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Published: 2007-04-23
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Scarabaeoidea (Coleoptera) of Portugal: genus-group names and  their type species

Rua de Camtes, 788, 2 Dto., P-4000-142 Porto, PORTUGAL
Coleoptera Scarabaeoidea Portugal nomenclature genus-group names type species

Abstract

The type species and nomenclature are discussed in detail of the genus-group names that have been used, correctly or incorrectly, in combination with species recorded from Portugal. This work strictly adheres to the rules of the International Code of Zoological Nomenclature, in order to promote nomenclatural stability. The contents are strictly nomenclatural as no view is taken on the taxonomic validity or rank of the genus-group names. A total of 171 available names are examined. Evidence is provided in each case for the reasons why the stated nominal species is believed to be the validly designated type species. Many instances were found in the modern literature of type species statements not in compliance with the requirements of the Code. In most cases it is a senior synonym that is stated as type species, instead of the nominal species originally included when the genus was established. That, fortunately, does not cause nomenclatural instability but should be corrected. In three cases where nomenclatural stability is threatened, Anisoplia Schönherr, 1817, Phyllopertha Stephens, 1830, and Scarabaeus Linnaeus, 1758, it is suggested that prevailing usage should be maintained until a case is made to the International Commission on Zoological Nomenclature and a ruling is published.

References

  1. ;dám, L. (1994) A checklist of the Hungarian Scarabaeoidea with the description of ten new taxa (Coleoptera). Folia Entomologica Hungarica, 55, 5–17.

    ;dám, L. (2003) Faunisztikai adatok a Kárpát-medencébÝl (Coleoptera: Scarabaeoidea). Folia Historico Naturalia Musei Matraensis, 27, 101–136.

    Agassiz, L. (1846) Nomenclatoris zoologici index universalis, continens nomina systematica classium, ordinum, familiarum et generum animalium omnium, tam viventium quam fossilium, secundum ordinem alphabeticum unicum disposita, adjectis homonymiis plantarum, nec non variis adnotationibus et emendationibus. Jent et Gassmann, Solothurn, viii + 393 pp.

    Allsopp, P.G. (1982) Hybosorus illigeri Reiche, 1853 (Insecta, Coleoptera): Proposed conservation by use of the plenary powers. Z.N.(S.) 2296. The Bulletin of Zoological Nomenclature, 39, 218–219.

    Arrow, G.J. (1910) The Fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia (Cetoniinae and Dynastinae). Taylor and Francis, London, xiv + 322 pp, 2 pls.

    Arrow, G.J. (1917) The Fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part II (Rutelinae, Desmonycinae and Euchirinae). Taylor and Francis, London, xiii + 387 pp, 5 pls.

    Arrow, G.J. (1931) The Fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part III (Coprinae). Taylor and Francis, London, xii + 428 pp, 13 pls, 1 map.

    Balthasar, V. (1959) Beitrag zur Kenntnis der Gattung Onthophagus Latr. Acta Entomologica Musei Nationalis Pragae, 33, 461–471.

    Balthasar, V. (1961) Eine neue Gattung und neue Arten der Unterfamilie Aphodiinae (104. Beitrag zur Kenntnis der Scarabaeidae, Coleoptera). Deutsche Entomologische Zeitschrift, Neue Folge, 8, 121–130.

    Balthasar, V. (1964) Monographie der Scarabeidae und Aphodiidae der palaearcktischen und orientalischen Region (Coleoptera: Lamellicornia). Band 3. Aphodiidae. Tschechoslowakischen Akademie der Wissenschaften, Prague, 652 pp, 2 pls.

    Baraud, J. (1962) Révision des espèces paléarctiques du genre Triodonta Muls. Actes de la Socté Linnnnée de Bordeaux, 100, 5–84.

    Baraud, J. (1965) Le genre Chasmatopterus Latr. (Col. Scarabaeoidea). Eos, Revista Española de Entomologia, 40, 263–287.

    Baraud, J. (1986) Nouvelle classification proposée pour les espèces du genre Anisoplia Fischer, 1824 (Col. Scarabaeoidea, Rutelidae). Première partie. L’Entomologiste, 42, 322–344.

    Baraud, J. (1987) Révision des Elaphocera d’Europe (Coléoptères, Melolonthidae). Annales de la Société Entomologique de France (Nouvelle Série), 23, 125–134.

    Baraud, J. (1991) Révision des espèces du genre Anisoplia Fischer, 1824 (Coleoptera Scarabaeoidea Rutelidae) (deuxième partie). Bulletin Mensuel de la Société Linnéenne de Lyon, 60, 309–344, 353–384.

    Baraud, J. (1992) Coléoptères Scarabaeoidea d’Europe. Faune de France et régions limitrophes. 78. Fédération FranHaise des Sociétés de Sciences Naturelles & Société Linnéenne de Lyon, Lyon, ix + 856 pp, xi pls.

    Bedel, L. (1892) Révision des Scarabaeus paléarctiques. L’Abeille, Journal d’Entomologie, 27, 281–288.

    Bedel, L. (1903) Synonymies de Coléoptères paléarctiques. L’Abeille, Journal d’Entomologie, 30, 152.

    Bedel, L. (1907) Synonymies de divers Aphodiens paléarctiques. L’Abeille, Journal d’Entomologie, 31, 56.

    Bedel, L. (1911) Synonymies de Scarabaeidae paléarctiques (Col.). Bulletin de la Société Entomologique de France, 1911, 377–381.

    Bezd.k, A. (2006) Tribe Melolonthini Samouelle, 1819. In: Löbl, I. & Smetana, A. (Ed.), Catalogue of Palaearctic Coleoptera. Volume 3: Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books, Stenstrup, 191–198.

    Blanchard, C.E. (1845) Histoirre Naturelle des Insectes, traitant de leurs moeurs et de leurs métamorphoses en général, et comprenant une nouvelle classification fondée sur leurs rapports naturels. Tome II. Coléoptères, Orthoptères, Névroptères, Lépidoptères, Hémiptères, Aphaniptères, Strépsiptères, Diptères, Anoplures et Thysanures. Firmin Didot Frères, Paris, 524 pp, pls. 11–20.

    Blanchard, C.E. (1851) Ordre des Coléoptères. In: Milne-Edwards, H., Blanchard, E. & Lucas, H., Muséum d’Histoire Naturelle de Paris: Catalogue de la Collection Entomologique. Classe des Insectes. Tome 1, Deuxième livraison. Gide & Baudry, Paris, 129–240.

    Boucomont, A. (1905)Étude sur les Enoplotrupes et Geotrupes d’Asie. Revue d’Entomologie, 23[1904], 209–252.

    Boucomont, A. (1911) Contribution à la classification des Geotrypidae (Col.). Annales de la Société Entomologique de France, 79[1910], 333–350.

    Boucomont, A. (1912) Scarabaeidae: Taurocerastinae, Geotrupinae. Pars 46. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Volume 19. W. Junk, Berlin, 47 pp.

    Branco, T. (2001) Coleoptera Scarabaeoidea new or otherwise noteworthy for the Portuguese fauna, with a nomenclatural note. Boletín de la Sociedad Entomologica Aragonesa, 29, 33–38.

    Branco, T. (2006) Authorship and date of the genus-group names Rhizotrogus and Amphimallon (Coleoptera: Melolonthidae). Elytron, 19[2005], 43–48.

    Branco, T. & Ruiz, J.L. (2003) Triodontella Reitter, 1919 or Triodonta Mulsant, 1842: a clarification (Coleoptera, Melolonthidae). Boletín de la Sociedad Entomologica Aragonesa, 33, 278.

    Branco, T. & Ziani, S. (2005) Cheironitis Lansberge, 1875 – its correct spelling and validity (Coleoptera, Scarabaeidae). Boletín de la Sociedad Entomologica Aragonesa, 37, 267–272.

    Branco, T. & Ziani, S. (2006) New acts and comments: Geotrupidae. In: Löbl, I. & Smetana, A. (Ed.), Catalogue of Palaearctic Coleoptera. Volume 3: Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books, Stenstrup, 28–30.

    Burmeister, H.C.C. (1842) Handbuch der Entomologie. Dritter Band. Coleoptera Lamellicornia Melitophila. T.C.F. Enslin, Berlin, xxii + 827 pp.

    Cartwright, O.L. (1953) Scarabeid beetles of the genus Bradycinetulus and closely related genera in the United States. Proceedings of the United States National Museum, 103, 95–120, pls. 3–4.

    Cartwright, O.L. (1974) Ataenius, Aphotaenius, and Pseudataenius of the United States and Canada. Smithsonian Contributions to Zoology, 154, 106 pp, 3 pls.

    Chapin, E.A. (1946) Necessary changes of names in the coleopterous family Scarabaeidae. Proceedings of the Biological Society of Washington, 59, 79–80.

    Clément, P. (1962) Un Anomius nouveau du Maroc (Col. Scarab. Aphodiini). Bulletin de la Société des Sciences Naturelles du Maroc, 42, 41–45.

    Costa, A. (1852) Coleotteri Pentameri. Famiglia de’ Cetoniidei. In: Costa, O.-G. (Ed.), Fauna del Regno di Napoli, ossia Enumerazione di tutti gli Animale che abitano le diverse regioni de questo Regno e le acque che le bagnano, contenente la descrizione de’ nuovi o poco esattamente conosciuti, con figure ricavate da originali viventi e dipinte al naturale. Gaetano Sautto, Napoli, 1–32, pl. 13. [Families separately paginated]

    Creutzer, C. (1799) Entomologische Versuche. K. Schaumburg und Comp., Vienna, 142 pp, 3pls.

    Crotch, G.R. (1870) The genera of Coleoptera studied chronologically (1802–21). Transactions of the Entomological Society of London, 1870, 213–241.

    Curtis, J. (1829) British Entomology; being illustrations and descriptions of the genera of insects found in Great Britain and Ireland: containing coloured figures from nature of the most rare and beautiful species, and in many instances of the plants upon which they are found. Volume 6. Published by the author, London, sheets and pls. 242–289.

    Curtis, J. (1834) British Entomology; being illustrations and descriptions of the genera of insects found in Great Britain and Ireland: containing coloured figures from nature of the most rare and beautiful species, and in many instances of the plants upon which they are found. Volume 11. Published by the author, London, sheets and pls. 482–529.

    Cuvier, G. (1797) Tableau élémentaire de l’Histoire Naturelle des Animaux. Baudoin, Paris, xvi + 710 pp, 14 pls.

    Dalla Torre, K. W. von (1912a) Scarabaeidae: Melolonthinae I. Pars 45. In: S. Schenkling (Ed.), Coleopterorum Catalogus. Volume 20. W. Junk, Berlin, 1–84.

    Dalla Torre, K.W. von (1912b) Scarabaeidae: Melolonthinae III. Pars 49. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Volume 20. W. Junk, Berlin, 134–290.

    Dechambre, R.-P. (2002) Du bon usage du principe de priorité ou pourquoi Gnorimus doit rester Gnorimus (Coleoptera, Cetoniidae, Trichiinae). L’Entomologiste, 57[2001], 229–232.

    Dejean, P.F.M.A. (1821) Catalogue de la collection de Coléoptères de M. le Baron Dejean. Crevot Libraire, Paris, viii + 136 + 2 pp.

    Dejean, P.F.M.A. (1833) Catalogue des Coléoptères de la collection de M. le Comte Dejean. Deuxième Édition,

    Deuxième livraison. Méquignon-Marvis Père et Fils, Paris, 97–176.

    Dellacasa, G. (1983) Sistematica e nomenclatura degli Aphodiini italiani (Coleoptera Scarabaeidae: Aphodiinae). Monografie I, Museo Regionale di Scienze Naturali, Torino, 465 pp.

    Dellacasa, G. & Baraud, J. (1978) Heptaulaculus nov. gen. per gli Heptaulacus paleartici del gruppo testudinarius (Fabr.). Bollettino della Società Entomologica Italiana, 110, 62–68.

    Dellacasa, G., Bordat, P. & Dellacasa, M. (2001) A revisional essay of world genus-group taxa of Aphodiinae (Coleoptera Aphodiidae). Memorie della Società Entomologica Italiana, 79, 1–482.

    Dellacasa, M. & Dellacasa, G. (2005) Comments on some systematic and nomenclatural questions in Aphodiinae with descriptions of new genera and on Italian taxa (Coleoptera Aphodiidae). Memorie della Società Entomologica Italiana, 84, 45–101.

    Dellacasa, M. & Dellacasa, G. (2006) Tribe Aphodiini Leach, 1815. In: Löbl, I. & Smetana, A. (Ed.), Catalogue of Palaearctic Coleoptera. Volume 3: Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books, Stenstrup, 105–143.

    Didier R. & Séguy, E. (1953) Catalogue illustré des Lucanides du Globe. Texte. Encyclopédie Entomologique, 27, 1–223.

    Erichson, W. F. (1841) Ueber die Insecten von Algier mit besonderer Bercksichtitung ihrer geographischen Verbreitung. In: Wagner, M. F. (Ed.), Reisen in der Regentschaft Algier in den Jahren 1836, 1837 und 1838 nebst einem naturhistorischen Anhang und einem Kupferatlas. Dritter Band. L. Voss, Leipzig, 140–194.

    Erichson, W.F. (1847) Naturgeschichte der Insecten Deutschlands. Erste Abteilung. Coleoptera. Dritter Band. Lieferung 5. Nicolaische Verlags-Buchhandlung, Berlin, 641–800.

    Eschscholtz, J.F. (1830) Nova genera Coleopterorum Faunae Europaeae. Bulletin de la Société Impériale des Naturalistes de Moscou, 2, 65–66.

    Evenhuis, N. L. (1997) Litteratura Taxonomica Dipterorum (17581930). Volumes III. Backhuys Publishers, Leiden, 871 pp.

    Fabricius, J.C. (1775) Systema Entomologicae, sistens Insectorum Classes, Ordines, Genera, Species, adjectis Synonymis, Locis, Descriptionibus, Observationibus. Kortii, Flensburg and Leipzig, xxxii + 832 pp.

    Fabricius, J.C. (1792) Entomologia systematica emendata et aucta. Secundum Classes, Ordines, Genera, Species adjectis, Synonymis, Locis, Observationibus, Descriptionibus. Tome I, Part 1. C. G. Proft, Copenhagen, xx + 330 pp.

    Fabricius, J.C. (1798) Supplementum Entomologiae systematicae. Proft et Storch, Copenhagen, 572 pp.

    Fabricius, J.C. (1801) Systema Eleutheratorum secundum Ordines, Genera, Species: adjectis Synonymis, Locis, Observationibus, Descriptionibus. Tomus I. Bibliopolii Academici Novi, Kiel, xxiv + 506 pp.

    Fallén, C.F. (1807) Observationes entomologicae. Pars III. Berlingianis, Lund, 31–42.

    Fischer von Waldheim, G. (1824) Entomographie de la Russie. Tome II. A.Semen, Moscow, xx + 264 pp, 40 pls.

    Fuente, J.M. de la (1926) Catálogo sistemático-geográfico de los Coleópteros observados en la península ibérica, Pirineos propriamente dichos y Baleares (continuación). Boletín de la Sociedad Entomològica de España, 9, 158–173.

    Gené, J. (1836) De quibusdam Insectis Sardiniae novis aut minus cognitis. Fasciculus I. Memorie della Accademia Realle delle Scienze di Torino, Classe di Scienze Naturali e Fisiche, 39, 1–199, 1 pl.

    Geoffroy, E.L. (1762) Histoire abrégée des insectes qui se trouvent aux environs de Paris. Tome premier. Durand, Paris, xxviii + 523 pp, 10 pls.

    Gozis, M. des (1886) Recherche de l’espèce typique de quelques anciens genres. Rectifications synonymiques et notes diverses. Herbin, MontluHon, 36 pp.

    Griffin, F.J. (1936) On the Dates of Publication of Motschulsky (V. de), ‘Etudes entomologiques’, I. – XI., 1853 – 1862. The Annals and Magazine of Natural History, Tenth Series, 12, 256–257.

    Guérin-Méneville, F.E. (1838a) Insectes du voyage de la Favorite. Revue et Magasin de Zoologie, Journal, 8, 1–80.

    Guérin-Méneville, F.E. (1838b) Crustacés, arachnides et insectes. In: Duperrey, L.E. (Ed.), Voyage Autour du Monde, exécuté par ordre du Roi, sur la Corvette de Sa Majesté, la Coquille, pendant les années 1822, 1823, 1824 et 1825, sous le Ministre et conformément aux instructions de S.E.M. Le Marquis de Clermont-Tonerre, Ministre de la Marine; et Publié sous les Auspices de son Excellence Mgr le Cte de Chabrol, Ministre de la Marine et des Colonies, vol. 2, Part 2 (Zoologie, M.Lesson editor). Division 1 (M.L.I. Duperrey, editor). Arthus Bertrand, Paris, xii + 319 pp. [dated 1830]

    Gyllenhal, L. (1808) Insecta Suecica. Classis I. Coleoptera sive Eleuterata. Tomus I. F.J. Leverentz, Scaris, viii + 4 unpaginate + 572 pp.

    Harold, E. von (1867a) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 1, 76–83.

    Harold, E. von (1867b) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 2, 94–100.

    Harold, E. von (1873) Literatur - Histoire naturelle des Coléoptères de France, par E. Mulsant, Lamellicornes – Pectinicornes. 1871. (2 Edit.). Coleopterologische Hefte, 11, 114–121.

    Heer, O. (1841) Fauna Coleopterorum Helvetica. Pars 1. Orelii, Fuesslini et Sociorum, Zrich, xii + 652 pp.

    Heyden, L. v., Reitter, E. & Weise, J. (1883) Catalogus Coleopterorum Europae et Caucasi. Editio Tertia. Libraria Nicolai, Berlin, 228 pp.

    Hollande, A. & Thérond, J. (1998) Aphodiidae du Nord de l’Afrique (Coleoptera Scarabaeoidea) (au soin de G.Dellacasa). Monografie XXI, Museo Regionale di Scienze Naturali, Torino, 280 pp.

    Hope, F.W. (1837) The Coleopterist’s Manual, containing the Lamellicorn Insects of Linnaeus and Fabricius. H.G.Bohn, London, xiii + 121 pp, 4 pls.

    Illiger, J.K.W. (1798) Verzeichniss der Käfer Preussens. J. J. Gebauer, Halle, xlii + 510 pp.

    Illiger, J.K.W. (1803) Verzeichniss der in Portugal einheimischen Käfer. Erste Lieferung. Magazin für Insektenkunde, 2, 186–258.

    International Commission on Zoological Nomenclature (1910) Opinion 11. The Designation of Genotypes by Latreille, 1810. Smithsonian Publication nr. 1938, 17–18.

    International Commission on Zoological Nomenclature (1922) Opinion 71. Interpretation of the Expression Typical Species in Westwood’s (1840) Synopsis. Smithsonian Miscellaneous Collections, 73, 200–202.

    International Commission on Zoological Nomenclature (1939) Opinion 136. Opinion supplementary to Opinion 11 on the interpretation of Latreille’s Considérations générales sur l’ordre naturel des animaux composant les classes des crustacés, des arachnides et des insectes avec un tableau méthodique de leurs genres disposés en familles, Paris, 1810. Opinions and Declarations Rendered by the International Commission on Zoological Nomenclature, 2, 15–19.

    International Commission on Zoological Nomenclature (1954) Direction 4. Addition to the Official Lists and Official Indexes of certain scientific names and of the titles of certain books dealt with in Opinions 134–160, exclusive of Opinion 140. Opinions and Declarations Rendered by the International Commission on Zoological Nomenclature, 2, 631–651.

    International Commission on Zoological Nomenclature (1955) Opinion 346. Designation, under the plenary powers, for the genus Geotrupes Latreille, 1796 (Class Insecta, Order Coleoptera) of a type species in harmony with current usage. Opinions and Declarations Rendered by the International Commission on Zoological Nomenclature, 10, 391–408.

    International Commission on Zoological Nomenclature (1956) Direction 32. Addition to the Official List of Works approved as available for Zoological Nomenclature of the Titles of certain Works dealt with in Opinions rendered in the period up to the end of 1936 and to the Official Index of Rejected and Invalid Works in Zoological Nomenclature of certain Works similarly dealt with in the same period ans matters incidental thereto. Opinions and Declarations Rendered by the International Commission on Zoological Nomenclature, 1C, 307–328.

    International Commission on Zoological Nomenclature (1956) Direction 46. Determination of the gender attributable to the generic names Acheta Linnaeus, 1758 (Class Insecta, Order Orthoptera) and Geotrupes (Class Insecta, Order Coleoptera) (Variation of rulings given in Opinions 299 and 346 respectively). Opinions and Declarations Rendered by the International Commission on Zoological Nomenclature, 12, 435–440.

    International Commission on Zoological Nomenclature (1957a) Direction 63. Completion and in certain cases correction of entries relating to the names of genera belonging to the class Insecta made in the Official List of Generic Names in Zoology in the period up to the end of 1936. Opinions and Declarations Rendered by the International Commission on Zoological Nomenclature, 1E, 21–60.

    International Commission on Zoological Nomenclature (1957b) Opinion 474. Determination of the dates to be assigned for the purposes of the Law of Priority to the names published in Dru Drury’s Illustrations of Natural History in the period 1770–1782. Opinions and Declarations Rendered by the International Commission on Zoological Nomenclature, 16, 297–306.

    International Commission on Zoological Nomenclature (1958) Opinion 516. Determination under the Plenary Powers of the relative precedence to be assigned to certain works on the Order Lepidoptera (Class Insecta) published in 1775 by Pieter Cramer, Michael Denis & Ignaz Schiffermüller, Johann Christian Fabricius, Johann Caspar Fuessly, and S.A. von Rottemburg respectively. The Bulletin of Zoological Nomenclature, 19, 1–44.

    International Commission on Zoological Nomenclature (1989) Opinion 1546. Chelonus Panzer, 1806 (Insecta, Hymenoptera) and Anomala Samouelle, 1819 (Insecta, Coleoptera): names conserved. The Bulletin of Zoological Nomenclature, 46, 149–150.

    International Commission on Zoological Nomenclature (1993) Opinion 1710. J.C. Megerle’s (1801–1805) auction catalogues of insects: suppressed for nomenclatural purposes, with the specific names of Aperda alboguttata Megerle, 1803 (currently Apomecyna alboguttata; Coleoptera) and Hippobosca variegata Megerle, 1803 (Diptera) conserved. The Bulletin of Zoological Nomenclature, 50, 79–82.

    International Commission on Zoological Nomenclature (1994) Opinion 1754. Histoire abrégée des insectes qui se trouvent aux environs de Paris (Geoffroy, 1762): some generic names conserved (Crustacea, Insecta). The Bulletin of Zoological Nomenclature, 51, 58–70.

    International Commission on Zoological Nomenclature (1999) International code of zoological nomenclature, Code international de nomenclature zoologique. Fourth Edition. International Trust for Zoological Nomenclature, London, xxix + 306 pp.

    International Commission on Zoological Nomenclature (2003) Opinion 2054 (Case 3201). Scarabaeus punctatus Villers, 1789 (currently Pentodon bidens punctatus; Insecta, Coloeptera); specific name conserved. The Bulletin of Zoological Nomenclature, 60, 247–248.

    International Commission on Zoological Nomenclature (2004), Opinion 2079 (Case 2926). Trichia Hartmann, 1840 (Mollusca, Gastropoda): proposed conservation; and TRICHIINAE LoÒek, 1956 (Gastropoda): proposed emendation of spelling to TRICHIAINAE, so removing homonymy with TRICHIIDAE Fleming, 1821 (Insecta, Coleoptera) not approved. The Bulletin of Zoological Nomenclature, 61, 177–181.

    International Commission on Zoological Nomenclature (2005a) Proposed conservation of the specific name of Nicrophorus tomentosus Weber, 1801 (Insecta, Coleoptera): case closed. The Bulletin of Zoological Nomenclature, 62, 124.

    International Commission on Zoological Nomenclature (2005b) New applications to the Commission. CASE 3349: Gnorimus Lepeletier de Saint-Fargeau & Serville, 1828, and Osmoderma Lepeletier de Saint-Fargeau & Serville, 1828 (Insecta, Coleoptera): proposed conservation. F.-T.Krell, A.Ballerio, A.B.T.Smith & P.Audisio. The Bulletin of Zoological Nomenclature, 62, 126.

    International Commission on Zoological Nomenclature (2006a) Opinion 2138 (Case 3097). Bolboceras Kirby, 1819 (July) (Insecta, Coleoptera): not conserved; priority maintained for Odonteus Samouelle, 1819 (June). The Bulletin of Zoological Nomenclature, 63, 62–64.

    International Commission on Zoological Nomenclature (2006b) OPINION 2150 (Case 3317), Scarabaeus arenarius Olivier, 1789 (currently Aphodius arenarius) and Scarabaeus fasciatus Olivier, 1789 (currently Aphodius fasciatus) (Insecta, Coleoptera): specific names conserved. Bulletin of Zoological Nomenclature, 63, 140–141.

    Janssens, A. (1937) Révision des Onitides. Mémoires du Musée Royal d’Histoire Naturelle de Belgique, Deuxième série, 11, 200 pp, 2 pls.

    Janssens, A. (1953) Exploration du Parc National de l’Upemba. Mission G. F. de Witte en colaboration avec W. Adam, A. Janssens, L. van Meel, et R. Verheyen (1946–1949). Fascicule 11. Oniticellini (Coleoptera Lamellicornia). Institut des Parcs Nationaux du Congo Belge, Bruxelles, 118 pp.

    Jekel, H. (1866) Essai sur la classification naturelle des Geotrupes Latreille et description d’espèces nouvelles. Annales de la Société Entomologique de France, 4e série, 5[1865], 513–618.

    Jessop, L. (1986) Dung Beetles and Chaffers. Coleoptera: Scarabaeoidea. New edition. Handbooks for the Identification of British Insects. Royal Entomological Society, London, 5, 53pp.

    Kabakov, O.N. (1979) Obzor plastinchatousyhk zhukov podsemeystva Coprinae (Scarabaeidae, Coleoptera) Dalnego Vostoka SSSR i Sopredelnykh territori. In: Krivolutskaya, G.O. (Ed.), Zhuki Dalnego Vostoka i vostochnoi Sibiri. Akademiya Nauk SSSR, Vladivostok, 58–98.

    Kabakov, O. N. (1980) Reviziya zhukov roda Scarabaeus L. (Coleoptera, Scarabaeidae) fauny SSSR. Entomologicheskoe Obozrenie, 59, 819–829, 2 pls.

    Kirby, W. (1819) A description of several new Species of Insects collected in New Holland by Robert Brown, Esq. - F.R.S.Lib.Linn.Soc. Transactions of the Linnean Society of London, 12[1818]: 454–482, pls. 21–23.

    Kirby, W. (1825a) A description of some Insects which appear to exemplify Mr. William S. MacLeay’s Doctrine of Affinity and Analogy. Transactions of the Linnean Society of London, 14, 93–110, pl. 3.

    Kirby, W. (1825b) A Description of such Genera and Species of Insects, alluded to in the Introduction to Entomology of Messrs. Kirby and Spencer, as appear not to have been before sufficiently noticed or described. Transactions of the Linnean Society of London, 14, 563–572.

    Kirby, W. (1827) A description of some new Genera and Species of Petalocerous Coleoptera. The Zoological Journal, 3, 145–158.

    Koshantschikoff, W.D. (1912) Vierter Beitrag zur Kenntnis der Aphodiini (Coleoptera Lamellicornia). Russkoe Entomologicheskoe Obozrenie, 12, 511–523.

    Krell, F.-T. (1990) Nomenklaturische Bemerkung zu diversen Taxa der Scarabaeoidea orb. terr. (Coleoptera). Entomologische Blätter, 86, 103–114.

    Krell, F.-T. (1991) Vorschlag zur Stabilisierung der Hoplia-Nomenklatur (Coleoptera: Scarabaeoidea: Hoplinae). Entomolomologische Blätter, 87, 186–192.

    Krell, F.-T. (1996) Type Catalogue of some Scarabaeoidea species described by Linnaeus (Insecta: Coleoptera). The Linnean, Newsletter and Proceedings of the Linnean Society of London, 11, 13–20.

    Krell, F.-T. (2002) On Nomenclature and Synonymy of Old World Dynastinae (Coleoptera: Scarabaeidae). Entomologische Blätter für Biologie und Systematik der Käfer, 98, 37–45.

    Krell, F.Th. (2006) New nomenclatural an taxonomic acts, and comments: Scarabaeidae: Dynastinae. In: Löbl, I. & A.Smetana (Ed.), Catalogue of Palaearctic Coleoptera. Volume 3: Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books, Stenstrup, 37.

    Krell, F.T., Ballerio, A., Smith, A.B.T & Audisio P. (2006) Case 3349. Gnorimus Le Peletier de Saint-Fargeau & Serville, 1828 and Osmoderma Le Peletier de Saint-Fargeau & Serville, 1828 (Insecta, Coleoptera): proposed conservation of the generic names. The Bulletin of Zoological Nomenclature, 63, 177–183.

    Kuijten, P.J. (1983) Revision of the genus Hybosorus MacLeay (Coleoptera: Scarabaeidae, Hybosorinae). Zoologische Verhandelingen, 203, 49 pp, 1 pl.

    Landin, B.-O. (1956) The Fabrician species of Aphodiini and Aegialiini (Col. Lamellicornia). Opuscula Entomologica, 21, 203–228.

    Landin, B.-O. (1960) The Lamellicorn Beetles of the Azores (Coleoptera), with some reflexions on the classifications of certain Aphodiini. Report N 5 from the Lund University Expedition in 1957 to the Azores and Madeira. Boletim do Museu Municipal do Funchal, 13, 49–84.

    Landin, B.-O. (1964) The identity of Scarabaeus arator Fabricius, 1775 (Col. Lamellicornia), with the designation of neotypes of arator Fabr. and arator Illig. and a list of the insects from the Cape Colony described by Fabricius 1775. Opuscula Entomologica, 29, 117–142.

    Lansberge, J.W. van (1875) Monographie des Onitides. Annales de la Société Entomologique de Belgique, 18, 1–148.

    Laporte, F.L. (1832a) Mémoire sur cinquante espèces nouvelles ou peu connues d’insectes. Annales de la Société Entomologique de France, 1, 386–415.

    Laporte, F.L. (1832b) Classe IX; Insectes: Calicnemis. Magasin de Zoologie, 1, pl. 7.

    Laporte, F.L. (1840) Histoire Naturelle des Insectes Coléoptères. Tome deuxième. P. Duménil, Paris, 564 pp, 38 pls. [Signed Comte de Castelnau]

    Latreille, P.A. (1797) Précis des caractères génériques des Insectes, disposés dans un ordre naturel. F.Bordeaux, Brive, xiii + 209 pp, 5 pls.

    Latreille, P.A, (1802) Histoire Naturelle, Générale et Particulière des Crustacés et des Insectes. Tome troisième. F.Dufart, Paris, xii + 464 pp.

    Latreille, P.A. (1807) Genera Crustaceorum et Insectorum secundum ordinen naturalem in familias disposita, iconibus exemplisque plurimus explicata. Tomus secundus. Amand Koenig, Paris, 280 pp.

    Latreille, P.A. (1810) Considérations Générales sur l’Ordre Naturel des Animaux Composant les Classes des Crustacés, des Arachnides, et des Insectes; avec un Tableau Méthodique de leurs Genres, disposés en Familles. F. Schoell, Paris, 444 pp.

    Latreille, P.A. (1825) Familles naturelles du règne animal, exposées succinctement et dans un ordre analytique, avec l’indication de leurs genres. J.-B. Baillière, Paris, 570 pp.

    Leach, W. E. (1815) Entomology. In: Brewster, D. (Ed.), Edinburgh Encyclopaedia. Volume 9, first part. Balfour, Edinburgh, 57–172.

    LePeletier de Saint-Fargeau, A.L.M. & Audinet-Serville, J.G. (1828a) Scarabé. In: Latreille, P.A., LePeletier de Saint-Fargeau, A.L.M., Audinet-Serville, J.G. & Guérin-Méneville, F.E., Encyclopédie Méthodique. Histoire Naturelle. Entomologie, ou Histoire Naturelle des Crustacés, des Arachnides et des Insectes. Tome dixième, deuxième livraison. Mme. Veuve Agasse, Paris, 346–382.

    LePeletier de Saint-Fargeau, A.L.M. & Audinet-Serville, J.G. (1828b) Trichie. In: Latreille, P.A., LePeletier de Saint-Fargeau, A.L.M., Audinet-Serville, J.G. & Guérin-Méneville, F.E., Encyclopédie Méthodique. Histoire Naturelle. Entomologie, ou Histoire Naturelle des Crustacés, des Arachnides et des Insectes. Tome dixième, deuxième livraison. Mme. Veuve Agasse, Paris, 701–704.

    Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Decima, Reformata. Tomus I. Laurentii Salvii, Stockholm, 823 pp.

    Linnaeus, C. (1761) Fauna Svecica sistens Animalia Sveciae Regni: Mammalia, Aves, Amphibia, Pisces, Insecta, Vermes. Distributa per classes & ordines, genera & species, cum Differentiis Specierum, Synonymis Auctorum, Nominibus Incolarum, Locis Natalium, Descriptionibus Insectorum. Editio Altera, Auctior. Laurentii Salvii, Stockholm, 587 pp.

    Linnaeus, C. (1764) Museum S:ae R:ae M:tis Ludovicae Ulricae Reginae Svecorum, Gothorum, Vandalorumque &c. &c. &c. in quo animalia rariora, exotica, Imprimis Insecta & Conchilia describuntur et determinantur. Laurentii Salvii, Stockholm, 7 (unpaged) + 720 + 2 (unpaged) pp.

    Linnaeus, C. (1767) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Duodecima, Reformata. Tomus I. Pars 2. Laurentii Salvii, Stockholm, 533–1327 + 36 (unpaged).

    Löbl, I., Nikolajev, G.V. & Král, D. (2006) Subfamily Geotrupinae Latreille, 1802. In: Löbl, I. & Smetana, A. (Ed.), Catalogue of Palaearctic Coleoptera. Volume 3: Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books, Stenstrup, 84–92.

    Lopéz-Colón, J.I. (1989) Algunas considerationes sobre la morfología de la armadura genital masculina en el género Thorectes Mulsant, 1842 y sus implicaciones filogenéticas (Col. Scarabaeoidea, Geotrupidae). Boletín do Grupo Entomològico de Madrid, 4, 69–82.

    Machatschke, J.W. (1957) Coleoptera Lamellicornia, Fam. Scarabaeidae, subfam. Rutelinae. Zweiter Teil. In Wytsman, P. (Ed.), Genera Insectorum, Fascicule 199B. Crainhem, Quatre-Bras. 219 pp, 6 pls.

    MacLeay, W.S. (1819) Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part I. S.Bagster, London, xxx + 160 pp.

    MacLeay, W.S. (1821) Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part II. S.Bagster, London, 161–524.

    Martín-Piera, F. (2000) Familia Scarabaeidae. In: Martín-Piera, F. & Lòpez-Colòn, J.I., Coleoptera, Scarabaeoidea I. Fauna Ibérica, Volume 14. Ramos, M.A. et al. (Ed.). Museo Nacional de Ciencias Naturales, CSIC, Madrid, 205–432.

    Martín Piera, F. & Zunino, M. (1983) Amphionthophagus, nuovo sottogenere di Onthophagus Latr. (Coleoptera, Scarabaeoidea). Bollettino del Museo Regionale di Scienze naturali, 1, 59–76.

    Medvedev, S.I. (1949) Fauna SSSR, Zhestkokrylye. Tom X, vyp. 3. Plastinchatousye (Scarabaeidae), Podsem. Rutelinae (khlebnye zhuki i blizkie gruppi). Zoologicheskii Institut Akademia Nauk SSSR, Moskow, Leningrad, 372 pp.

    Medvedev, S.I. (1951) Fauna SSSR, Zhestkokrylye. Tom X, vyp. 1. Plastinchatousye (Scarabaeidae), Podsem. Melolonthinae, ch. 1 (khrushi). Zoologicheskii Institut Akademia Nauk SSSR, Moskow, Leningrad, 514 pp.

    Medvedev, S.I. (1952) Fauna SSSR, Zhestkokrylye. Tom X, vyp. 2. Plastinchatousye (Scarabaeidae), Podsem. Melolonthinae, ch. 2 (khrushi). Zoologicheskii Institut Akademia Nauk SSSR, Moskow, Leningrad, 276 pp.

    Medvedev, S.I. (1964) Fauna SSSR, Zhestkokrylye. Tom X, vyp. 5. Plastinchatousye (Scarabaeidae), Podsem. Cetoniinae, Valginae. Zoologicheskii Institut Akademia Nauk SSSR, Moskow, Leningrad, 376 pp.

    Motschulsky, V. (1860) Sur les Collections Coléoptèrologiques de Linné et de Fabricius. (Continuation). XXII. Lamellicornes. Études Entomologiques, 8[1859], 147–162.

    Mulsant, E. (1842) Histoire Naturelle des Coléoptères de France. Lamellicornes. Maison, Savy Jeune, Paris, Lyon, viii + 623 pp, 3 pls.

    Mulsant, E. & Godart, A. (1855) Description de quelques Coléoptères nouveaux ou peu connus. Opuscules Entomomologiques, 6, 1–8.

    Mulsant, E. & Rey, C. (1870) Histoire Naturelle des Coléoptères de France. Tribu des Lamellicornes. Annales de la Société d’Agriculture, Histoire Naturelle et Arts Utiles de Lyon, Quatrième Série, 2[1869], 241–650.

    Mulsant, E. & Rey, C. (1871) Histoire Naturelle des Coléoptéres de France. Tribu des Lamellicornes - suite. Annales de la Société d’Agriculture, Histoire Naturelle et Arts Utiles de Lyon, Quatrième Série, 3[1870], 155–530.

    Olivier, G. A. (1789) Entomologie, ou Histoire Naturelle des Insectes, avec leurs caractères génériques et spécifiques, leur description, leur synonymie, et leur figure enluminée. Coléoptères. Tome premier. Baudouin, Paris, xx + 433 pp. [Genera separately paginated]

    Panzer, G.W.F. (1796) Faunae Insectorum Germaniae Initia. oder Deutschlands Insecten. Heft 35. Felsecker, Nürnberg, 24 pp, 24 pls.

    Paulian, R. (1942) Exploration du Parc National Albert. Mission G. F. de Witte (193335). Fascicule 35. Aphodiinae (Coleoptera Lamellicornia) Fam. Scarabaeidae. Institut des Parcs Nationaux du Congo Belge, Bruxelles, 143 pp, 23 pls.

    Paulian, R. (1979) Un nouvel Aphodiidae du Portugal. Bulletin de la Société Entomologique de France, 84, 66–67.

    Pierotti, H. (1982) Gli Aphodius italiani del sottogenere Phalacronotus Motschulsky (Coleoptera Aphodiidae). Società Veneziana di Scienze Naturali - Lavori, 7, 17–44.

    Reiche, L.J. (1841) Tableau d’une division systématique de la tribu des Coprophages dans la famille des Lamellicornes. Revue Zoologique, 4, 211–213.

    Reiche, L.J. (1871) Examen de quelques espéces de Cétonides d’Europe et pays limitrophes et description de quatre espèces nouvelles. Annales de la Société Entomologique de France, 5e série, 1, 83–87.

    Reitter, E. (1890) Uebersicht der bekannten Hymenoplia-Arten. Wiener Entomologische Zeitung, 9, 259–263.

    Reitter, E. (1892) Bestimmungstabelle der Lucaniden und coprophagen Lamellicornen. XXIV. Heft, (Sonderabdruck aus dem XXX. Bande der Verhandlungen des naturforscheden Vereins in Brünn). Verlag des Verfassers, Brno, 230 pp.

    Reitter, E. (1896) Uebersicht der mir bekannten palaearctischen, mit der Coleopteren-Gattung Serica verwandten Gattungen und Arten. Wiener Entomologische Zeitung, 15, 180–188.

    Reitter, E. (1901) Weitere Beiträge zur Coleopteren-Fauna des russischen Reiches. Deutsche Entomologische Zeitschrift, 1901, 65–84.

    Reitter, E. (1902) Bestimmungs-Tabelle der Melolonthidae aus der Europaischen Fauna und den angrenzenden Ländern. III. Teil, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforscheden Vereins in Brünn, 40[1901], 93–303.

    Reitter, E. (1903) Bestimmungs-Tabelle der Melolonthidae aus der europäischen Fauna und den angrenzenden Ländern. IV. Teil (Schluss): enthaltend die Rutelini, Hopliini, und Glaphyrini. LI. Heft (51). Sonderabdruck aus dem XLI Bande der Verhandlungen des Naturforschenden Vereiner in Brünn. Verlag von Edm. Reitter, Druck von W. Burkart, Brno, 131 pp. [numbered 28–158]

    Reitter, E. (1919) Coleopterologische Notizen. Entomologische Blätter, 15, 220–221.

    Samouelle, G. (1819) The Entomologist’s Useful Compendium; or an introduction to the knowledge of British insects. Thomas Boys, London, 496 pp, 12 pls.

    Schmidt, A. (1913) Erster Versuch einer Einteilung der exotischen Aphodien in Subgenera und als Anhang einige Neubeschreibungen. Archiv für Naturgeschichte. Abtheilung A, Original-Arbeiten, 79, 117–178, 203–212.

    Schönherr, C.J. (1817) Synonymia Insectorum, oder Versuch einer Synonymie aller bisher bekannten Insecten nach Fabricii Systema Eleutheratorum &:c geordnet. Erster Band. Eleutherata oder K@fer. Dritter Theil: Hispa ... Molorchus. Lewerentzischen Buchdruckerey, Skara, xi + 506 pp, pls. 5–6.

    Scopoli, J.A. (1763) Entomologia Carniolica exhibens Insecta Carnioliae indigena et distributa in Ordines, Genera, Species, Varietates, Methodo Linneana. I. T. Trattner, Vienna, xxxvi + 420 + 2 (unpaged) pp, 37 pls.

    Scriba, L.G. (1790) Verzeichnis der Insekten in der Darmst@dter Gegend. Journal für die Liebhaber der Entomologie, 1, 40–73.

    Seabra, A.F. (1909) Estudos sobre os Animaes uteis ou nocivos à Agricultura. VI - EsboHo Monographico sobre os Scarabaeideos de Portugal (Aphodiini e Hybosorini). DirecHno Geral de Agricultura, PublicaHtes do Laboratorio de Pathologia Vegetal, Lisbon, 126 pp, 4 pls.

    Shipp, J.W. (1895) A revised classification of the genus Ateuchus, Weber. The Entomologist, 28, 218–221.

    Smetana, A. & Smith, A.B.T. (2006) Type species designations and other nomenclatural notes on Palaearctic Melolonthinae and Cetoniinae (Coleoptera: Scarabaeoidea: Scarabaeidae). Zootaxa, 1220, 47–53.

    Smith, A.B.T. (2004) Clarification on the nomenclatural status of six genus-group names in the tribe Trichiini (Coleoptera: Scarabaeidae: Cetoniinae). The Coleopterists Bulletin, 58, 285–291.

    Stephens, J. F. (1829) A Systematic Catalogue of British Insects: Being an attempt to arrange all the hitherto discovered indigenous insects in accordance with their natural affinitis. Containing also the references to every English writer on Entomology, and on the principal Foreign Authors. With all the published British genera to the present time. Baldwin and Cradock, London, xxxix + 416 + 388 pp.

    Stephens, J.F. (1830) Illustrations of British Entomology; or a Synopsis of Indigenous Insects: Containing their Generic and Specific Distinctions; with an Account of their Metamorphoses, Times of Appearance, Localities, Food and Economy as Far as Practicable. Mandibulata. Vol. III. Baldwin and Cradock, London, 374 pp, pls. 16–19.

    Stephens, J.F. (1839) A manual of British Coleoptera, or Beetles; Containing a Brief Description of all the Species of Beetles Hitherto Ascertained to Inhabit Great Britain. Longman, Orme Brown, Green and Longmans, London, xii + 443 pp.

    Tauzin, P. (2001) Le genre Aleurostictus Kirby, 1827. Contribution à sa connaissance et précision sur la distribution des espèces (Coleoptera, Cetoniidae, Trichiinae, Trichiini). L’Entomologiste, 56[2000], 231–281.

    TesaÕ , Z. (1957) Brouci ListorozR Lamellicornia, II. Scarabaeidae Laparostictí. Fauna ÑSR, 11. NakladatelstvR ÑeskoslovenskJ Akademie Vd, Prague, 326 pp, 16 pls.

    Thomson, C.G. (1859) Skandinaviens Coleoptera. Synoptiskt bearbetade. I. Tom. Berlingska Boktryckeriet, Lund, vi + 290 pp.

    Villa, A. & Villa, G. B. (1833) Coleoptera Europae dupleta in collectione Villa, quae pro mutua commutatione offeri possunt. Mediolani, 36 pp.

    Weber, F. (1801) Observationies Entomologicae, continentes novorum quae condidit Generum Characteres, et nuper detectarum Specierum Descriptiones. Bibliopolii Academici Novi, Kiel, xii + 116 pp.

    Weise, J. (1883) Noch einmal Geoffroy. Wiener Entomlogische Zeitung, 2, 150–151.

    Westwood, J.O. (1838) Stirps 2. Lamellicornes Latr. In: Westwood, J. O. – 1838–1840, Synopsis of the genera of British insects. Longman, Orme, Brown, Green & Longmans, London, 22–24.

    [Westwood, J.O.] (1845) A Catalogue of the Lucanoid Coleoptera in the collection of the Rev. F. W. Hope, M.A. F.R.S. & c. President of the Entomological Society of London, together with descriptions of the new species therein contained. J. C. Bridgewater, London, 31 pp. [Published anonymously].

    Ziani, S. (2002a) Sulle specie appartenenti al genere Scarabaeus Linnaeus, 1758 (sensu lato) presenti in Romagna (Insecta Coleoptera Scarabaeidae). Quaderno di Studi e Notizie di Storia Naturale della Romagna, 16, 27–35.

    Ziani, S. (2002b) On the question of priority between the names Parentius Zunino, 1979 and Relictonthophagus Kabakov, 1979 (Coleoptera: Scarabaeidae: Onthophagini). Elytron, 15[2001], 79–81.

    Zídek, J. & Pokornv , S. (2005) Supplemental comments on Scarabaeus Linn (Scarabaeinae: Scarabaeini). Animma.x, 9, 1–12.

    Zunino, M. (1979) Gruppi artificiali e grupi naturali negli Onthophagus (Coleoptera, Scarabaeoidea). Bollettino del Museo di Zoologia dell’Università di Torino, 1, 1–18.

    Zunino, M. (1984) Sistematica generica dei Geotrupinae (Coleoptera, Scarabaeoidea: Geotrupidae), filogenesi della sottofamiglia e considerazioni biogeografiche. Bollettino del Museo Regionale di Scienze Naturali - Torino, 2, 9–162.