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Published: 2007-04-25
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A review of the Aphodiinae (Coleoptera: Scarabaeidae) of southern South America

Research Division, Canadian Museum of Nature, P.O. Box 3443, Station D, Ottawa, ON, K1P 6P4, CANADA.
Florida Department of Agriculture and Consumer Services, Florida State Collection of Arthropods, 1911 SW 34th St., P.O. Box 147100, Gainesville, FL, 32614-7100, USA.
Coleoptera Argentina Chile Aphodiinae nomenclature classification

Abstract

The 27 Aphodiinae (Coleoptera: Scarabaeidae) species that occur in southern South America are reviewed. This study region consists of the southern parts of Argentina and Chile including the Central Chilean, Patagonian, and Subantarctic biogeographical provinces. Keys, illustrations, and distributional records with maps for all species are presented. Seventeen species are native to southern South America, nine species are introduced, and the status of one species could not be determined due to a lack of study material. Nine of the 17 native species are endemic to southern South America, and the relationship between these endemic species and similar Australian groups is discussed. A number of classification and nomenclatural changes are made. Paranimbus Schmidt and Pseudopodotenus Dellacasa are considered to subgenera of Podotenus Schmidt. As a result, the following new combinations are created: Podotenus (Paranimbus) longitarsis (Harold), Podotenus (Paranimbus) penai (Petrovitz), Podotenus (Paranimbus) peruanus (Erichson), Podotenus (Paran- imbus) zoiai (Dellacasa), and Podotenus (Pseudopodotenus) fulviventris (Fairmaire and Germain). Ataenius chilensis (Solier) is removed from synonymy with Ataenius gracilis (Melsheimer) and is considered a valid species. Ataenius modestus Schmidt is synonymized under Ataenius chilensis (Solier). Psammobius indefensus Schmidt is transferred to the genus Leiopsammodius and Leiopsammodius chilensis Rakovic is placed in synonymy with this name. Lectotypes are designated for the following names: Aphodius chilensis Solier (now Ataenius chilensis), Aphodius reflexus Schmidt (now Orodaliscoides reflexus), Ataenius modestus (now a synonym of Ataenius chilensis), and Psammobius indefensus Schmidt (now Leiopsammodius indefensus). A neotype is designated for Aphodius angustus Philippi and Philippi, a synonym of Pleurophorus caesus (Panzer). Records considered to be erroneous for several Aphodiinae species in southern South America are also discussed.

References

  1. Arrow, G.J. (1903) On the Laparostict Lamellicorn Coleoptera of Grenada and St. Vincent. Transaction of the Entomological Society of London, 1903, 509–520.

    Balthasar, V. (1936) Neue Aphodius-Arten aus China und Syrien. Entomologisches Nachrichtenblatt, 11, 1–6.

    Balthasar, V. (1941) Gli Scarabeidi coprofagi e Trogini dell’Africa orient. Ital. del Museo di Milano. Atti de la Societa Italiana di Scienze Naturali del Museo Civico di Soria, 80, 103–154.

    Balthasar, V. (1942) Neue Arten und Gattungen der coprophagen Scarabaeiden. 75. Beitrag zur Kenntnis der Scarabaeiden (Col.). Acta Entomologica Musaei Pragae, 20, 188–206.

    Balthasar, V. (1946) De novis generis Aphodius Illig. speciebus. Acta Entomologica Musei Nationalis Pragae, 24, 53–68.

    Balthasar, V. (1947) De novis generis Ataenius Har. speciebus. Acta Entomologica Musei Nationalis Pragae, 25, 49–54.

    Balthasar, V. (1960) Beiträge zur Kenntnis der Insektenfauna Boliviens Teil XIII. Coleoptera XII. Neue Aphodiinae-Arten (100. Beitrag zur Kenntnis der Scarabaeidae). Opuscula Zoologica, 48, 1–7.

    Balthasar, V. (1961) Eine neue Gattung und neue Arten der Unterfamilie Aphodiinae. Deutsche Entomologische Zeitschrift, 8(1/2), 121–130.

    Balthasar, V. (1965) Eine neue Gattung und Untergattung und neue Arten der Scarabaeoidea. (117. Beitrag zur Kenntnis der Scarabaeoidea, Col.). Deutsche Entomologische Zeitschrift, 12, 311–323.

    Bates, H.W. (1887) Pectinicornia and Lamellicornia, pp 25–160. In: F. D. Godman and O. Salvin (eds.), Biologia Centrali-Americana. Insecta. Coleoptera, volume 2, part 2. 432 pp, 24 plates.

    Blackburn, T. (1892) Further notes on Australian Coleoptera, with descriptions of new genera and species. XI. Transactions of the Royal Society of South Australia, 15, 20–73.

    Blackburn, T. (1904) Further notes on Australian Coleoptera, with descriptions of new genera and species. XXXIV. Transactions of the Royal Society of South Australia, 28, 281–297.

    Blanchard, C.É. (1847) Insectes de l’Amérique Méridionale. In: A. d’Orbigny, C. É. Blanchard, and A. Brullé, Voyage dans l’Amérique Méridionale, volume 6, part 2, livraison 90. P. Bertrand, Paris. Pp 185–232.

    Boheman, C.H. (1858) Kongliga Svenska Fregatten Eugenies Resa omkring Jorden unter Befäll af C. A. Virgin 1851–53. Vetenskapliga Jakttagelser (Coleoptera). Norstedt, Stockholm. 112 pp.

    Bouskell, F. (1901) The variation and distribution of the genus Aphodius (Illiger). Transactions of the Leicester Literary and Philosophical Society, 5, 1–35.

    Bruch, C. (1911) Catálogo Sistemático de los Coleópteros de la República Argentina. Pars IV. Familias Lucanidae, Scarabaeidae (Coprini-Cetonini), Passalidae. Revista del Museo de La Plata, 17, 181–225.

    Bruch, C. (1938) Algunas especies nuevas de coleopteros. In: Miscelaneas entomologicas. Notas del Museo de La Plata, 3, 160–164.

    Campos, L. & Peña, L.E. (1973) Los insectos de Isla de Pascua. Revista Chilena de Entomologia, 7, 217–229.

    Cartwright, O.L. (1946) Two new Psammodius from southeastern United States (Coleoptera: Scarabaeidae). Bulletin of the Brooklyn Entomological Society, 41, 89–91.

    Cartwright, O.L. (1948) The American species of Pleurophorus (Coleopera: Scarabaeidae). Transactions of the American Entomological Society, 74, 131–145.

    Cartwright, O.L. (1955) Scarab beetles of the genus Psammodius in the Western Hemisphere. Proceedings of the United States National Museum, 104(3344), 413–462.

    Cartwright, O.L. (1973) Additional lectotype designations in the Aphodiinae (Coleoptera: Scarabaeidae). Coleopterists Bulletin, 27, 41–43.

    Chalumeau, F. (1983) Batesiana et Martinezia, nouveaux genres d’Eupariini (Coleoptera: Scarabaeidae: Aphodiinae) du nouveau monde. Bulletin Mensuel de la Société Linnéenne de Lyon, 52, 142–153.

    Chalumeau, F. (1990) Un nouvel Ataenius du Bresil: A. degallieri (Coleoptera Scarabaeidae Aphodiinae). Bulletin Mensuel de la Société Linnéenne de Lyon, 59, 304–305.

    Chalumeau, F. (1992) Eupariini du nouveau monde: un mise au point (Coleoptera, Scarabaeidae) (1re partie). Nouvelle Revue d'Entomologie, 9(3), 189–206.

    Chapin, E.A. (1940) A revision of the West Indian beetles of the scarabaeid subfamily Aphodiinae. Proceedings of the United States National Museum, 89, 1–41

    Creutzer, C. (1799) Entomologische Versuche (Kleine Beitrage zur nahren Berichtigungen einiger Kaferarten mit 3 ausgemahften Kupfertafeln von Sturm J.). Wien, Austria.

    Crisci, J.V., Cigliano, M.M., Morrone, J.J. & Roig Juñent, S. (1991) Historical biogeography of southern South America. Systematic Zoology, 40, 152–171.

    von Dalla Torre, K.W. (1879) Systematisches Verzeichnis der in Oberösterreich bisher beobachteten Käfer. Jahres-Bericht des Vereines für Naturkunde in Oesterreich ob der Ens, 10, 1–125.

    Darlington, P.J. (1965) Biogeography of the Southern End of the World. Harvard University Press, Cambridge, Massachusetts.

    Dellacasa, G. (1990) A systematic revision of the Aphodius subgenus Paranimbus A. Schmidt with description of a new species. Bollettino del Museo Regionale di Scienze Naturali, Torino, 8(2), 417–428.

    Dellacasa, G. (1992) On some little known species of Aphodius from Australia and South America with description of a new subgenus. Frustula Entomologica, 26, 145–158. [Dated 1990].

    Dellacasa, G., Bordat P. & Dellacasa, M. (2001) A revisional essay of world genus-group taxa of Aphodiinae (Coleoptera: Aphodiidae). Memorie della Societa Entomologica Italiana, 79, 1–482.

    Dellacasa, G. & Gordon, R.D. (1997) Nearctic and neotropical genus-group taxa of Aphodiini and their type species (Coleoptera: Aphodiidae). Annali del Museo Civico di Storia Naturale “Giacomo Doria”, 91, 355–382.

    Dellacasa, M. (1988) Contribution to a world-wide catalogue of Aegialiidae, Aphodiidae, Aulonocnemidae, Termitotrogidae (Coleoptera Scarabaeoidea). Memorie della Societa Entomologica Italiana, 66, 1–455. [Dated 1987].

    Dellacasa, M., Gordon, R.D. & Dellacasa, G. (2002a) Aphodiinae described or recorded by Bates in Biologia Centrali-Americana (Coleoptera Scarabaeoidea: Aphodiidae). Acta Zoologica Mexicana, 86, 155–223.

    Dellacasa, M., Gordon, R.D., Harpootlian, P.J., Stebnicka, Z. & Dellacasa, G. (2002b) Systematic redefinition of the New World Didactyliini (Coleoptera: Scarabaeidae: Aphodiinae) with descriptions of two new species of Aidophus Balthasar. Insecta Mundi, 15(4), 193–216. [Dated 2001].

    Dellacasa, M. & Stebnicka, Z.T. (2001a) Acanthaphodius bruchi A. Schmidt, 1909, femina nondum cognita (Coleoptera: Scarabaeidae: Aphodiinae). Acta Zoologica Cracoviensia, 44(4), 431–433.

    Dellacasa, M. & Stebnicka, Z.T. (2001b) A new genus for Oxyomus morosus Harold, 1869 (Eupariini) and redefinition of Oxyomus setosopunctatus A. Schmidt, 1911 (Aphodiini) (Coleoptera Scarabaeidae: Aphodiinae). Acta Zoologica Mexicana, 83, 29–34.

    Dury, C. (1902) A revised list of the Coleoptera observed near Cincinnati, Ohio, with notes on localities, bibliographical references, and description of new species. Journal of the Cincinnati Society of Natural History, 20, 107–189.

    Erichson, W.F. (1834) Beitrage zur Zoologie: gesammelten auf einer Reise um die Erde, von Dr. F. J. F. Meyen, Insekten : Coleoptera. Nova Acta Leopoldina, 16, 219–284.

    Fabricius, J.C. (1798) Supplementum Entomologiae Systematicae. Proft et Storgh, Copenhagen. 572 pp.

    Fairmaire, L. (1884) Note sur quelques Coléoptères de Magellan et de Santa Cruz. Annales de la Société Entomologique de France, (6)3, 483–506.

    Fairmaire, L. & Germain, P. (1860) Coleoptera chilensia. Revue et Magasin de Zoologie, (2)12, 267–269.

    Faldermann, F. (1835) Fauna entomologica trans-caucasia. Coleoptera: Bd. 1 Coleoptera persico-armeniaca. Nouveaux Mémoires de la Société Impériale des Naturalistes de Moscou, 4, 1–310, 10 plates.

    Fall, H.C. (1930) On Ataenius strigatus Say and allied species (Coleoptera). Journal of the New York Entomological Society, 38, 93–108.

    Fallén, C.F. (1807) Observationes Entomologicae, part 3. Litteris Berlingianis, Lund. Pp. 31–42.

    Gordon, R.D. & Cartwright, O.L. (1988) North American representatives of the tribe Aegialiini (Coleoptera: Scarabaeidae). Smithsonian Contributions to Zoology, 461, 1–37.

    Gordon, R.D. & Pittino, R. (1992) Current status of the American genera and species of Psammodiini (Coleoptera: Scarabaeidae: Aphodiinae). Coleopterists Bulletin, 46, 260–273.

    Gutiérrez, R. (1947) Escarabajos comunes a Chile y la Argentina (Col. Scarabaeidae). Revista de la Sociedad Entomológica Argentina, 13, 309–314.

    Gutiérrez, R. (1949) Notas sobre Scarabaeidae neotrópicos (Coleoptera Lamellicornia). Anales de la Sociedad Científica Argentina, 148(6), 9–35.

    Haldeman, (1848) Descriptions of North American species of Coleoptera, chiefly in the cabinet of J. L. LeConte, M. D., with references to described species. Journal of the Academy of Natural Sciences of Philadelphia, 1, 95–110.

    Harold, E. (1859) Beitrage zur Kenntnis einiger coprophagen Lamellicornien (Erstes Stuck). Berliner Entomologische Zeitschrift, 3, 193–224.

    Harold, E. (1860) Diagnoses de nouvelles espèces d’Aphodius. Annales de la Société Entomologique de France, (3)8, 614–616.

    Harold, E. (1861) Beiträge zur Kenntniss einiger coprophagen Lamellicornia. Berliner Entomologische Zeitschrift, 5, 92–115.

    Harold, E. (1867) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 1, 76–83.

    Harold, E. (1868a) Die chilenischen Aphodiden. Berliner Entomologische Zeitschrift, 11, 278–282. [Dated 1867].

    Harold, E. (1868b) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 3, 80–86.

    Harold, E. (1869) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 5, 94–104.

    Harold, E. (1871) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 8, 114–118.

    Harold, E. (1874) Beitrage zur Kenntniss einiger coprophagen Lamellicornien (Achtes Stuck). Berliner Entomologische Zeitschrift, 18,177–208.

    Heyden, L., Reitter, E. & Weise, J. (1906) Catalogus Coleopterorum Europae, Caucasi et Armeniae Rossicae, Edition 2. Paskau, Berlin. 774 pp.

    Hinton, H.E. (1937) Descriptions of new American Ataenius, with notes on others (Coleoptera, Scarabaeidae). Annals and Magazine of Natural History, series 10, volume 20, 177–196.

    Hope, F.W. (1837) The Coleopterist’s Manual containing the lamellicorn insects of Linnaeus and Fabricius. London. 121 pp.

    Horn, G.H. (1887) A monograph of the Aphodiini inhabiting the United States. Transactions of the American Entomological Society, 14, 1–110.

    Jameson, M.L. & Smith, A.B.T. (2002) Revision of the South American genus Brachysternus Guérin-Méneville (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini: Brachysternina). Coleopterists Bulletin, 56, 321–366.

    Johnson, C. (1975) Nine species of Coleoptera new from Britain. Entomologist’s Monthly Magazine, 111, 177–183.

    Klug, J.C.F. (1845) Coleoptera XV–XVI, 8 unnumbered pages, plate XLI–XLII. In: C. G. Eherenberg (ed.), Symbolae Physicae seu Icones et Descriptiones Insectorum quae ex itinere Africam Borealem et Asiam Occidentalem, volume 5. G. Reimeri, Berlin.

    Kolbe, H.J. (1908) Dynastidae, Cetoniidae und Scarabaeidae. Denkschriften der medizinisch-naturwissenschaftlichen Gesellschaft zu Jena, 13, 121–132.

    Kugelann, J.G. & Illiger, J.K.W. (1798) Verzeichniss der Käfer Preussens. Johann Jacob Gebauer, Halle. 510 pp.

    Landin, B.-O. (1955) Reports of the Lund University Chile Expeditions 1948–49. 22. Coleoptera; Lamellicornia. Lunds Universitets Arsskrift N. F., Avd. 2. Bd. 51. Nr. 14, 1–14.

    Laporte, F.L. (Comte de Castelnau) (1840) Histoire Naturelle des Insectes Coléoptères (Volume 2 of Histoire Naturelle des Animaux Articulés). P. Duménil, Paris. 564 pp.

    Latreille, P.A. (1807) Genera Crustaceorum et insectorum Secundum Ordinem Naturalem in Familias Disposita, Iconibus Exemplisque Plurimis Explicata, volume 2. Amand Koenig, Paris, 280 pp.

    Lea, A.M. (1904) Descriptions of new species of Australian Coleoptera. Proceedings of the Linnean Society of New South Wales, 29, 60–107.

    Lea, A.M. (1923) On Australian aphodiides (Coleoptera). Proceedings of the Royal Society of Victoria, 36(1), 1–20.

    Leach, W.E. (1815) Entomology [pp. 57–172]. In: Brewster (ed.), Edinburgh Encyclopaedia, volume 9, part 1. Edinburgh.

    LeConte, J.L. (1857) Report upon insects collected on the survey. In: Reports of Explorations and Surveys for a Railroad Route from the Mississippi River to the Pacific Ocean, volume 12, part 3, pp. 1–72, 2 plates.

    LePeletier, A.L.M. & Serville, J.G.A. (1828) Scarabé, Scarabaeus (pp. 346–382). In: P. A. Latreille, A. L. M. LePeletier, J. G. A. Serville, and F. E. Guérin-Méneville (eds.) Encyclopédie Méthodique. Histoire Naturelle. Entomologie, ou Histoire Naturelle des Crustaces, des Arachnides et des Insectes, volume 10, part 2. Mme veuve Agasse, Paris, pp. 345–832.

    Linder, H.P. & Crisp, M.D. (1995) Nothofagus and Pacific biogeography. Cladistics, 11, 5–32.

    Linell, M.L. (1896) New species of North American Coleoptera of the family Scarabaeidae. Proceedings of the United States National Museum, 18, 721–731.

    Linnaeus, C. (1758) Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis, edition 10, volume 1. Laurentii Salvii, Stockholm. 823 pp.

    Linnaeus, C. (1767) Systema Naturae, per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis, edition 12, volume 1, part 2. Laurentii Salvii, Stockholm. Pp. 533–1327

    Löbl, I. & Smetana, A. (eds.) (2006) Catalogue of Palaearctic Coleoptera, Volume 3. Scarabaeoidea – Scirtoidea – Dascilloidea – Buprestoidea – Byrrhoidea. Apollo Books, Stenstrup. 690 pp.

    Marseul, S.A. de. (1878) Nouvelles et faits divers de l’Abeille. Mélanges (suite). Deuxième série. Nos. 14 et 15; 16 et 17; 18. L’Abeille, Journal d’Entomologie, 16, 53–60, 63–68, 69–72.

    Martínez, A. (1952) Scarabaeidae nuevos o poco conocidos, III. Publicaciones Mision de Estudios de Patalogia Regional Argentina, Buenos Aires, 23(81–82), 53–118.

    Martínez, A., Periera, F.S. & Vulcano M.A. (1970) Aegialiini, nueva Tribu de Apodiinae para la region neotropical. Acta Zoologica Lilloana, 25, 335–341.

    Melsheimer, F.E. (1846) Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 2, 134–160.

    Ménétries, E. (1832) Catalogue Raisonné des Objets de Zoologie Recueillis dans un Voyage au Caucase et Jusqu’aux Frontières Actuelles de la Perse Entrepris par Ordre de S. M. l’Empereur. L’Académie Impériale des Sciences, St. Petersburg. 271 pp.

    Morrone, J.J. (1996) The biogeographical Andean subregion: a proposal exemplified by Arthropod taxa (Arachnida, Crustacea, and Hexapoda). Neotrópia, 42, 103–114.

    Morrone, J.J. (2001) Review of the biogeographic provinces of the Patagonian subregion. Revista de la Sociedad Entomológica Argentina, 60, 1–8.

    Morrone, J.J., Katinas, L. & Crisci, J.V. (1996) On temperate areas, basal clades and biodiversity conservation. Oryx, 30(3), 187–194.

    Motschulsky, V. (1858) Entomologie spéciale. Insectes des Indes orientales. Etudes Entomologiques, 7, 20–122.

    Motschulsky, V. (1859) Sur les collections coléoptèrologiques de Linné et de Fabricius (continuation). XXII. Lamellicornes. Etudes Entomologiques, 8, 147–162.

    Motschulsky, V. (1862) Synonymie. Etudes Entomologiques, 11, 55.

    Mulsant, E. (1842) Histoire Naturelle des Coléoptères de France. Lamellicornes. Maison, Libraire, Paris, 623 pp.

    Mulsant, E. & Rey, C. (1870) Description d’une espece nouvelle constituant un genre nouveau dans la famille des Aphodiens. Opuscules Entomologiques, 15, 200–202.

    Nakane, T. (1966) Coleoptera of Afghanistan. Results of the Kyoto University Scientific Expedition to the Karakoram and Hindukush, 1955, 8, 231–248.

    Olivier, A.G. (1789) Coléoptères, part 1. Number 3, Scarabé, Scarabaeus, 190 pp. In: Entomologie, ou Histoire Naturelle des Insectes, avec leurs Caractères Génériques et Spécifiques, leur Description, leur Synonymie, et leur Figure Enluminée. Baudouin, Paris.

    d'Orbigny, H. (1896) Synopsis des aphodiens d’Europe et du basin de la Méditerranèe. L’Abeille Journal d’Entomologie, 18, 197–271.

    Panzer, G.W.F. (1796) Scarabaeus caesus. Der Dungkäfer mit der Brustschramme. Faunae Insectorum Germanicae initia oder Deutschlands Insecten, 35, 2.

    Paulian, R. (1934) Deux nouveaux coprophages des Nouvelles-Hébrides. Revue Française d’Entomologie, 1, 219–221.

    Paulian, R. (1937) Trois nouveaux Ataenius de la région australienne. Miscellanea Entomologica, 38, 41–43.

    Paulian, R. (1979) Un nouvel Aphodiidae du Portugal (Col.). Bulletin de la Société Entomologique de France, 84, 66–67.

    Paulsen, M.J. & Mondaca, J. (2005) Revision of the South American Ceratognathini (Coleoptera: Lucanidae: Aesalinae) with the description of a new genus and a new species. Zootaxa, 1191, 1–19.

    Péringuey, L. (1901) Descriptive catalogue of the Coleoptera of South Africa. Fam. Lucanidae. Fam. Scarabaeidae. Sub-families: Coprinae, Aphodiinae, Troginae, Chironinae, Geotrupinae, Hybosorinae, Orphninae, Dynastinae. Transactions of the South African Philosophical Society, 12, 1–560.

    Petrovitz, R. (1963) Neue und verkannte Aphodiinae aus allen Erdteilen (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey, 14, 630–647.

    Petrovitz, R. (1970) Neue neotropische Aphodiinae und Hybosoridae (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 225–243.

    Petrovitz, R. (1971) Neue Aphodiidae der äthiopischen und australischen Region (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 22, 99–104.

    Petrovitz, R. (1973) Neue Aphodiinae, Hybosoridae und Acanthocerinae aus der Neotropischen Region (Coleoptera: Scarabaeidae). Studia Entomologica, 16, 141–202.

    Philippi, R.A. & Philippi, F. (1864) Beschreibung einiger neuen Chilenischen Käfer. Stettiner Entomologische Zeitung, 25, 313–406.

    Pittino, R. & Mariani, G. (1986) Revision of the Old World species of the genus Diastictus Muls) and its allies (Platytomus Muls., Pleurophorus Muls., Afrodiastictus n. gen., Bordatius n. gen.) (Coleoptera, Aphodiidae, Psammodiini). Giorale Italiano di Entomologie, 3(12), 1–165.

    Posadas, P., Miranda-Esquivel, D.R. & Crisci, J.V. (2001) Using phylogenetic diversity measures to set priorities in conservation: an example from southern South America. Conservation Biology, 15(5), 1325–1334.

    Ragusa, E. (1883) Catalogo ragionato dei coleotteri di Sicilia. Palermo. 469 pp.

    Rakovic, M. (1981a) Revision of species of the tribe Psammodiini from the Australian Region (Coleoptera, Scarabaeidae, Aphodiinae). I. Key to genera and revision of the genera Aphodopsammobius Endrodi, Psammodius Fallen, Diastictus Mulsant, Phycochus Broun and Tesarius g. n. Annotationes Zoologicae et Botanicae, 139, 1–37.

    Rakovic, M. (1981b) A revision of the Psammodius Fallen species from Europe, Asia and Africa. Rozpravy Ceske Akademie Ved a Umeni, 91, 1–82.

    Rakovic, M. (1984) A review of the genus Tesarius Rakovic (Coleoptera, Scarabaeidae, Aphodiinae). Acta Entomologica Bohemoslovaca, 81, 448–452.

    Rakovic, M. (1990) Review of the genus Leiopsammodius Rakovic on the world basis with a key to species from the Western Hemisphere and description of a new species (Coleoptera, Scarabaeidae, Aphodiinae). Annotationes Zoologicae et Botanicae, 197, 1–18.

    Reitter, E. (1892) Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen des palaearctischen Faunengebietes. Verhandlungen des Naturforschenden Vereins in Brünn, 24, 1–230.

    Richards, A.M. (1959) Revision of the genus Saprosites Redtenbacher, 1858 (Coleoptera: Aphodiinae) in New Zealand. Transactions of the Royal Society of New Zealand, 87, 35–50.

    Schmidt, A. (1907) Namensanderungen in der Gattung Aphodius und eine Neubescrheibung. Deutsche Entomologische Zeitschrift, 1907, 201–202.

    Schmidt. A. (1909a) Eine Serie neuer Aphodiinen und eine neue Gattung (Fortsetzung VII). Societas Entomologica, 24, 60–62.

    Schmidt, A. (1909b) Eine Serie neuer Apodiinen und eine neue Gattung (Fortsetzung VIII). Societas Entomologica, 24, 66–67.

    Schmidt, A. (1910) Neue Arten aus den Gattungen Aphodius Illig., Ataenius Har., Saprosites Redtenb. (Col.). Deutsche Entomologische Zeitschrift, 1910, 353–361.

    Schmidt, A. (1911) Eine Serie neuer Aphodiinen. Entomologische Zeitung, 72, 3–51.

    Schmidt, A. (1912) Neue Scarabaeiden aus den Subfamilien Aphodiinae, Orphinae und Hybosorinae. Archiv für Naturgeschichte, 78(A), 196–202.

    Schmidt, A. (1913) Erster Versuch einer Einteilung der exotischen Aphodien in Subgenera und als Anhang einige Neubeschreibungen. Archiv für Naturgeschichte, 79(A), 117–178.

    Schmidt, A. (1916) Namenänderungen und Beschreibung neuer Aphodiinen (Col.). Archiv für Naturgeschichte 82(A) Heft 1: 95–116.

    Schmidt, A. (1922) Coleoptera. Aphodiinae. Das Tierreich, 45, I–XXXVI + 1–614 pp.

    Schmidt, A. (1931) Coleoptera-Scarabaeidae-Aphodiinae von Juan Fernandez, p. 638. In: Skottsberg, C. (ed.) The Natural History of Juan Fernandez and Easter Island, volume 3, Zoology. Almquist and Wiksells, Upsalla. 688 pp.

    Skelley, P.E. (2001) New record for Tesarius Rakovic in South America (Scarabaeidae: Aphodiinae: Psammodiini). Insecta Mundi, 14, 240. [Dated 2000].

    Smith, A.B.T. (2002) Revision of the southern South American endemic genus Aulacopalpus Guérin-Méneville with phylogenetic and biogeographic analyses of the subtribe Brachysternina (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini). Coleopterists Bulletin, 56(3), 379–437.

    Solier, A.J.J. (1851) Orden III. Coleopteros, pp. 5–285. In: C. Gay (ed.), Historia Fisica y Politica de Chile. Zoología, vol. 5. C. Gay, Paris. 564 pp.

    Stebnicka, Z.T. (1977) A revision of the world species of the tribe Aegialiini (Coleoptera: Scarabaeidae: Aphodiinae). Acta Zoologica Cracoviensia, 22(11), 397–506.

    Stebnicka, Z.T. (2001) Aphodiinae (Insecta: Coleoptera: Scarabaeidae). Fauna of New Zealand, 42, 1–59.

    Stebnicka, Z.T. (2004) The New World species of Ataenius Harold, 1867. IV. Revision of the A. strigicauda-group (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia, 47(3–4), 211–228.

    Stebnicka, Z.T. (2005) The New World species of Ataenius Harold, 1867. VI. Revision of the A. aequalis-platensis-group (Coleotpera: Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia, 48B(1–2), 99–138.

    Stebnicka, Z.T. (2006) Tribe Eupariini, pp. 143–144. In: I. Löbl and A. Smetana (eds.), Catalogue of Palaearctic Coleoptera, Volume 3. Scarabaeoidea – Scirtoidea – Dascilloidea – Buprestoidea – Byrrhoidea. Apollo Books, Stenstrup. 690 pp.

    Stebnicka, Z.T., Dellacasa, M. & Skelley, P.E. (2004) Review of the New World Aegialiini (Coleoptera: Scarabaeidae: Aphodiinae), with the description of two new genera from South America. Insecta Mundi, 17(1–2), 73–83. [Dated 2003].

    Stebnicka, Z.T. & Howden, H.F. (1994) A revision of the Australian genus Podotenus A. Schmidt (Coleoptera: Scarabaeoidea: Aphodiini). Invertebrate Taxonomy, 8, 17–62.

    Stebnicka, Z.T. & Howden, H.F. (1995) Revision of Australian genera in the tribes Aphodiini, Aegialiini and Proctophanini (Coleoptera: Scarabaeidae: Aphodiinae). Invertebrate Taxonomy, 9, 709–766.

    Stephens, J.F. (1830) Illustrations of British Entomology; or, a Synopsis of Indigenous Insects: Containing their Generic and Specific Distinctions; with an Account of their Metamorphoses, Times of Appearance, Localities, Food, and Economy, as Far as Practicable. Mandibulata, volume 3. Baldwin and Cradock, London. 379 pp.

    Sturm, J. (1805) Deutschlands Fauna in Abbildungen nach der Natur mit Beschreibungen. Nurnberg. 268 pp.

    Waltl, J. (1835) Reise durch Tyrol, Oberitalien und Piemont nach dem sudlichen Spanien. Passau. 247 pp.

    Weinelt, M. (2006) Online Map Creation. Version 4.1. Available from: http://www.aquarius.geomar.de/omc/omc_intro.html (Accessed: 28 August 2006).