy Sibiri) Novosibirsk, 1983. Verslagen en Technische Gegevens Instituut voor Taxonomische Zoologie (Zoologische Museum) Universiteit van Amsterdam, No. 43, 1–228. [English translation of Violovitsh (1983)]

    Vujić, A. & Milankov, V. (1999) New data for the tribes Milesiini and Xylotini (Diptera, Syrphidae) on the Balkan peninsula. Dipteron, 2, 113–132.

    Westwood, J.O. (1840) Order XIII. Diptera Aristotle. (Antliata Fabricius. Halteriptera Clairv.). In: Synopsis of the genera of British Insects. An Introduction to the Modern classification of Insects; founded on the natural habits and corresponding organization of the different families. Longman, Orme, Brown, Green, and Longmans, London, 158 pp.

    https://doi.org/10.5962/bhl.title.12455

    Williston, S.W. (1882) Contribution to a monograph of the North American Syrphidae. Proceedings of the American Philosophical Society, 20, 299–332.

    https://doi.org/10.5962/bhl.title.40965

    Williston, S.W. (1891) Fam. Syrphidae. In: Godman, F.D. & Salvin, O. (Eds.), Biologia Centrali-Americana. Zoologia-Insecta-Diptera. Taylor & Francis, London, pp. 1–56.

    Wirth, W.W., Sedman, Y.S. & Weems, H.V. (1965) Syrphidae. In: Stone, A., Sabrosky, C.W., Wirth, W.W., Foote, R.H. & Coulson, J.R. (Eds.), A catalog of the Diptera of America north of Mexico. USDA Agricultural Handbook, 276, USDA Agricultural, Washington, D.C., pp. 1–1096.

    Yang, C. & Cheng, X. (1993) A study of the genus Milesia Latreille from Fujian, with descriptions of three new species (Diptera: Syrphidae). Wuyi Science Journal, 10, 41–49.

    Yang, C. & Cheng, X. (1998) Syrphidae. In: Xue, W. & Chao, C. (Eds.), Flies of China. Vol. 1. Liaoning Science & Technology Press, Shenyang, pp. 118–223. [1996]

    Yoon, I.B. & Nam, S.H. (1980) Insect fauna of Mts. Chilgab and Gyeryong. Report of Korean Association for Conservation of Nature, 17, 129–158. [in Korean]

    Young, A.D., Lemmon, A.R., Skevington, J.H., Mengual, X., Ståhls, G., Reemer, M., Jordaens, K., Kelso, S., Lemmon, E.M. & Hauser, M. (2016) Anchored enrichment dataset for true flies (order Diptera) reveals insights into the phylogeny of flower flies (family Syrphidae). BMC evolutionary biology, 16, 143.

    https://doi.org/10.1186/s12862-016-0714-0

    Zetterstedt, J.W. (1838) Dipterologis Scandinaviae. Sect. 3: Diptera. In: Zetterstedt, J.W. (Ed.), Insecta Lapponica. Voss, Lipsiae [= Leipzig], pp. 477–868.

    Zoological Society of Korea (ZSK) (1968) Nomina Animalium Koreanorum (2) Insecta. Zoological Society ofKorea, Seoul, 334 pp. [pp. 172–187, in Korean]